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{{Short description|Species of carnivorous plant}}
{{taxobox
{{speciesbox
|image = Roraima Heliamphora nutans1.JPG
|image_caption = ''Heliamphora nutans'' on [[Mount Roraima]]
|genus = ''[[Heliamphora]]''▼
|regnum = [[Plantae]]
|species = '''''H. nutans'''''▼
|unranked_divisio = [[Angiosperms]]
|binomial_authorityauthority = [[George Bentham|Benth.]] (1840)<ref name=Bentham>Bentham, G. (June 1840). [httphttps://www.biodiversitylibrary.org/page/4862618 XXV. On the ''Heliamphora nutans'', a new pitcher-plant from British Guiana]. ''The Transactions of the Linnean Society of London'' '''18'''(3): 429–433. {{DOIdoi|10.1111/j.1095-8339.1838.tb00190.x}}</ref>▼
|unranked_classis = [[Eudicots]]
|synonyms = *''Heliamphora heterodoxa''<br><small>''auct. non'' [[Julian Alfred Steyermark|Steyerm.]]: [[Gordon Cheers|G.Cheers]] (1992)</small>▼
|unranked_ordo = [[Asterids]]
*''Sarracenia nutans''<br><small>([[George Bentham|Benth.]]) [[David Nathaniel Friedrich Dietrich|D.Dietr.]] (1843)</small>▼
|ordo = [[Ericales]]
|familia = [[Sarraceniaceae]]
▲|genus = ''[[Heliamphora]]''
▲|species = '''''H. nutans'''''
|binomial = ''Heliamphora nutans''
▲|binomial_authority = [[George Bentham|Benth.]] (1840)<ref name=Bentham>Bentham, G. (June 1840). [http://biodiversitylibrary.org/page/4862618 XXV. On the ''Heliamphora nutans'', a new pitcher-plant from British Guiana]. ''The Transactions of the Linnean Society of London'' '''18'''(3): 429–433. {{DOI|10.1111/j.1095-8339.1838.tb00190.x}}</ref>
▲|synonyms = *''Heliamphora heterodoxa''<br><small>''auct. non'' Steyerm.: G.Cheers (1992)</small>
▲*''Sarracenia nutans''<br><small>(Benth.) D.Dietr. (1843)</small>
}}
 
'''''Heliamphora nutans''''' ([[Latin]]: ''nutans'' = nodding) is a species of [[Heliamphora|marsh pitcher plant]] native to the border area between [[Venezuela]], [[Brazil]] and [[Guyana]], where it grows on several [[tepui]]s, including [[Mount Roraima|Roraima]], [[Kukenán Tepui|Kukenán]], [[Yuruaní Tepui|Yuruaní]], [[Maringma Tepui|Maringma]], and [[Wei Assipu Tepui|Wei Assipu]]. ''Heliamphora nutans'' was the first ''Heliamphora'' to be described and is the best known species.<ref name=McPherson2011>McPherson, S., A. Wistuba, A. Fleischmann & J. Nerz 2011. ''[[Sarraceniaceae of South America]]''. Redfern Natural History Productions, Poole.</ref>
 
''Heliamphora nutans'' was originally discovered in 1839 on [[Mount Roraima]] by the two brothers [[Robert Hermann Schomburgk|Robert]] and [[Moritz Richard Schomburgk|Richard Schomburgk]],<ref name = "Orchids">{{cite web| url= http://www.orchids.co.in/orchidologists/david-burke.shtm|title = David Burke (1854 – 1897) |author= |publisher = www.orchids.co.in|accessdate=7 November 2008}}</ref> although they did not collect samples to return to Europe. The plant was [[Species description|formally described]] by [[George Bentham]] in 1840,<ref name=Bentham /> becoming the [[type species]] of the genus. In 1881, [[David Burke (botanist)|David Burke]] was plant-hunting in the same area of [[British Guiana]] where he collected specimens of the plant and introduced it to [[England]].<ref name = "HV87">{{cite book | author=[[James Herbert Veitch]]| title=[[Hortus Veitchii]]| publisher=[[Caradoc Doy]] | year=2006 |edition=reprint| isbn=978-0-9553515-0-21|page=87| title-link=Hortus Veitchii| author-link=James Herbert Veitch}}</ref>
 
This species employs an '[[aquaplaning]]' trapping mechanism (whereby prey animals slip into the pitchers on the downwards-pointing hairs, which are significantly more slippy when wet) similar to that of many tropical pitcher plants of the genus ''[[Nepenthes]]''.<ref>Bauer, U., M. Scharmann, J. Skepper & W. Federle 2013. 'Insect aquaplaning' on a superhydrophilic hairy surface: how ''Heliamphora nutans'' Benth. pitcher plants capture prey. ''Proceedings of the Royal Society B'' '''280'''(1753): 20122569. {{DOIdoi|10.1098/rspb.2012.2569}}</ref><ref>[http://www.cam.ac.uk/research/news/ants-aquaplaning-on-a-pitcher-plant Ants aquaplaning on a pitcher plant]. University of Cambridge.</ref>
 
<gallery>
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==References==
{{reflistReflist}}
 
==Further reading==
{{refbegin|40em}}
* Adlassnig, W., K. Pranji, E. Mayer, G. Steinhauser, F. Hejjas & I.K. Lichtscheidl (2010). The abiotic environment of ''Heliamphora nutans'' (Sarraceniaceae): pedological and microclimatic observations on Roraima Tepui. ''Brazilian Archives of Biology and Technology'' '''53'''(2): 425–430. {{DOIdoi|10.1590/S1516-89132010000200022}}
* Barrera, R., D. Fish & C.E. Machado-Allison (1989). [https://web.archive.org/web/20141221004256/http://ecotropicos.saber.ula.ve/db/ecotropicos/Edocs/vol2_n1/articulo4.pdf Ecological patterns of aquatic insect communities in two ''Heliamphora'' pitcher-plant species of the Venezuelan highlands.] ''Ecotropicos'' '''2'''(1): 31–44. <!--https://www.webcitation.org/6UyKztF8n-->
* {{esin iconlang|es}} Brewer-Carías, C. (2012). {{PDFlinkcite web |[url= http://www.plymouth.ac.uk/files/extranet/docs/I_UBC/RORAIMA_Madre_de_todos_los_rios.pdf |title= Roraima: madre de todos los ríos.] |access-date= 2013-03-07 |archive-url= https://web.archive.org/web/20160304025504/http://www6.plymouth.ac.uk/files/extranet/docs/I_UBC/RORAIMA_Madre_de_todos_los_rios.pdf |archive-date= 2016-03-04 |url-status= dead }} ''Río Verde'' '''8''': 77–94. <!--actually H. glabra-->
* Fleischmann, A., A. Wistuba & S. McPherson (21 December 2007). ''Drosera solaris'' (Droseraceae), a new sundew from the Guayana Highlands. ''Willdenowia'' '''37'''(2): 551–555. {{DOIdoi|10.3372/wi.37.37214}}
* Gonzalez, J.M., K. Jaffe & F. Michelangeli (1991). Competition for Prey Between the Carnivorous Bromeliaceae ''Brocchinia reducta'' and Sarraceniaceae ''Heliamphora nutans''. ''Biotropica'' '''23'''(4B): 602–604.
* JoelGonzalez, DJ.M., B.EK. JuniperJaffe & AF. DafniMichelangeli (December 19851991). UltravioletCompetition patternsfor inprey between the trapscarnivorous ofBromeliaceae carnivorous''Brocchinia plants.reducta'' and Sarraceneacea<!--sic--> ''NewHeliamphora Phytologistnutans''. ''Biotropica'101' '''23'''(44B): 585–593602–604. {{DOIJSTOR|10.1111/j.1469-8137.1985.tb02864.x2388398}}
* Joel, D.M., B.E. Juniper & A. Dafni (December 1985). Ultraviolet patterns in the traps of carnivorous plants. ''New Phytologist'' '''101'''(4): 585–593. {{doi|10.1111/j.1469-8137.1985.tb02864.x}}
* Kelloff, C.L., S.N. Alexander, V.A. Funk & H.D. Clarke (2011). [http://si-pddr.si.edu/jspui/bitstream/10088/17550/1/SCB97_Kelloff_web-FINAL.pdf Smithsonian Plant Collections, Guyana: 1995–2004, H. David Clarke.]{{Dead link|date=July 2024 |bot=InternetArchiveBot |fix-attempted=yes }} ''Smithsonian Contributions to Botany'' '''97''': i–viii, 1–307. <!--plant identified as H. nutans on Waukauyengtipu (Cerro Venamo)-->
* Kok, P.J.R. (20 October 2008). [http://www.mapress.com/zootaxa/2008/2/zt01909p015.pdf A new highland species of ''Arthrosaura'' Boulenger, 1885 (Squamata: Gymnophthalmidae) from Maringma tepui on the border of Guyana and Brazil.] ''Zootaxa'' '''1909''': 1–15. [{{PDFlinkcite web |[url= http://www.mapress.com/zootaxa/2008/f/z01909p015f.pdf |title= first page] }}]
* Lindley, J. (1846). [https://www.biodiversitylibrary.org/page/7433438 Order CLV. Sarraceniaceæ.—Sarraceniads.] In: [https://dx.doi.org/10.5962/bhl.title.7756 ''The Vegetable Kingdom; or, The Structure, Classification, and Uses of Plants, Illustrated upon the Natural System'']. Bradbury & Evans, London. p.&nbsp;429.
* McPherson, S. (2007). ''Pitcher Plants of the Americas''. The McDonald & Woodward Publishing Company, Blacksburg, Virginia.
* Nerz, J. (December 2004). [http://www.carnivorousplants.org/cpn/Species/v33n4p111_116.html ''Heliamphora elongata'' (Sarraceniaceae), a new species from Ilu-Tepui]. ''[[Carnivorous Plant Newsletter]]'' '''33'''(4): 111–116.
* Steyermark, J.A. ''et al.'' (18 May 1951). [httphttps://www.biodiversitylibrary.org/page/2455368 Sarraceniaceae]. [pp.&nbsp;239–242] In: [httphttps://www.biodiversitylibrary.org/page/2455345 Contributions to the flora of Venezuela. Botanical exploration in Venezuela - 1.] ''Fieldiana: Botany'' '''28'''(1): 1–242.
* Wistuba, A., T. Carow & P. Harbarth (September 2002). [http://www.carnivorousplants.org/cpn/Species/v31n3p78_82.html ''Heliamphora chimantensis'', a new species of ''Heliamphora'' (Sarraceniaceae) from the ‘Macizo de Chimanta’ in the south of Venezuela]. ''[[Carnivorous Plant Newsletter]]'' '''31'''(3): 78–82.
* [http://www.brucemeans.com/journal/journalVenezula2006.htm Journal of Venezuela Expedition January/February 2006 by D. Bruce Means]. Journals: D. Bruce Means, PhD. <!--https://www.webcitation.org/6UU2Mcfae-->
<!--http://www.cpukforum.com/forum/index.php?showtopic=33030 - Mt Roraima H. nutans + misidentified plants from Cerro Venamo (actually H. heterodoxa)
http://botany.si.edu/bdg/gallery_coll.cfm?lastname=Clarke - Trip 15 (misidentified Cerro Venamo H. heterodoxa)-->
https://www.youtube.com/watch?feature=player_embedded&v=t-n0-NFNFmM#!-->
{{refend}}
 
{{commonsCommons}}
 
{{Taxonbar|from=Q146102}}
{{Heliamphora}}
 
[[Category:Heliamphora|nutans]]
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[[Category:Flora of Brazil]]
[[Category:Flora of Guyana]]
[[Category:Flora of the Tepuis]]
 
{{Ericales-stub}}
 
[[pt:Heliamphora nutans]]
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