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{{Short description|LargeThird-largest species of baleen whale}}
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The '''sei whale''' ({{IPAc-en|s|eɪ}} {{respell|SAY}},<ref>{{OED|sei whale}}</ref> {{IPA-no|sæɪ|lang}}; '''''Balaenoptera borealis''''') is a [[baleen whale]]. It is one of ten [[rorqual]] species, and the third-largest member after the [[blue whale|blue]] and [[fin whale]]s. TheyIt can grow up to {{cvt|19.5|m}} in length and weigh as much as {{cvt|28|t}}. Two subspecies are recognized: ''B. b. borealis'' and ''B. b. schlegelii''. The whale's [[Anatomical terms of location#Dorsal and ventral|ventral]] surface has sporadic markings ranging from light grey to white, and its body is usually dark steel grey in colorcolour. It is among the fastest of all [[cetaceans]], and can reach speeds of up to {{cvt|50|km/h}} over short distances. 
 
It inhabits most oceans and adjoining seas, and prefers deep offshore waters. It avoids  [[polar water|polar]] and tropical waters and semi-enclosed bodies of water. The sei whale migrates annually from cool, subpolar waters in summer to [[temperate]], subtropical waters in winter with a lifespan of 70 years. It is a [[filter feeder]], with its diet consisting primarily of [[copepods]], [[krill]], and other [[zooplankton]]. It is typically solitary or can be found in groups numbering half a dozen. During the breeding period, a mating pair will remain together. Sei whale vocalizations usually lasts half a second, and occurs at 240–625 hertz.
 
Following large-scale [[commercial whaling]] during the late 19th and 20th centuries, when over 255,000 whales were killed, the sei whale is now internationally protected. It is listed as [[Endangered species|endangered]] by the [[IUCN Red List]], despite increasing populations. The Northern Hemisphere population is listed under [[CITES]] Appendix II, which indicates they are not threatened with [[extinction]], while the Southern Hemisphere population is listed under CITES Appendix I, indicating that they are threatened and are given the highest levels of protection.
 
== Etymology ==
"Sei whale" is an anglicization of the Norwegian ''seihval'', meaning "[[Pollachius virens|pollock]] whale".<ref>{{Cite OED|sei whale|1406967721}}</ref> The species was so called because it "appeared off the coast of Norway at the same time each year as the pollock that came to feed on the abundant plankton".<ref name="acs">{{cite web | url = http://www.acsonline.org/factpack/SeiBrydesWhales.htm | title = Sei Whale & Bryde's Whale ''Balaenoptera borealis'' & ''Balaenoptera edeni'' | publisher = American Cetacean Society |date=March 2004 | access-date = 8 November 2006 | archive-url = https://web.archive.org/web/20060927212353/http://www.acsonline.org/factpack/SeiBrydesWhales.htm <!--Added by H3llBot--> | archive-date = 27 September 2006 }}</ref> In the Pacific, the whale has been called the '''Japan finner'''; "finner" was a common term used to refer to rorquals. It has also been referred to as the '''lesser fin whale''' because it somewhat resembles the fin whale.<ref name=allen>{{cite book| url=https://books.google.com/books?id=30YZAAAAYAAJ&q=Balaenoptera+borealis&pg=PA234 | title=Whalebone Whales of New England | author = Glover Morrill Allen | year = 1916 | page = 234 | volume = 8 | access-date=24 June 2009 }}</ref>
 
== Taxonomy ==
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Two [[subspecies]] have been identified—the northern sei whale (''B. b. borealis'') and southern sei whale (''B. b. schlegelii'').<ref name="ITIS">{{ITIS | id = 180526 | taxon = Balaenoptera borealis | access-date = 10 November 2006 }}</ref>
 
=== EtymologyDescription ===
{{See also|Whale anatomy}}
[[File:Sei whale dorsal fin visible.jpg|thumb|right|A sei whale showing distinctive upright dorsal fin]]
The sei whale's body is typically a dark steel grey with irregular light grey to white markings on the ventral surface, or towards the front of the lower body. The whale has a relatively short series of 32–60 [[pleat]]s or grooves along its ventral surface that extend halfway between the pectoral fins and umbilicus (in other species it usually extends to or past the umbilicus), restricting the expansion of the buccal cavity during feeding compared to other species.<ref>{{cite journal | last1 = Brodie | first1 = P. | last2 = Víkingsson | first2 = G. | year = 2009 | title = On the feeding mechanism of the sei whale (''Balaenoptera borealis'')". | journal = Journal of Northwest Atlantic Fishery Science | volume = 42 | pages = 49–54 | doi=10.2960/j.v42.m646| doi-access = free }}</ref> The [[Rostrum (anatomy)|rostrum]] is pointed and the [[Fish anatomy#Types of fin|pectoral fins]] are relatively short, only 9–10% of body length, and pointed at the tips.<ref name="acs" /> Sei whales have a solitary ridge extending from the tip of the rostrum to the paired [[Blowhole (biology)|blowholes]] that are a distinctive characteristic of baleen whales.<ref>{{Cite web |last=Barnes |first=Morvan |title=Sei whale (Balaenoptera borealis) - MarLIN - The Marine Life Information Network |url=https://www.marlin.ac.uk/species/detail/103 |access-date=2023-12-28 |website=www.marlin.ac.uk |language=en-GB}}</ref>
 
Its skin is often marked by pits or wounds, which after healing become white scars. These are now known to be caused by [[Cookiecutter shark|cookie-cutter sharks]] (''Isistius brasiliensis'').<ref name="shevchenko">{{cite journal | author = Shevchenko, V.I. | year = 1977 | title = Application of white scars to the study of the location and migrations of sei whale populations in Area III of the Antarctic | journal = Rep. Int. Whal. Comm. | volume = Spec. Iss. 1 | pages = 130–134}}</ref> It has a tall, [[sickle]]-shaped [[dorsal fin]] that ranges in height from {{cvt|38|-|90|cm}} and averages {{cvt|53|-|56|cm}}, about two-thirds of the way back from the tip of the rostrum.<ref name=Matthews1938>{{cite journal| last1 = Matthews | first1 = L.H. | year = 1938 | title = The sei whale ''Balaenoptera borealis''". | journal = Discovery Reports | volume = 17 | pages = 183–290}}</ref> Dorsal fin shape, [[pigment]]ation pattern, and scarring have been used to a limited extent in [[Wildlife photo-identification|photo-identification]] studies.<ref name="schilling">{{cite journal | author = Schilling, M.R. | author2 = I. Seipt | author3 = M.T. Weinrich | author4 = S.E. Frohock | author5 = A.E. Kuhlberg | author6 = P.J. Clapham | year = 1992 | title = Behavior of individually identified sei whales ''Balaenoptera borealis'' during an episodic influx into the southern Gulf of Maine in 1986 | journal = Fish. Bull. | volume = 90 | pages = 749–755 | url = https://secure2.nni.com/whalecenter/pdfs/Sei_whales_FishBull92.pdf | url-status = dead | archive-url = https://web.archive.org/web/20210831001911/https://secure2.nni.com/whalecenter/pdfs/Sei_whales_FishBull92.pdf | archive-date = 31 August 2021 | df = dmy-all }}</ref> The tail is thick and the [[Whale#Appendages|fluke]], or lobe, is relatively small in relation to the size of the whale's body.<ref name="acs" />
The origin of ''sei'' is derived from [[pollock]] in Norwegian.<ref name="acs">{{cite web | url = http://www.acsonline.org/factpack/SeiBrydesWhales.htm | title = Sei Whale & Bryde's Whale ''Balaenoptera borealis'' & ''Balaenoptera edeni'' | publisher = American Cetacean Society |date=March 2004 | access-date = 8 November 2006 | archive-url = https://web.archive.org/web/20060927212353/http://www.acsonline.org/factpack/SeiBrydesWhales.htm <!--Added by H3llBot--> | archive-date = 27 September 2006 }}</ref> The specific name is the Latin word ''borealis'', meaning northern. The genus name ''Balaenoptera,'' is Latin for whale.<ref>{{Cite web |title=Definition of BALAENOPTERA |url=https://www.merriam-webster.com/dictionary/Balaenoptera |access-date=2024-01-17 |website=www.merriam-webster.com |language=en}}</ref><ref>{{Cite web |title=borealis {{!}} Etymology of borealis by etymonline |url=https://www.etymonline.com/word/borealis |access-date=2024-01-17 |website=www.etymonline.com |language=en}}</ref> In the Pacific, the whale has been called the Japan finner; "finner" was a common term used to refer to rorquals. In Japanese, the whale was called ''iwashi kujira'', or sardine whale, a name originally applied to [[Bryde's whale]]s by early Japanese whalers. Later, as modern whaling shifted to [[Sanriku]]—where both species occur—it was confused for the sei whale. Now the term only applies to the latter species.<ref>Omura, Hidero. "Review of the Occurrence of the Bryde's Whale in the Northwest Pacific". ''Rep. Int. Commn.'' (Special Issue 1), 1977, pp. 88–91.</ref><ref name="natlgeo1911">{{cite journal | url = http://www.edwardtbabinski.us/whales/shore_whaling_industry.html | archive-url = https://web.archive.org/web/20061004121048/http://www.edwardtbabinski.us/whales/shore_whaling_industry.html | url-status = dead | archive-date = 2006-10-04 | journal = National Geographic Magazine |date=May 1911 | title = Shore Whaling: A World Industry | author = Andrews, R.C. }}</ref> It has also been referred to as the '''lesser fin whale''' because it somewhat resembles the fin whale.<ref name=allen>{{cite book| url=https://books.google.com/books?id=30YZAAAAYAAJ&q=Balaenoptera+borealis&pg=PA234 | title=Whalebone Whales of New England | author = Glover Morrill Allen | year = 1916 | page = 234 | volume = 8 | access-date=24 June 2009 }}</ref>
 
[[File:Baleen.jpg|thumb|left|Close-up view of baleen plates, used to strain food from the water|alt=Photo displaying dozens of baleen plates: The plates face each other, and are evenly spaced at approximately {{cvt|0.25|in}} intervals. The plates are attached to the jaw at the top, and have hairs at the bottom end.]]
== Size ==
 
Adults have 300–380 ashy-black baleen plates on each side of the mouth, up to {{cvt|80|cm}} long. Each plate is made of fingernail-like [[keratin]], which is bordered by a fringe of very fine, short, curly, wool-like white bristles.<ref name="adw">{{cite web | author = Shefferly, N. | year = 1999 | title = Balaenoptera borealis | publisher = Animal Diversity Web | url = http://animaldiversity.ummz.umich.edu/site/accounts/information/Balaenoptera_borealis.html | access-date = 4 November 2006}}</ref> The sei's very fine baleen bristles, about {{cvt|0.1|mm}}, are the most reliable characteristic that distinguishes it from other rorquals.<ref name="mead">{{cite journal | author = Mead, J.G. | year = 1977 | title = Records of sei and Bryde's whales from the Atlantic coast of the United States, the Gulf of Mexico, and the Caribbean | journal = Rep. Int. Whal. Comm. | volume = Spec. Iss. 1 | pages = 113–116}}</ref>
 
The sei whale looks very similar to other large rorquals, especially to its smaller relative the Bryde's whale. Exceptional individuals may resemble a fin whale, which leads to confusion. They are usually differentiated from the fin whale by the colour of their head. WhenContrary viewed fromto the side,fin thewhale's smooth rostrum appears slightly arched (accentuated at, the tip),sei while fin and Brydewhale's whales have relativelyrostrum flatis rostrumscurved.<ref name="nmfs" />
 
=== Size ===
[[File:Balaenoptera borealis.jpg|thumb|left|Painting of a sei whale.]]
[[File:Ballena del MNHN.jpg|thumb|Sei whale skeleton]]
The sei whale is the third-largest balaenopterid, after the [[blue whale]] and the [[fin whale]].<ref name="av">{{cite journal | author = S.L. Perry | author2 = D.P. DeMaster | author3 = G.K. Silber | year = 1999 | title = Special Issue: The Great Whales: History and Status of Six Species Listed as Endangered Under the U.S. Endangered Species Act of 1973 | journal = Marine Fisheries Review | volume = 61 | issue = 1 | url = http://spo.nwr.noaa.gov/mfr611/mfr611.htm | pages = 52–58 | access-date = 26 April 2011 | archive-url = https://web.archive.org/web/20110721055959/http://spo.nwr.noaa.gov/mfr611/mfr611.htm | archive-date = 21 July 2011 | url-status = dead | df = dmy-all }}</ref> Adults usually weigh between {{cvt|15|-|20|MT}}.<ref name="Lockyer1976" /> FemalesThey areexhibit considerably[[sexual largerdimorphism]], with females outweighing and being longer than malestheir male counterparts.<ref name="nmfs">{{cite book | author = Reeves, R. | author2 = G. Silber | author3 = M. Payne | title = Draft Recovery Plan for the Fin Whale ''Balaenoptera physalus'' and Sei Whale ''Balaenoptera borealis'' | publisher = National Marine Fisheries Service | date = July 1998 | location = Silver Spring, Maryland | url = http://www.cresli.org/cresli/pdf%20documents/finwhale.pdf | access-date = 5 November 2006 | archive-url = https://web.archive.org/web/20160303204014/http://www.cresli.org/cresli/pdf%20documents/finwhale.pdf | archive-date = 3 March 2016 | url-status = dead }}</ref> At birth, a calf typically measures {{cvt|4.4|-|4.5|m}} in length.<ref>{{Cite book |last= |first= |url=https://books.google.com/books?id=gfopb_WsW4YC&dq=info:f3100ZalxFIJ:scholar.google.com/&pg=RA3-PA25 |title=Marine Fisheries Review |date=1984 |publisher=NMFS, Scientific Publications Office |pages=27 |language=en}}</ref>
 
In the Northern Hemisphere, males reach up to {{cvt|17.1|m}} and females up to {{cvt|18.6|m}},<ref name=Klinowska>{{cite book | last = Klinowska| first = M.| title = Dolphins, Porpoises and Whales of the World: The IUCN Red Data Book| publisher = IUCN| year = 1991| location = Cambridge, U.K.}}</ref> while in the Southern Hemisphere males reach a maximum of {{cvt|18.6|m}} and females of {{cvt|19.5|m}}. The authenticity of an alleged {{cvt|22|m}} female caught {{cvt|80|km}} northwest of [[St Kilda, Scotland|St. Kilda]] in July 1911 is doubted.<ref name="Skinner2006">Skinner, J.D. and Christian T. Chimimba. (2006). ''The Mammals of the Southern African Sub-region''. Cambridge University Press, Third Edition.</ref><ref name=WentworthThompson1919>Thompson, D'Arcy Wentworth. "On whales landed at the Scottish whaling stations, especially during the years 1908–1914—Part VII. The sei-whale". ''The Scottish Naturalist'', nos. 85-96 (1919), pp. 37–46.</ref><ref name="martin83" /> Five specimens taken off [[Iceland]] exceeded {{cvt|14.6|m}} in length.<ref name="martin83">{{cite journal | author = Martin, A.R. | year = 1983 | title = The sei whale off western Iceland. I. Size, distribution and abundance | journal = Rep. Int. Whal. Comm. | volume = 33 | pages = 457–463 }}</ref><ref name="Mitchell1975">{{cite journal | last1 = Mitchell | first1 = E.D. | year = 1975 | title = Preliminary report on Nova Scotia fishery for sei whales (''Balaenoptera borealis'')". | journal = Reports of the International Whaling Commission | volume = 25 | pages = 218–225 }}</ref> The longest measured during JARPN II cruises in the North Pacific were a {{cvt|16.32|m}} female and a {{cvt|15|m}} male.<ref name="JARPNII2005">{{cite journal| last1 = Tamura | display-authors = etal | year = 2005 | title = Cruise report of the second phase of the Japanese Research Program under Special Permit in the Western North Pacific (JAPRN II) in 2005 – Offshore component | journal = Rep. Int. Whal. Comm. | volume = 58 | pages = 1–52}}</ref><ref name="JAPRNII2006">{{cite journal | last1 = Tamura | display-authors = etal | year = 2006 | title = Cruise report of the second phase of the Japanese Research Program under Special Permit in the Western North Pacific (JAPRN II) in 2006 (part I) – Offshore component | journal = Rep. Int. Whal. Comm. | volume = 59 | pages = 1–26 }}</ref>
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In the North Pacific, males weigh an average of {{cvt|15|t}} and females {{cvt|18.5|t}}. North Atlantic sei whale males average {{cvt|15.5|t}} and females {{cvt|17|t}}. Southern Hemisphere whales average {{cvt|17|-|18.5|t}} in body weight.<ref name="Lockyer1976" /><ref name="evans1987"/>
 
== Anatomy ==
{{See also|Whale anatomy}}
[[File:Sei whale dorsal fin visible.jpg|thumb|right|A sei whale showing distinctive upright dorsal fin]]
The sei whale's body is typically a dark steel grey with irregular light grey to white markings on the ventral surface, or towards the front of the lower body. The whale has a relatively short series of 32–60 [[pleat]]s or grooves along its ventral surface that extend halfway between the pectoral fins and umbilicus (in other species it usually extends to or past the umbilicus), restricting the expansion of the buccal cavity during feeding compared to other species.<ref>{{cite journal | last1 = Brodie | first1 = P. | last2 = Víkingsson | first2 = G. | year = 2009 | title = On the feeding mechanism of the sei whale (''Balaenoptera borealis'')". | journal = Journal of Northwest Atlantic Fishery Science | volume = 42 | pages = 49–54 | doi=10.2960/j.v42.m646| doi-access = free }}</ref> The [[Rostrum (anatomy)|rostrum]] is pointed and the [[Fish anatomy#Types of fin|pectoral fins]] are relatively short, only 9–10% of body length, and pointed at the tips.<ref name="acs" /> Sei whales have a solitary ridge extending from the tip of the rostrum to the paired [[Blowhole (biology)|blowholes]] that are a distinctive characteristic of baleen whales.<ref>{{Cite web |last=Barnes |first=Morvan |title=Sei whale (Balaenoptera borealis) - MarLIN - The Marine Life Information Network |url=https://www.marlin.ac.uk/species/detail/103 |access-date=2023-12-28 |website=www.marlin.ac.uk |language=en-GB}}</ref>
 
Its skin is often marked by pits or wounds, which after healing become white scars. These are now known to be caused by [[Cookiecutter shark|cookie-cutter sharks]] (''Isistius brasiliensis'').<ref name="shevchenko">{{cite journal | author = Shevchenko, V.I. | year = 1977 | title = Application of white scars to the study of the location and migrations of sei whale populations in Area III of the Antarctic | journal = Rep. Int. Whal. Comm. | volume = Spec. Iss. 1 | pages = 130–134}}</ref> It has a tall, [[sickle]]-shaped [[dorsal fin]] that ranges in height from {{cvt|38|-|90|cm}} and averages {{cvt|53|-|56|cm}}, about two-thirds of the way back from the tip of the rostrum.<ref name=Matthews1938>{{cite journal| last1 = Matthews | first1 = L.H. | year = 1938 | title = The sei whale ''Balaenoptera borealis''". | journal = Discovery Reports | volume = 17 | pages = 183–290}}</ref> Dorsal fin shape, [[pigment]]ation pattern, and scarring have been used to a limited extent in [[Wildlife photo-identification|photo-identification]] studies.<ref name="schilling">{{cite journal | author = Schilling, M.R. | author2 = I. Seipt | author3 = M.T. Weinrich | author4 = S.E. Frohock | author5 = A.E. Kuhlberg | author6 = P.J. Clapham | year = 1992 | title = Behavior of individually identified sei whales ''Balaenoptera borealis'' during an episodic influx into the southern Gulf of Maine in 1986 | journal = Fish. Bull. | volume = 90 | pages = 749–755 | url = https://secure2.nni.com/whalecenter/pdfs/Sei_whales_FishBull92.pdf | url-status = dead | archive-url = https://web.archive.org/web/20210831001911/https://secure2.nni.com/whalecenter/pdfs/Sei_whales_FishBull92.pdf | archive-date = 31 August 2021 | df = dmy-all }}</ref> The tail is thick and the [[Whale#Appendages|fluke]], or lobe, is relatively small in relation to the size of the whale's body.<ref name="acs" />
 
[[File:Baleen.jpg|thumb|left|Close-up view of baleen plates, used to strain food from the water|alt=Photo displaying dozens of baleen plates: The plates face each other, and are evenly spaced at approximately {{cvt|0.25|in}} intervals. The plates are attached to the jaw at the top, and have hairs at the bottom end.]]
 
Adults have 300–380 ashy-black baleen plates on each side of the mouth, up to {{cvt|80|cm}} long. Each plate is made of fingernail-like [[keratin]], which is bordered by a fringe of very fine, short, curly, wool-like white bristles.<ref name="adw">{{cite web | author = Shefferly, N. | year = 1999 | title = Balaenoptera borealis | publisher = Animal Diversity Web | url = http://animaldiversity.ummz.umich.edu/site/accounts/information/Balaenoptera_borealis.html | access-date = 4 November 2006}}</ref> The sei's very fine baleen bristles, about {{cvt|0.1|mm}}, are the most reliable characteristic that distinguishes it from other rorquals.<ref name="mead">{{cite journal | author = Mead, J.G. | year = 1977 | title = Records of sei and Bryde's whales from the Atlantic coast of the United States, the Gulf of Mexico, and the Caribbean | journal = Rep. Int. Whal. Comm. | volume = Spec. Iss. 1 | pages = 113–116}}</ref>
 
The sei whale looks very similar to other large rorquals, especially to its smaller relative the Bryde's whale. Exceptional individuals may resemble a fin whale, which leads to confusion. They are usually differentiated from the fin whale by the colour of their head. When viewed from the side, the rostrum appears slightly arched (accentuated at the tip), while fin and Bryde's whales have relatively flat rostrums.<ref name="nmfs" />
 
== Life history ==
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=== Surface behaviours ===
Very little is known about the sei whale [[social structure]]. They have been documented traveling alone or in pods of up to six individuals; larger groups may assemble at particularly abundant feeding grounds.<ref name="edds84">{{cite journal | author = Edds, P.L. |author2 = T.J. MacIntyre|author3 = R. Naveen | year = 1984 | title = Notes on a sei whale (''Balaenoptera borealis'' Lesson) sighted off Maryland | journal = Cetus | volume = 5 | issue = 2 | pages = 4–5 }}</ref><ref name="schilling" /> During the southern Gulf of Maine influx in mid-1986, groups of at least three sei whales were observed "milling" on four occasions – i.e. moving in random directions, rolling, and remaining at the surface for over 10 minutes. One whale would always leave the group during or immediately after such socializing bouts.<ref name="schilling" />
The sei whale is among the fastest [[cetacea]]ns. The American [[naturalist]] [[Roy Chapman Andrews]] compared the sei whale to the [[cheetah]], because it can swim at great speeds "for a few hundred yards", but it "soon tires if the chase is long" and "does not have the strength and staying power of its larger relatives".<ref>Andrews, Roy Chapman. 1916. ''Whale hunting with gun and camera; a naturalist's account of the modern shore-whaling industry, of whales and their habits, and of hunting experiences in various parts of the world''. New York: D. Appleton and Co., p. 128.</ref> It can reach speeds of up to {{cvt|50|km/h}} over short distances.<ref name="adw" /> However, it is not a remarkable diver, reaching relatively shallow depths for 5 to 15 minutes. Between dives, the whale surfaces for a few minutes, remaining visible in clear, calm waters, with blows occurring at intervals of about 60 seconds (range: 45–90 sec.). UnlikeWhen theabout finto whaledive, the sei whale tendsusually notjust tosinks risebelow highthe outsurface; ofonly the waterdorsal asfin it dives, usually just sinking below the surface. Theand [[Blowhole (biology)|blowholes]] and dorsal fin are often exposed above the water surface almost simultaneouslyprotrude. The whale almost never lifts its flukes above the surface, and is generally less [[WhaleCetacean surfacing behaviour|active on water surfaces]] than theother closelywhale related Bryde's whalespecies; itthey rarely exhibit [[Whale#Behavior|breacheslobtail]] behaviour.<ref name="nmfs" />
 
=== Feeding ===
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=== Vocalizations ===
{{See also|Whale sound}}
The sei whale makes long, loud, low-frequency sounds. Relatively little is known about specific calls, but in 2003, observers noted sei whale calls in addition to sounds that could be described as "growls" or "whooshes" off the coast of the [[Antarctic Peninsula]].<ref name="mcdonald05">{{cite journal | author = McDonald, M. |author2=Hildebrand, J. |author3=Wiggins, S. |author4=Thiele, D. |author5=Glasgow, D. |author6=Moore, S. | title = Sei whale sounds recorded in the Antarctic | journal = The Journal of the Acoustical Society of America |date=December 2005 | volume = 118 | issue = 6 | pmid = 16419837 | pages = 3941–3945 | url = https://escholarship.org/content/qt6z22w129/qt6z22w129.pdf?t=lnrzou | doi = 10.1121/1.2130944|bibcode=2005ASAJ..118.3941M |s2cid=2094987 }}</ref> Many calls consisted of multiple parts at different frequencies. This combination distinguishes their calls from those of other whales. Most calls lastedlast about a half second, and occurredoccur in the 240–62537–98 [[hertz]] range, well within the range of human hearing. The maximum volume of the vocal sequences is reported as 156 [[decibel]]s relative to 1 [[Pascal (unit)|micropascal (μPa)]] at a reference distance of one metre.<ref name="mcdonald05" /> An observer situated one metre from a vocalizing whale would perceive a volume roughly equivalent to the volume of a jackhammer operating two meters away.<ref>Direct comparisons of sounds in water to sounds in air can be complicated, see [http://www.surtass-lfa-eis.com/Terms/index.htm this description] {{webarchiveusurped|url1=[https://web.archive.org/web/20090725234254/http://www.surtass-lfa-eis.com/Terms/index.htm |date=25 July 2009this description]}} for more information.</ref>
 
In November 2002, scientists recorded calls in the presence of sei whales off [[Maui]]. All the calls were downswept tonal calls, all but two ranging from a mean high frequency of 39.1&nbsp;Hz down to 21&nbsp;Hz of 1.3 second duration – the two higher frequency downswept calls ranged from an average of 100.3&nbsp;Hz to 44.6&nbsp;Hz over 1 second of duration. These calls closely resembled and coincided with a peak in "20- to 35-Hz irregular repetition interval" downswept pulses described from seafloor recordings off [[Oahu]], which had previously been attributed to fin whales.<ref>{{cite journal | last1 = Rankin | first1 = S. | last2 = Barlow | first2 = J. | year = 2007 | title = Vocalizations of the sei whale ''Balaenoptera borealis'' off the Hawaiian Islands | journal = Bioacoustics | volume = 16 | issue = 2| pages = 137–145 | doi=10.1080/09524622.2007.9753572| bibcode = 2007Bioac..16..137R | s2cid = 85413269 }}</ref> Between 2005 and 2007, low frequency downswept vocalizations were recorded in the Great South Channel, east of [[Cape Cod]], [[Massachusetts]], which were associated with the presence of sei whales. These calls averaged 82.3&nbsp;Hz down to 34&nbsp;Hz over about 1.4 seconds in duration. This call has also been reported from recordings in the Gulf of Maine, [[New England]] shelf waters, the mid-Atlantic Bight, and in [[Davis Strait]]. It likely functions as a contact call.<ref>{{cite journal | last1 = Baumgartner | first1 = M.F. | last2 = Van Parijs | first2 = S.M. | last3 = Wenzel | first3 = F.W. | last4 = Tremblay | first4 = C.J. | last5 = Esch | first5 = H.C. | last6 = Ward | first6 = A.A. | year = 2008 | title = Low frequency vocalizations attributed to sei whales (''Balaenoptera borealis'')". | journal = Journal of the Acoustical Society of America | volume = 124 | issue = 2| pages = 1339–1349 | doi=10.1121/1.2945155| pmid = 18681619 | hdl = 1912/4618 | bibcode = 2008ASAJ..124.1339B | hdl-access = free }}</ref>
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Sei whales live in all oceans, although rarely in polar or tropical waters.<ref name="nmfs" /> The difficulty of differentiating them at sea from their close relatives, Bryde's whales and in some cases from fin whales, creates confusion about their range and population, especially in warmer waters where Bryde's whales are most common.<ref>{{Cite book |last=Tinker |first=Spencer Wilkie |url=https://books.google.com/books?id=ASIVAAAAIAAJ&dq=Sei+whale&pg=PA283 |title=Whales of the World |date=1988-01-01 |publisher=Brill Archive |isbn=978-0-935848-47-2 |pages=281 |language=en}}</ref><ref>{{Cite book |url=https://books.google.com/books?id=2D03AQAAMAAJ&dq=Sei+whale&pg=SA4-PA61 |title=Amending the Atlantic Large Whale Take Reduction Plan, Broad-based Gear Modifications: Environmental Impact Statement |publisher=Environmental Impact Statement |year=2007 |pages=4–62 |language=en}}</ref>
 
In the North Atlantic, its range extends from [[southern Europe]] or [[Northwest Africa|northwestern Africa]] to Norway, and from the [[southern United States]] to [[Greenland]].<ref name="Gambell85a">{{cite book | author = Gambell, R. | year = 1985 | chapter = Sei Whale ''Balaenoptera borealis'' Lesson, 1828 | title = Handbook of Marine Mammals, Vol. 3 | editor = S.H. Ridgway | editor2 = R. Harrison | publisher = Academic Press | location = London | pages = 155–170 }}</ref> The southernmost confirmed records are strandings along the northern [[Gulf of Mexico]] and in the [[Greater Antilles]].<ref name="mead" /> ThroughoutIt itsrarely range, the whale tends not to frequent semienclosed bodies of water, such as the Gulf of Mexico,enters the [[Gulf of Saint Lawrence]], [[Hudson Bay]], the [[NorthMediterranean Sea]], and the [[Mediterranean Sea]].<ref name="nmfs" /> It occurs predominantly in deep water, occurring most commonly over the [[continental shelf|continental slope]],<ref name="CETAP">{{cite report | author = CETAP | year = 1982 | title = Final ReportGulf of the Cetacean and Turtle Assessment ProgramMexico, Universitywhich ofare Rhodeboth Island,considered to Bureaube ofsmall Landwaterbodies. ManagementSei |whales publisherare =pelagic U.S.and Departmentare oftypically the Interior. Ref. No. AA551-CT8–48}}</ref>found in basins situated between banks,<ref name="sutcliffe">{{cite book | author1= Sutcliffe, W.H. Jr. |author2=P.F. Brodie | year = 1977 | chapter = Whale distributions in Nova Scotia waters | title = Fisheries & Marine Service Technical Report No. 722 }}</ref>oceans or [[submarine canyon]]open areasseas.<ref name="kenney87nmfs">{{cite journal | author1= Kenney, R.D. |author2=H.E. Winn | year = 1987 | title = Cetacean biomass densities near submarine canyons compared to adjacent shelf/slope> areas | journal = Cont. Shelf Res. | volume = 7 | pages = 107–114 | doi = 10.1016/0278-4343(87)90073-2 | issue=2|bibcode=1987CSR.....7..107K }}</ref>
 
In the North Pacific, it ranges from [[20th parallel north|20°N]] to [[23rd parallel north|23°N]] [[latitude]] in the winter, and from [[35th parallel north|35°N]] to [[50th parallel north|50°N]] latitude in the summer.<ref name="masaki76">{{cite journal | author = Masaki, Y. | year = 1976 | title = Biological studies on the North Pacific sei whale | journal = Bull. Far Seas Fish. Res. Lab. | volume = 14 | pages = 1–104 }}</ref> Approximately 75% of the North Pacific population lives east of the [[International Date Line]].<ref name="horwood87">{{cite book |author=Horwood, J. |year=1987 |title=The sei whale: population biology, ecology, and management |publisher=Croom Helm Ltd. |location=Kent, England |isbn=978-0-7099-4786-8}}</ref> {{As of|2017|February}}, the U.S. [[National Marine Fisheries Service]] estimated that the eastern North Pacific population stood at 374 whales.<ref>{{cite web |url=https://www.fisheries.noaa.gov/webdam/download/70095913 |title=SEI WHALE (Balaenoptera borealis borealis): Eastern North Pacific Stock |work=[[National Marine Fisheries Service]] Marine Mammal Stock Assessment Reports |date=2017-02-08 |access-date=2019-05-23}}</ref> Two whales tagged in deep waters off California were later recaptured off [[Washington (state)|Washington]] and [[British Columbia]], revealing a possible link between these areas,<ref name="rice74">{{cite book | author = Rice, D.W. | year = 1974 | chapter = Whales and whale research in the North Pacific | editor = Schervill, W.E. | title = The Whale Problem: a status report | pages = [https://archive.org/details/whaleproblemstat0000inte/page/170 170–195] | publisher = Harvard University Press | location = Cambridge, Massachusetts | isbn = 978-0-674-95075-7 | chapter-url = https://archive.org/details/whaleproblemstat0000inte/page/170 }}</ref> but the lack of other tag recovery data makes these two cases inconclusive. Occurrences within the [[Gulf of California]] have been fewer.<ref>Gendron D.. Rosales C. S.. 1996. [http://www.aquaticmammalsjournal.org/share/AquaticMammalsIssueArchives/1996/AquaticMammals_22-02/22-02_Gendron.pdf Recent sei whale (Balaenoptera borealis) sightings in the Gulf of California, Mexico]. Aquatic Mammals 1996, 22.2, pp.127-130 (pdf). Retrieved on February 24, 2017</ref> In [[Sea of Japan]] and [[Sea of Okhotsk]], whales are not common, although whales were more commonly seen than today in southern part of Sea of Japan. There had been a sighting in [[Golden Horn Bay]],<ref>{{cite web|url=http://www.zoosite.com.ua/animal-sei-whale-642.html|title=Сейвал / Balaenoptera borealis|website=www.zoosite.com.ua}}</ref> and whales were much more abundant in the triangle area around [[Kunashir Island]] in whaling days, making the area well known as sei – ground.<ref>Uni Y.,2006 [http://www.cho.co.jp/natural-h/download/archive/shiretoko/2703s_UNI.pdf Whales, Dolphins and Porpoises off Shiretoko]. Bulletin of the
Shiretoko Museum 27: pp.37-46. Retrieved on 16 December 2015</ref> There had been sightings of the species off the [[Sea of Japan]] during cetacean surveys.<ref>Omura, H. and Fujino, K. (1954) [https://www.icrwhale.org/pdf/SC00989-103.pdf Sei Whales in the Adjacent Waters of Japan. II. Further Studies on the External Characters.] Scientific Report of the Whales Research Institute, 9, 89-103.</ref>
 
Sei whales have been recorded from northern Indian Ocean as well such as around [[Sri Lanka]] and [[India]]n coasts.<ref>Sathasivam K.. 2015. [https://www.cbd.int/doc/meetings/mar/ebsaws-2015-01/other/ebsaws-2015-01-gobi-submission5-en.pdf A CATALOGUE OF INDIAN MARINE MAMMAL RECORDS] (pdf)</ref> In the Southern Hemisphere, summer distribution based upon historic catch data is between [[40th parallel south|40°S]] and [[50th parallel south|50°S]] latitude in the South Atlantic and southern Indian Oceans and [[45th parallel south|45°S]] and [[60th parallel south|60°S]] in the South Pacific, while winter distribution is poorly known, with former winter whaling grounds being located off northeastern Brazil ([[7th parallel south|7°S]]) and Peru ([[6th parallel south|6°S]]).<ref name="iucn" /> The majority of the "sei" whales caught off Angola and Congo, as well as other nearby areas in equatorial West Africa, are thought to have been predominantly misidentified [[Bryde's whale]]s. For example, Ruud (1952) found that 42 of the "sei whale" caught off [[Gabon]] in 1952 were actually Bryde's whales, based on examination of their baleen plates. The only confirmed historical record is the capture of a {{convert|14|m|ft|abbr=on}} female, which was brought to the Cap Lopez whaling station in Gabon in September 1950. During cetacean sighting surveys off Angola between 2003 and 2006, only a single confirmed sighting of two individuals was made in August 2004, compared to 19 sightings of Bryde's whales.<ref>{{cite journal| last1 = Weir | first1 = C.R. | year = 2010 | title = A review of cetacean occurrence in West African waters from the Gulf of Guinea to Angola | journal = Mammal Review | volume = 40 | pages = 2–39 | doi=10.1111/j.1365-2907.2009.00153.x}}</ref> Sei whales are commonly distributed along west to southern Latin America, including the entire Chilean coast down to the [[Beagle Channel]].<ref>Daniel.Bisson – [https://www.panoramio.com/photo/71888436 SEI Whale at Beagle canal, Ushuaia, Argentina] {{webarchive|url=https://web.archive.org/web/20160413175552/http://www.panoramio.com/photo/71888436 |date=13 April 2016 }}</ref><ref>OceanSounds e.V. - [http://ocean-sounds.org/en_US/whales-in-patagonia/marine-mammals-in-patagonia/sei-whale/ Sei whale] {{Webarchive|url=https://web.archive.org/web/20160413012011/http://ocean-sounds.org/en_US/whales-in-patagonia/marine-mammals-in-patagonia/sei-whale/ |date=13 April 2016 }}</ref> The Falkland Islands appear to be a regionally important area for the sei whale, as a small population exists in coastal waters off the eastern Falkland or Malvinas archipelago. For reasons unknown, the whales prefer to stay inland here, even venturing into large bays.<ref>{{Cite journal |last1=Baines |first1=Mick |last2=Weir |first2=Caroline R. |date=2020-12-23 |title=Predicting suitable coastal habitat for sei whales, southern right whales and dolphins around the Falkland or Malvinas Islands |journal=PLOS ONE |language=en |volume=15 |issue=12 |pages=e0244068 |doi=10.1371/journal.pone.0244068 |doi-access=free |issn=1932-6203 |pmc=7757899 |pmid=33362235|bibcode=2020PLoSO..1544068B }}</ref>
 
=== Migration ===
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== Whaling ==
{{Main|Whaling|History of whaling}}
The development of explosive [[harpoon]]s and steam-powered whaling ships in the late nineteenth century brought previously unobtainable large whales within the reach of commercial [[Whaling|whalers]]. Initially, the sei whale's speed and elusiveness partially protected them,<ref name="sigurjónsson88">{{cite journal | author = Sigurjónsson, J.|year = 1988 |title= Operational factors of the Icelandic large whale fishery | journal = Rep. Int. Whal. Comm. |volume = 38 |pages = 327–333}}</ref> and later the comparatively small yield of [[Whale oil|oil]] and [[whale meat|meat]]. Once stocks of more profitable [[right whale]]s, blue whales, fin whales, and humpback whales became depleted, sei whales were hunted in earnest, particularly from 1950 to 1980.<ref name="av" />
 
=== North Atlantic ===
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In the North Atlantic between 1885 and 1984, 14,295 sei whales were taken.<ref name="horwood87" /> They were hunted in large numbers off the coasts of Norway and Scotland beginning in the late 19th and early 20th centuries,<ref name="jonsgard and darling" /> and in 1885 alone more than 700 were caught off [[Finnmark]].<ref name="andrews16">{{cite journal | author = Andrews, R.C.| year= 1916 | title= The sei whale (''Balaenoptera borealis'' Lesson)| journal = Mem. Am. Mus. Nat. Hist. |series=New Series | volume = 1 | issue = 6 | pages = 291–388}}</ref> Their meat was a popular Norwegian food. The meat's value made the hunting of this difficult-to-catch species profitable in the early twentieth century.<ref>{{cite book | author = Ingebrigtsen, A.| year = 1929 | chapter = Whales caught in the North Atlantic and other seas | title = Rapports et Procès-verbaux des réunions, Cons. Perm. Int. L'Explor. Mer, Vol. LVI| publisher = Høst & Fils | location = Copenhagen }}</ref>
 
In Iceland, a total of 2,574 whales were taken from the [[Hvalfjörður]] whaling station between 1948 and 1985. Since the late 1960s to early 1970s, the sei whale has beenwas second only to the fin whale as the preferred target of Icelandic whalers, with meat in greater demand than [[whale oil]], the prior target.<ref name="sigurjónsson88" />
 
Small numbers were taken off the [[Iberian Peninsula]], beginning in the 1920s by Spanish whalers,<ref name="aguilar and lens">{{cite journal | author = Aguilar, A. |author2=S. Lens | year = 1981 | title = Preliminary report on Spanish whaling operations | journal = Rep. Int. Whal. Comm. |volume = 31 | pages = 639–643}}</ref> off the [[Nova Scotia]]n shelf in the late 1960s and early 1970s by Canadian whalers,<ref name="mitchell and chapman" /> and off the coast of West Greenland from the 1920s to the 1950s by Norwegian and Danish whalers.<ref name="kapel85" />
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[[Category:Cetaceans of the Arctic Ocean]]
[[Category:Baleen whalesBalaenoptera]]
[[Category:Mammals of Japan]]
[[Category:Cetaceans of the Indian Ocean]]