Goniatite: Difference between revisions

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{{Short description|Extinct order of molluscs}}
{{Automatic taxobox
| fossil_range = {{fossil range|390|251.4}}[[Middle Devonian]] – [[Late Permian]]
| image = Fossile J 2.jpg
| image_caption = A polished Goniatitegoniatite [[fossil]]
| taxon = Goniatitida
| authority = [[Alpheus Hyatt|Hyatt]], 1884
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}}
 
'''Goniatids''', informally Goniatitesgoniatites, are [[Ammonoidea|ammonoid]] cephalopods[[cephalopod]]s that form the order '''Goniatitida''', derived from the more primitive [[Agoniatitida]] during the Middle [[Devonian]] some 390 million years ago (around [[Eifelian]] stage). Goniatites (goniatitidagoniatitids) survived the [[Late Devonian extinction]] to flourish during the [[Carboniferous]] and [[Permian]] only to become [[Permian–Triassic extinction event|extinct at the end of the Permian]] some 139 million years later.
 
== Morphology ==
[[File:Imitoceras rotatorium (ammonite).jpg|left|thumb|217x217px|The [[Prionoceratidae|prionoceratid]] ''[[Imitoceras|Imitoceras rotatorium]]'', showing characteristic goniatitic sutures. Early [[Carboniferous]], [[Rockford Limestone]], [[Rockford, Illinois|Rockford]], [[Illinois]].]]
 
All goniatites possessed an external shell, which is divided internally into chambers filled with gas giving it [[buoyancy]] during the life of the animal. An open chamber at the front of the shell provided living space for the goniatitid animal, with access to open water through a ventral siphuncle.
The general morphology and habit of goniatites was probably similar to that of their later relatives the [[Ammonitida|ammonites]], being free swimming and possessing a head with two well developed eyes and arms (or [[tentacle]]s).
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== Ecology ==
 
Ecologically, goniatites were limited to environments of normal-marine salinity—as appears to be the case for all cephalopods throughout their history. Goniatites are much more abundant and speciose in sediments that represent called [[Inland sea (geology)|epicontinental]] seas than they are in those that represent the open ocean. Within these inland seas, goniatites' greatest abundance and diversity appears to have been achieved in deeper offshore and [[Oceanic basin|basinal]] environments rather than in nearshore environments. Known nearshore (e.g., [[lagoon]]al) occurrences have generally been ascribed to wash-in of shells from offshore waters.
 
Due to lack of strong evidence for any particular life mode (e.g., [[benthos|benthic]] [[nekton]]ic, [[plankton]]ic), it remains unclear what resources goniatites were capitalizing on in these offshore environments. Only a few goniatites' full [[wikt:Special:Search/trophic|trophic]] apparatuses have ever been described, and reports of stomach contents in these creatures' fossils remain questionable at best. However, goniatites clearly lacked the calcified jaw apparatuses developed in later [[Ammonitida|ammonites]]; this has been cited as evidence against their having a [[durophagy|durophagous]] (shell-crushing) diet.
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[[Category:Middle Devonian first appearances]]
[[Category:Lopingian extinctions]]
[[Category:FossilPrehistoric taxacephalopod described in 1884orders]]