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Miles Warren was a [[professor]] of [[biology]] at [[Empire State University]].<ref name="mnyc">{{cite book | last = Sanderson | first = Peter | authorlink = | coauthors = | title = The Marvel Comics Guide to New York City | publisher = [[Pocket Books]] | date = 2007 | location = New York City | pages = 30–33 | url = | doi = | id = | isbn = 1-14653-141-6}}</ref> There, he met [[Spider-Man|Peter Parker]] and [[Gwen Stacy]].<ref>''Amazing Spider-Man'' #31</ref> Warren secretly fell in love with the much younger Stacy, and after she was killed by the original [[Green Goblin]], Warren swore vengeance on Spider-Man, falsely believing it was he who caused her death. After his lab assistant, Anthony Serba, revealed that he had successfully cloned a frog, Warren gave Serba tissue samples taken from Gwen, telling him that they were rat cells. Sometime later, Serba confronts Warren, telling him that the clones are human and have to be destroyed immediately. Panicking, Warren covers Serba's mouth to shut him up, accidentally suffocated him. Unable to accept responsibility for his action, Warren creates an alter ego he calls "The Jackal", which he took after overhearing a professor describing a [[jackal]] as a "cowardly predator". Warren further develops his alter ego by fashioning a green suit and training himself athletically.
The Jackal harasses Spider-Man numerous times, setting him against various other adversaries. He allied himself with the [[Punisher]] against Spider-Man, but his alliance with the Punisher was quickly dissolved.<ref>''Amazing Spider-Man'' #129</ref> He then attempts to incite a gang war between [[Hammerhead (comics)|Hammerhead]] and [[Doctor Octopus]].<ref>''Amazing Spider-Man'' #130</ref> The Jackal equipped wrestler Maxwell Markham with the costume and powerful exoskeleton of the [[Grizzly (comics
Meanwhile, his cloning experiments had varying results; out of numerous Spider-Man clones he created, only one was a perfect copy of the original. He also created two clones of himself, one a direct copy, the other a modified clone harboring the [[Carrion (comics)|Carrion]] virus. His most prized creation however was a perfect copy of Gwen Stacy. Realizing the end of his plan was near, the Jackal went out against Spider-Man. With the assistance of the [[Tarantula (Marvel Comics)
It was much later revealed that the clone of Peter had survived the explosion and gone into hiding. The Jackal who died at Shea stadium was also a clone. Many months later, another Jackal clone would marry the Gwen Stacy clone and they would live under the assumed names Warren and Gwen Miles. This clone of Warren died of the clone degeneration that afflicted most of the clones created by the Jackal. Another
The Peter Parker clone, unaware that the Jackal is alive, takes the name [[Ben Reilly]], and goes into exile. He returns years later to [[New York City]], allied himself with Spider-Man, and became the [[Scarlet Spider]]. The Jackal returned at this point <ref>''Amazing Spider-Man'' #399, March, 1995</ref> and convinced both Parker and Reilly that Reilly was the real Peter Parker and that the other man was the clone, and created a number of other Peter Parker clones who came into conflict with Spider-Man, the Scarlet Spider, and [[Kaine]]. Ultimately, the Jackal, in the process of attempting to kill and replace millions of people with clones he could control, was killed falling off a tall building while trying to save the Gwen clone. <ref>''Maximum Clonage Omega'', July, 1995</ref>
However, another clone of Miles Warren has appeared, who was even further genetically altered to the point where he frequently displays animalistic tendencies and his body is always cold, causing him to wear a thick fur coat even in the hottest weather. He became a crime-lord calling himself "The Professor" and allied himself with
Near the end of the Clone Saga it was revealed that the Jackal and the other major players of the Clone Saga had unknowingly been duped by Norman Osborn
It was also revealed that Miles had previously been an assistant of the [[High Evolutionary]]. He married a woman named Monica who bore him two small children and that they were killed in what was originally believed to be a car crash. But
Miles had a brother who was a science teacher at Peter's High School (revealed in ''Untold Tales of Spider-Man'') named Aaron.
==Other versions==
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In Marvel Zombies Universe, when the Zombie Galacti left the Earth (after eating Galactus), Wilson Fisk ([[Kingpin (comics)|Kingpin]]) makes an empire. The zombiefied Jackal plays an important part in it, creating human clones to feed the remaining Marvel Zombies with Inhuman technology.<ref>''Marvel Zombies 3'' #2</ref>
==
===Television===
* Miles Warren appears in the two-part ''[[Spider-Man
* Miles Warren appears in ''[[The Spectacular Spider-Man (TV series)|The Spectacular Spider-Man]]'' episode "Blueprints" voiced by [[Brian George]]. Miles, along with his assistant [[Debra Whitman]], joins his brother Aaron, [[Lizard (comics)|Dr. Curt Connors]], and Dr. Martha Connors in scientific research at the ESU labs with a grant from [[Green Goblin|Norman Osborn]]. Warren bases his genetic research from Curt's work with lizard DNA, but
===Video games===
* Jackal appears in the Playstation 2 and PSP
==References==
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{{Spider-Man}}
[[Category:1965 comics characters debuts]]
[[Category:Fictional canines]]
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