Eli Greenbaum

Eli Greenbaum, U.S. herpetologist.

• Associate Professor of Evolutionary Genetics in the Department of Biological Sciences at UTEP, Texas, United States
• Website

Taxon names authored

(List may be incomplete)

• 31 taxon names authored by Eli Greenbaum

Publications

(List may be incomplete)

2001

• Greenbaum, E., and J. L. Carr., 2001: Sexual differentiation in the spiny softshell turtle (Apalone spinifera), a species with genetic sex determination. Journal of Experimental Zoology 290:190–200.

2002

• Greenbaum, E., 2002: A standardized series of embryonic stages for the emydid turtle Trachemys scripta. Canadian Journal of Zoology 80:1150–1170.
• Greenbaum, E., and J. L. Carr., 2002: Staging criteria for embryos of the spiny softshell turtle, Apalone spinifera (Testudines: Trionychidae). Journal of Morphology 254:272–291.

2003

• Greenbaum, E., N. Galeva, and M. Jorgensen., 2003: Venom variation and chemoreception of the viperid Agkistrodon contortrix: evidence for adaptation? Journal of Chemical Ecology 29:1741–1755.

2004

• Greenbaum, E., J. L. Carr, and A. Almendáriz., 2004: Taxonomic status of Tantilla equatoriana Wilson and Mena 1980 (Serpentes: Colubridae). The Southwestern Naturalist 49:457–464.
• Greenbaum, E., 2004: A new species of Bolitoglossa (Amphibia: Caudata: Plethodontidae) from montane forests in Guatemala and El Salvador. Journal of Herpetology 38:411–421.
• Greenbaum, E., and M. Jorgensen., 2004: Envenomated-invertebrate prey preference of Agkistrodon contortrix (Serpentes: Viperidae) during strike-induced chemosensory searching. Amphibia-Reptilia 25:165–172.
• Greenbaum, E., 2004: The influence of prey-scent stimuli on predatory behavior of the North American copperhead Agkistrodon contortrix (Serpentes: Viperidae). Behavioral Ecology 15:345–350.

2005

• Greenbaum, E., and O. Komar., 2005: Threat assessment and conservation prioritization of the herpetofauna of El Salvador. Biodiversity and Conservation 14:2377–2395.
• Greenbaum, E., 2005: Systematics of West African skinks in the Chalcides thierryi group: composition, distribution, and redescription of types. African Journal of Herpetology 54:17–29.
• Greenbaum, E., and J. L. Carr., 2005: The herpetofauna of Upper Niger National Park, Guinea, West Africa. Scientific Papers, Natural History Museum, The University of Kansas 37:1–21.
• Sheil, C. A., and E. Greenbaum., 2005: Reconsideration of skeletal development of Chelydra serpentina (Reptilia: Testudinata: Chelydridae): evidence for intraspecific variation. Journal of Zoology, London 265:235–267.

2006

• Bauer, A. M., T. Jackman, E. Greenbaum, and T. J. Papenfuss., 2006: Confirmation of the occurrence of Hemidactylus robustus Heyden, 1827 (Reptilia: Gekkonidae) in Iran and Pakistan. Zoology in the Middle East 39:59–62.
• Greenbaum, E., A. C. Campbell, and C. J. Raxworthy., 2006: A revision of sub-Saharan Chalcides (Squamata: Scincidae), with redescriptions of two East African species. Herpetologica 62:71–89.

2007

• Greenbaum, E., A. M. Bauer, and T. R. Jackman., 2007: Homopholis and Blaesodactylus (Squamata: Gekkonidae) revisited: new insights from a molecular phylogeny. African Journal of Herpetology 56:101–114.
• Jackman, T. R., A. M. Bauer, E. Greenbaum, M. Vences, and F. Glaw., 2007: Molecular phylogenetic relationships among species of the Malagasy-Comoran gecko genus Paroedura (Sauria: Gekkonidae). Molecular Phylogenetics and Evolution 46:74–81.
• Greenbaum, E., A. M. Bauer, T. R. Jackman, M. Vences, and F. Glaw., 2007: A phylogeny of the enigmatic Madagascan geckos of the genus Uroplatus (Sauria: Gekkonidae). Zootaxa 1493:41–51.
• Pramuk, J. B., M. J. Grose, A. L. Clarke, E. Greenbaum, E. Bonaccorso, J. M. Guayasamin, A. H. Smith, B. W. Benz, B. R. Harris, E. Siegfreid, Y. R. Reid, N. Holcroft, and E. O. Wiley., 2007: Phylogeny of finescale shiners of the genus Lythrurus (Cypriniformes: Cyprinidae) inferred from four mitochondrial genes. Molecular Phylogenetics and Evolution 42:287–297.
• Bauer, A. M., T. R. Jackman, E. Greenbaum, and T. J. Papenfuss., 2007: First record of Lepidodactylus lugubris in Suriname. Applied Herpetology 4:84–85.
• Herrera, N., V. Henríquez, and E. Greenbaum., 2007: New department and country records for amphibians and reptiles from El Salvador. Herpetological Review 38:222–226.
• Bauer, A. M., A. De Silva, E. Greenbaum, and T. R. Jackman., 2007: A new species of day gecko from high elevation in Sri Lanka, with a preliminary phylogeny of Sri Lankan Cnemaspis (Reptilia: Squamata: Gekkonidae). Mitteilungen aus dem Museum für Naturkunde in Berlin. Zoologische Reihe 83 (Sonderheft):22–32.

2008

• Bauer, A. M., V. B. Giri, E. Greenbaum, T. R. Jackman, M. S. Dharne, and Y. S. Shouche., 2008: On the systematics of the gekkonid genus Teratolepis Günther, 1869: another one bites the dust. Hamadryad 33:13–27.
• Jackman, T. R., A. M. Bauer, and E. Greenbaum., 2008: Phylogenetic relationships of geckos of the genus Nactus and their relatives (Squamata: Gekkonidae). Acta Herpetologica 3:1–18.
• Simmons, J. E., A. C. Campbell, and E. Greenbaum., 2008: Presence of elemental mercury in fluid preserved specimens. Collection Forum 22:32–36.
• McCranie, J. R., and E. Greenbaum., 2008: Bolitoglossa heiroreias Greenbaum. Catalogue of American Amphibians and Reptiles 842:1–3.
• Gamble, T., Bauer, A.M., Greenbaum, E. & Jackman, T.R. 2008. Evidence for Gondwanan vicariance in an ancient clade of gecko lizards. Journal of Biogeography 35(1): 88–104. DOI: 10.1111/j.1365-2699.2007.01770.x Reference page. 
• Gamble, T., Simons, A.M., Colli, G.R. & Vitt, L.J. 2008. Tertiary climate change and the diversification of the Amazonian gecko genus Gonatodes (Sphaerodactylidae, Squamata). Molecular Phylogenetics and Evolution 46(1): 269–277. DOI: 10.1016/j.ympev.2007.08.013 Reference page. 
• Jackman, T.R., Bauer, A.M., Greenbaum, E., Glaw, F. & Vences, M. 2008. Molecular phylogenetic relationships among species of the Malagasy-Comoran gecko genus Paroedura (Squamata: Gekkonidae). Molecular Phylogenetics and Evolution 46(1): 74–81. DOI: 10.1016/j.ympev.2007.10.018 Reference page. 
• Greenbaum, E., C. Kusamba, M. Aristote, and K. D. Reed., 2008: Amphibian chytrid fungus infections in Hyperolius (Anura: Hyperoliidae) from eastern Democratic Republic of Congo. Herpetological Review 39:70–73.

2010

• Heinicke, M.P., Greenbaum, E., Jackman, T.R. & Bauer, A.M. 2010. Molecular Phylogenetics of Pacific Nactus (Squamata: Gekkota: Gekkonidae)and the Diphyly of Australian Species. Proceedings of the California Academy of Sciences. Series 4 61(17): 633–646. BHL Reference page. 
• Bauer, A. M., T. R. Jackman, E. Greenbaum, A. de Silva, V. B. Giri, and I. Das., 2010: Molecular evidence for the taxonomic status of Hemidactylus brookii group taxa (Squamata: Gekkonidae). Herpetological Journal 20:129–138.
• Bauer, A. M., T. R. Jackman, E. Greenbaum, V. B. Giri, and A. de Silva., 2010: South Asia supports a major endemic radiation of Hemidactylus geckos. Molecular Phylogenetics and Evolution 57:343–352.

2011

• Sinsch, U., E. Greenbaum, C. Kusamba and E. Lehr. 2011: Rapid assessment of montane anuran communities in the Albertine Rift: Hyperolius castaneus Ahl, 1931 as a focal species for conservation. African Zoology 46:320–333.
• Greenbaum, E., Villanueva, C.O., Kusamba, C,, Aristote, M.M. & Branch, W.R. 2011. A molecular phylogeny of Equatorial African Lacertidae, with the description of a new genus and species from eastern Democratic Republic of the Congo. Zoological Journal of the Linnean Society 163(3): 913–942. DOI: 10.1111/j.1096-3642.2011.00732.x Reference page. 
• Heinicke, M. P., E. Greenbaum, T. Jackman, and A. M. Bauer. 2011: Phylogeny of a trans-Wallacean radiation (Squamata, Gekkonidae, Gehyra) supports a single early colonization of Australia. Zoologica Scripta 40:584–602.
• Roelke, C. E., E. Greenbaum, C. Kusamba, M. M. Aristote, and E. N. Smith. 2011: Systematics and conservation status of two distinct Albertine Rift treefrogs, Leptopelis karissimbensis and Leptopelis kivuensis (Anura: Arthroleptidae). Journal of Herpetology 45:343–351.
• Greenbaum, E., E. N. Smith, and R. O. de Sá. 2011: Molecular systematics of the Middle American genus Hypopachus (Anura: Microhylidae). Molecular Phylogenetics and Evolution 61:265–277.
• Rösler, H., Bauer, A.M., Heinicke, M.P., Greenbaum, E., Jackman, T., Nguyen, T.Q. & Ziegler, T. 2011. Phylogeny, taxonomy, and zoogeography of the genus Gekko Laurenti, 1768 with the revalidation of G. reevesii Gray, 1831 (Sauria: Gekkonidae). Zootaxa 2989(1): 1–50. DOI: 10.11646/zootaxa.2989.1.1 Reference page. 
• Gamble, T., Bauer, A.M., Colli, G.R., Greenbaum, E., Jackman, T.R., Vitt, L.J. & Simons, A.M. 2011. Coming to America: multiple origins of New World geckos. Journal of Evolutionary Biology 24(2): 231–244. DOI: 10.1111/j.1420-9101.2010.02184.x Reference page. 
• Evans, B. J., E. Greenbaum, C. Kusamba, T. F. Carter, M. L. Tobias, S. A. Mendel, and D. B. Kelley. 2011: Description of a new octoploid frog species (Anura: Pipidae: Xenopus) from the Democratic Republic of the Congo, with a discussion of the biogeography of African clawed frogs in the Albertine Rift. Journal of Zoology, London 283: 276–290.

2012

• Eli Greenbaum, Frank Portillo, Chifundera Kusamba, 2012: Systematics of the poorly known treefrog Leptopelis fiziensis (Anura: Arthroleptidae), with a description of its call. African journal of herpetology, 61(2), 113-127.
• Heinicke, M. P., E. Greenbaum, T. R. Jackman, and A. M. Bauer. 2012: Evolution of gliding in Southeast Asian geckos and other vertebrates is temporally congruent with dipterocarp forest development. Biology Letters 8:994–997.
• de Sá, R. O., J. W. Streicher, R. Selenkoyela, M. C. Forlani, S. P. Loader, E. Greenbaum, S. Richards, and C. F. B. Haddad. 2012: Molecular phylogeny of microhylid frogs (Anura: Microhylidae) with emphasis on relationships among New World genera. BMC Evolutionary Biology 12:241 (21 pp).
• Anderson*, C. G., and E. Greenbaum., 2012: Phylogeography of northern populations of the black-tailed rattlesnake (Crotalus molossus Baird and Girard, 1853), with the revalidation of C. ornatus Hallowell, 1854. Herpetological Monographs 26:19–57.
• Greenbaum, E., and C. Kusamba., 2012: Conservation implications following the rediscovery of four frog species from the Itombwe Natural Reserve, eastern Democratic Republic of the Congo. Herpetological Review 43:253–259.
• Gamble, T., E. Greenbaum, T. R. Jackman, A. P. Russell, and A. M. Bauer. 2012: Repeated evolution of adhesive toepads in geckos. PLoS ONE 7:e39429.
• Wagner, P., E. Greenbaum, and A. M. Bauer. 2012: A new species of the Acanthocercus atricollis (Squamata: Agamidae) complex from Zambia. Salamandra 48:21–30.
• Greenbaum, E., Stanley, E.L., Kusamba, C., Moninga, W.M., Goldberg, S.R. & Bursey, C.R. 2012. A new species of Cordylus (Squamata: Cordylidae) from the Marungu Plateau of south-eastern Democratic Republic of the Congo. African Journal of Herpetology 61(1): 14–39. DOI: 10.1080/21564574.2012.666505 Reference page. 
• Oliver, P.M., Bauer, A.M., Greenbaum, E., Jackman, T. & Hobbie, T. 2012. Molecular phylogenetics of the arboreal Australian gecko genus Oedura Gray 1842 (Gekkota: Diplodactylidae): Another plesiomorphic grade? Molecular Phylogenetics and Evolution 63(2): 255–264. DOI: 10.1016/j.ympev.2011.12.013 Reference page. 
• Greenbaum, E., K. A. Tolley, A. Joma*, and C. Kusamba. 2012: A new species of chameleon (Sauria: Chamaeleonidae: Kinyongia) from the northern Albertine Rift, Central Africa. Herpetologica 68:60–75.

2013

• Greenbaum, E., U. Sinsch, E. Lehr, F. Valdez, and C. Kusamba. 2013: Phylogeography of the reed frog Hyperolius castaneus (Anura: Hyperoliidae) from the Albertine Rift of Central Africa: Implications for taxonomy, biogeography and conservation. Zootaxa 3731: 473–494.

2014

• Frank Portillo, Eli Greenbaum, 2014: At the Edge of a Species Boundary: A New and Relatively Young Species of Leptopelis (Anura: Arthroleptidae) from the Itombwe Plateau, Democratic Republic of the Congo. Herpetologica, 70(1), 100-119 (2014)
• Portillo, F.; Greenbaum, E.. A New Species of the Leptopelis modestus Complex (Anura: Arthroleptidae) from the Albertine Rift of Central Africa. Journal of herpetology, 48(3): 394–406. DOI: 10.1670/13-043
• Leaché, A. D., P. Wagner, C. W. Linkem, W. Böhme, T. J. Papenfuss, R. Chong, B. R. Lavin, A. M. Bauer, S. Nielsen, E. Greenbaum, M. -O. Rödel, A. Schmitz, M. LeBreton, I. Ineich, L. Chirio, E. A. Eniang, S. Baha El Din, A. R. Lemmon, and F. T. Burbrink., 2014: Sexual selection drives adaptive radiation in African lizards. Molecular Phylogenetics and Evolution, 79: 215–230.
• Leaché, A.D., Wagner, P., Linkem, C.W., Böhme, W., Papenfuss, T.J., Chong, R.A., Lavin, B.R., Bauer, A.M., Nielsen, S.V., Greenbaum, E., Rödel, M.O., Schmitz, A., LeBreton, M., Ineich, I., Chirio, L., Ofori-Boateng, C., Eniang, E.A., El Din, S.B., Lemmon, A.R. & Burbrink, F.T. 2014. A hybrid phylogenetic-phylogenomic approach for species tree estimation in African Agama lizards with applications to biogeography, character evolution, and diversification. Molecular Phylogenetics and Evolution 79: 215–230. DOI: 10.1016/j.ympev.2014.06.013 Reference page. 
• Wagner, P., E. Greenbaum, P. Malonza, and W. R. Branch. 2014: Resolving sky island speciation in populations of East African Adolfus alleni (Sauria, Lacertidae). Salamandra 50:1–17.
• Heinicke, M.P., Daza, J.D., Greenbaum, E., Jackman, T.R. & Bauer, A.M. 2014. Phylogeny, taxonomy and biogeography of a circum-Indian Ocean clade of leaf-toed geckos (Reptilia: Gekkota), with a description of two new genera. Systematics and Biodiversity 12(1): 23–42. DOI: 10.1080/14772000.2013.877999 Reference page. 

2015

• Greenbaum, E., Portillo, F., Jackson, K. & Kusamba, C. 2015. A phylogeny of Central African Boaedon (Serpentes: Lamprophiidae), with the description of a new cryptic species from the Albertine Rift. African Journal of Herpetology 64(1): 18–38. DOI: 10.1080/21564574.2014.996189 Reference page. 
• Lehr, E., Dehling, J.M., Greenbaum, E. & Sinsch, U. 2015. Embryogenesis and tadpole description of Hyperolius castaneus Ahl, 1931 and H. jackie Dehling, 2012 (Anura, Hyperoliidae) from montane bog pools. ZooKeys 546: 125–152. DOI: 10.3897/zookeys.546.6044 Reference page. 

2016

• Blackburn, D.C., Boix, C., Greenbaum, E., Fabrezi, M., Meirte, D., Plumptre, A.J. & Stanley, E.L. 2016. The distribution of the Bururi Long-fingered Frog (Cardioglossa cyaneospila, family Arthroleptidae), a poorly known Albertine Rift endemic. Zootaxa 4170(2): 355–364. DOI: 10.11646/zootaxa.4170.2.8. Reference page. 
• Medina, M.F., Bauer, A.M., Branch, W.R., Schmitz, A., Conradie, W., Nagy, Z.T., Hibbitts, T.J., Ernst, R., Portik, D.M., Nielsen, S.V., Colston, T.J., Kusamba, C., Behangana, M., Rödel, M.-O. & Greenbaum, E. 2016. Molecular phylogeny of Panaspis and Afroablepharus skinks (Squamata: Scincidae) in the savannas of sub-Saharan Africa. Molecular Phylogenetics and Evolution 100: 409–423. DOI: 10.1016/j.ympev.2016.04.026 Reference page. 

2017

• Agarwal, I., Biswas, S., Bauer, A.M., Greenbaum, E., Jackman, T.R., De Silva, A. & Batuwita, S. 2017. Cryptic species, taxonomic inflation, or a bit of both? New species phenomenon in Sri Lanka as suggested by a phylogeny of dwarf geckos (Reptilia, Squamata, Gekkonidae, Cnemaspis). Systematics and Biodiversity 15(5): 427–439. DOI: 10.1080/14772000.2017.1282553 Reference page. 
• Hughes, D.F., Kusamba, C., Behangana, M. & Greenbaum, E. 2017. Integrative taxonomy of the Central African forest chameleon, Kinyongia adolfifriderici (Sauria: Chamaeleonidae), reveals underestimated species diversity in the Albertine Rift. Zoological Journal of the Linnean Society 181(2): 400–438. DOI: 10.1093/zoolinnean/zlx005. Reference page. 
• Leaché, A.D., Grummer, J.A., Miller, M., Krishnan, S., Fujita, M.K., Böhme, W., Schmitz, A., LeBreton, M., Ineich, I., Chirio, L., Ofori-Boateng, C., Eniang, E.A., Greenbaum, E., Rödel, M.O. & Wagner, P. 2016. Bayesian inference of species diffusion in the West African Agama agama species group (Reptilia, Agamidae). Systematics and Biodiversity 15(3): 192–203. DOI: 10.1080/14772000.2016.1238018 Reference page. 

2018

• Broadley, D.G.†, Tolley, K.A., Conradie, W., Wishart, S., Trape, J.-F., Burger, M., Kusamba, C., Zassi-Boulou, A.-G. & Greenbaum, E. 2018. A phylogeny and genus-level revision of the African file snakes Gonionotophis Boulenger (Squamata: Lamprophiidae). African Journal of Herpetology 67(1): 43–60. DOI: 10.1080/21564574.2018.1423578 Reference page. 
• Greenbaum, E., Beer, S.D., Hughes, D.F., Wagner, P., Anderson, C.G., Villanueva, C.O., Malonza, P.K., Kusamba, C., Muninga, W.M., Aristote, M.M. & Branch, W.R. 2018. Phylogeography of Jackson's Forest Lizard Adolfus jacksoni (Sauria: Lacertidae) Reveals Cryptic Diversity in the Highlands of East Africa. Herpetological Monographs 32(1): 51–68. DOI: 10.1655/HERPMONOGRAPHS-D-18-00005.1 Reference page. 
• Wüster, W., Chirio, L., Trape, J.-F., Ineich, I., Jackson, K., Greenbaum, E., Barron, C., Kusamba, C., Nagy, Z.T., Storey, R., Hall, C., Wüster, C.E., Barlow, A. & Broadley, D.G.† 2018. Integration of nuclear and mitochondrial gene sequences and morphology reveals unexpected diversity in the forest cobra (Naja melanoleuca) species complex in Central and West Africa (Serpentes: Elapidae). Zootaxa 4455(1): 68–98. DOI: 10.11646/zootaxa.4455.1.3 Reference page. 

2019

• Keates, C., Conradie, W., Greenbaum, E. & Edwards, S. 2019. A snake in the grass: Genetic structuring of the widespread African grass snake (Psammophylax Fitzinger 1843), with the description of a new genus and a new species. Journal of Zoological Systematics and Evolutionary Research 57(4): 1039–1066. DOI: 10.1111/jzs.12337 Reference page. 
• Portillo, F., Branch, W.R., Tilbury, C.R., Nagy, Z.T., Hughes, D.F., Kusamba, C., Muninga, W.M., Aristote, M.M., Behangana, M. & Greenbaum, E. 2019. A Cryptic New Species of Polemon (Squamata: Lamprophiidae, Aparallactinae) from the Miombo Woodlands of Central and East Africa. Copeia 107(1): 22–36. DOI: 10.1643/CH-18-098 Reference page. 

2020

• Engelbrecht, H.M., Branch, W.R., Greenbaum, E., Burger, M., Conradie, W. & Tolley, K.A. 2020. African Herald snakes, Crotaphopeltis, show population structure for a widespread generalist but deep genetic divergence for forest specialists. Journal of Zoological Systematics and Evolutionary Research 58(4): 1220–1233. DOI: 10.1111/jzs.12361 Reference page. 
• Greenwood, L., Loader, S.P., Lawson, L.P., Greenbaum, E. & Zimkus, B.M. 2020. A new species of Phrynobatrachus (Amphibia: Anura: Phrynobatrachidae) from the northern mountains of Tanzania. Journal of Natural History 54(1–4): 63–85. DOI: 10.1080/00222933.2020.1757171 Reference page. 

2021

• Ceríaco, L.M.P., Bauer, A.M., Kusamba, C., Agarwal, I. & Greenbaum, E. 2021. A New Species of Ground-Dwelling Hemidactylus (Squamata: Gekkonidae) from Southwestern Democratic Republic of the Congo. Journal of Herpetology 55(2): 105–111. DOI: 10.1670/20-094 Reference page. 
• Greenbaum, E., Allen, K.E., Vaughan, E.R., Pauwels, O.S.G., Wallach, V., Kusamba, C., Muninga, W.M., Aristote, M.M., Mali, F.M.M., Badjedjea, G., Penner, J., Rödel, M.-O., Rivera, J., Sterkhova, V., Johnson, G., Tapondjou, W.P. & Brown, R.B. 2021. Night stalkers from above: A monograph of Toxicodryas tree snakes (Squamata: Colubridae) with descriptions of two new cryptic species from Central Africa. Zootaxa 4965(1): 1–44. DOI: 10.11646/zootaxa.4965.1.1 Reference page. 

2023

• Greenbaum, E., Pauwels, O.S.G., Gvoždík, V., Vaughan, E.R., Chaney, T., Buontempo, M., Aristote, M.M., Muninga, W.M. & Engelbrecht, H.M. 2023. Systematics of the Thirteen-scaled Green Snake Philothamnus carinatus (Squamata: Colubridae), with the description of a cryptic new species from Central and East Africa. African Journal of Herpetology 72(2): 119–144. DOI: 10.1080/21564574.2023.2245840 Reference page. 

2024

• Das, S., Greenbaum, E., Brecko, J., Pauwels, O.S.G., Ruane, S., Pirro, S. & Merilä, J. 2024. Phylogenomics of Psammodynastes and Buhoma (Elapoidea: Serpentes), with the description of a new Asian snake family. Scientific Reports 14: 9489. DOI: 10.1038/s41598-024-60215-2 Reference page. 
• Hughes, D.F., Behangana, M., Lukwago, W., Menegon, M., Dehling, J.M., Wagner, P., Tilbury, C.R., South, T., Kusamba, C. & Greenbaum, E. 2024. Taxonomy of the Rhampholeon boulengeri Complex (Sauria: Chamaeleonidae): Five New Species from Central Africa’s Albertine Rift. Zootaxa 5458(4): 451–494. DOI: 10.11646/zootaxa.5458.4.1 Reference page.