Although Nayarit has been considered a "well-known" Mexican state (GONZALEZ SORIANO & NOVELO GUTI... more Although Nayarit has been considered a "well-known" Mexican state (GONZALEZ SORIANO & NOVELO GUTIERREZ, 1996), almost the entire published record from the state consists of records from the 19th century (CALVERT, 1899, 1901-08). Only a few subsequent papers have mentioned specimens from Nayarit (BELLE, 1987;
is the only species of Odonata presently known to be restricted to the Columbia River Basin and i... more is the only species of Odonata presently known to be restricted to the Columbia River Basin and is the only member of the subgenus Gomphurus known from west of the Rocky Mountains. It is a dark-colored species related to G. exrernus but differing from that species and other Gomphurus in numerous structural and color-pattern features. It is peculiar in having abundant pruinosity on the head, thorax, legs and base of abdomen and is perhaps the most pruinose gomphid. Relationships among the species of Gomphurus are discussed and species pairs suggested. G. lynnae inhabits a large, rocky to muddy river in sagebrush steppe and is known to fly from 2 June to 18 July. It may be theatened because of its restricted range and the considerable human alteration of rivers in the Columbia Basin. 60 D.R. Paulson I examined specimens of all the North American species of this subgenus except ozarkensis Westfall and ventricosus Walsh, which are well described in the literature (WESTFALL, 1975; WALKER, 1958). I have discounted 'Gomphurus' adelphus Selys entirely in all the following discussions, as it is now known that it is not a valid species (M.J. Westfall, Jr., pers. comm.), and it and all Old World species are excluded when 1 refer to 'all Gomphurus.' GOMPHUS (GOMPHURUS) LYNNAE SP.N. Figures 1 a-f Material (all from Benton County, Washington).
Leptobasis guanacaste is described from seasonal wetlands in dry forest in Guanacaste, Costa Rica... more Leptobasis guanacaste is described from seasonal wetlands in dry forest in Guanacaste, Costa Rica. It is unique among the five species of the genus in thoracic color pattern and the structure of the male terminal appendages and female mesostigmal laminae and appears to be closest to L. candelaria through similarities in genital ligula, male metafemur, and female ovipositor.
Seventeen species of Odonata were collected in 1994-1996 from 21 localities in the Kuril Islands.... more Seventeen species of Odonata were collected in 1994-1996 from 21 localities in the Kuril Islands. Mnais pruinosa, Aeshna nigroflava, Cordulia aenea, and Pseudothemis zonata are first records from the archipelago, and the last species represents a significant range extension from Honshu. Enallagma belyshevi is synonymized with E. circulatum, which is considered a valid species rather than a subspecies of E. boreale.
... 101 ISCHNURA PERPARVA MCLACHLAN (ZYGOPTERA: COENAGRIONIDAE) HAS AN ANDROMORPHIC FEMALE, AND A... more ... 101 ISCHNURA PERPARVA MCLACHLAN (ZYGOPTERA: COENAGRIONIDAE) HAS AN ANDROMORPHIC FEMALE, AND ANOTHER SUGGESTION TO MODIFY THE TERMINOLOGY OF ... females were collected in Washington state, from a pond 0.5 mi E Beverly, Grant Co., 5 ...
The sibling species Orthemis discolor and 0. ferruginea are distinguished by coloration, although... more The sibling species Orthemis discolor and 0. ferruginea are distinguished by coloration, although one structural characteristic can be used to differentiate many females. Further information on the distribution and relative abundance of the two species in North and Middle America is presented.
ABSTRACT Herein I respond to a critique of my paper on wing positions in Zygoptera. The author of... more ABSTRACT Herein I respond to a critique of my paper on wing positions in Zygoptera. The author of that critique suggested that most of the hypotheses presented in that paper were flawed and questioned some of the facts brought to bear on them. In addition, he presented his own ideas in support of hypotheses I had rejected. I take this opportunity to clarify my reasoning further. Although I did not elaborate sufficiently in some cases, no statement made in my paper was incorrect. My critic and I are in agreement that this is a complicated matter, and all hypotheses continue to be worth further testing.
ABSTRACT This report summarizes progress that has been made during the past five years toward the... more ABSTRACT This report summarizes progress that has been made during the past five years toward the understanding of Neotropical Odonata. It also presents a list of critical species and sites, threats to Odonata conservation in the region, and priorities for further research. This region, the richest in the world for Odonata, must be a focus of intense research and conservation efforts.
Abstract Zygoptera show two perching modes, one with wings closed and one with wings open. These ... more Abstract Zygoptera show two perching modes, one with wings closed and one with wings open. These perching modes are distributed unequally through the suborder; most Zygoptera perch with closed wings, but species in 43 genera of eight families at least occasionally - in most cases usually - perch with open wings. Alternative hypotheses to explain this dichotomy are assessed. The dichotomy does not seem to be explicable by the Phylogenetic Inertia Hypothesis (PIH), the Wing Display Hypothesis (WDH), or the Thermoregulation Hypothesis (TH). I propose a hypothesis that the openwing position used by some zygopterans facilitates either more rapid takeoff or quicker orientation toward flying prey: the Quick Takeoff Hypothesis (QTH). That openwing species usually take flying prey furnishes support for the QTH, although many closedwing species also take flying prey. However, as most zygopterans perch with closed wings, that behavior needs explanation too, and I propose a hypothesis that perching with wings spread may make a zygopteran more conspicuous to predators and thus may be disadvantageous: the Shiny Wing Hypothesis (SWH). Larger species are less at risk of predation than smaller species, open wings in shade should be less conspicuous than in sunlight, and the majority of zygopterans with open wings are large tropical shade perchers, furnishing support for the SWH.
The accuracy of the Content should not be relied upon and should be independently verified with p... more The accuracy of the Content should not be relied upon and should be independently verified with primary sources of information. Taylor and Francis shall not be liable for any losses, actions, claims, proceedings, demands, costs, expenses, damages, and other liabilities whatsoever or howsoever caused arising directly or indirectly in connection with, in relation to or arising out of the use of the Content. This article may be used for research, teaching, and private study purposes. Any substantial or systematic reproduction, redistribution, reselling, loan, sub-licensing, systematic supply, or distribution in any form to anyone is expressly forbidden. Terms &
A recent collection of 69 specimens together with survey counts and incidental observations durin... more A recent collection of 69 specimens together with survey counts and incidental observations during June-July 2002 provide new information on the odonate fauna of Kosrae, Micronesia. The fauna comprises one zygopteran (Isehnura aurora) and six anisopterans. It appears to have remained stable with no known extinctions or colonizations over the past half century. The fauna is nearly a subset of that of Pohnpei and the islands to the west, and it comprises six widespread weedy species and one endemic, Hemieordulia erieo. Upland aquatic habitats appear largely unexploited or underutilized by odonates, and the absence of any Teinobasis species on Kosrae is in marked contrast to the presence of six species on the nearest high island, Pohnpei.
Fifteen species of Odonata are recorded from Yap, Micronesia-two Zygoptera and 13 Anisoptera. Non... more Fifteen species of Odonata are recorded from Yap, Micronesia-two Zygoptera and 13 Anisoptera. None is endemic to Yap. Hemicordulia lulico occurs elsewhere only in Palau, whereas most of the other species are widespread in the western Pacific and Indo-Australian regions. Macrodiplax cora and Tramea loewi, both recorded by Lieftinck in 1962, were the only species not encountered during this study; Tramea loewi remains known in Micronesia only from a single male collected in Yap by R. J. Goss in 1950. Six of the breeding species in Yap that are widespread in Indo-Australia occur no farther east in the Caroline Islands except possibly as unusual extralimital records in the cases of Agriocnemis femina and Neurothemis terminata; the four other species reaching only as far east as Yap are Anaciaeschna jaspidea, Agrionoptera insignis, Orthetrum serapia, and Rhyothemis phyllis. Orthetrum serapia is reported from Micronesia for the first time, although a very old single specimen record of O. sabina from Tobi Island may possibly pertain to O. serapia. The odonate fauna of the outer islands of Yap State is poorly known; only six species have been recorded from among four of the 15 island groups. In addition, Tramea transmarina euryale rather than T. t. propinqua was found to be the subspecies occurring in the Chuuk Islands, contrary to earlier publications.
... m. We were surprised by the proximity of the sandpipers to the plovers, because there seemed ... more ... m. We were surprised by the proximity of the sandpipers to the plovers, because there seemed to be many similar areas of tundra that could ... Lesser Golden-Plovers (Pluvi-alis dominica) were less likely to attack avian predators; from the five pairs in our study area, we observed ...
Page 1. SHORT COMMUNICATIONS 99 Condor 85:99-101 ? The Cooper Ornithological Society 1983 FLEDGIN... more Page 1. SHORT COMMUNICATIONS 99 Condor 85:99-101 ? The Cooper Ornithological Society 1983 FLEDGING DATES AND SOUTHWARD MIGRATION OF JUVENILES OF SOME CALIDRIS SANDPIPERS DENNIS R. PAULSON ...
This article appeared in a journal published by Elsevier. The attached copy is furnished to the a... more This article appeared in a journal published by Elsevier. The attached copy is furnished to the author for internal non-commercial research and education use, including for instruction at the authors institution and sharing with colleagues. Other uses, including reproduction and distribution, or selling or licensing copies, or posting to personal, institutional or third party websites are prohibited. In most cases authors are permitted to post their version of the article (e.g. in Word or Tex form) to their personal website or institutional repository. Authors requiring further information regarding Elsevier's archiving and manuscript policies are encouraged to visit: http://www.elsevier.com/copyright
BioOne sees sustainable scholarly publishing as an inherently collaborative enterprise connecting... more BioOne sees sustainable scholarly publishing as an inherently collaborative enterprise connecting authors, nonprofit publishers, academic institutions, research libraries, and research funders in the common goal of maximizing access to critical research.
Although Nayarit has been considered a "well-known" Mexican state (GONZALEZ SORIANO & NOVELO GUTI... more Although Nayarit has been considered a "well-known" Mexican state (GONZALEZ SORIANO & NOVELO GUTIERREZ, 1996), almost the entire published record from the state consists of records from the 19th century (CALVERT, 1899, 1901-08). Only a few subsequent papers have mentioned specimens from Nayarit (BELLE, 1987;
is the only species of Odonata presently known to be restricted to the Columbia River Basin and i... more is the only species of Odonata presently known to be restricted to the Columbia River Basin and is the only member of the subgenus Gomphurus known from west of the Rocky Mountains. It is a dark-colored species related to G. exrernus but differing from that species and other Gomphurus in numerous structural and color-pattern features. It is peculiar in having abundant pruinosity on the head, thorax, legs and base of abdomen and is perhaps the most pruinose gomphid. Relationships among the species of Gomphurus are discussed and species pairs suggested. G. lynnae inhabits a large, rocky to muddy river in sagebrush steppe and is known to fly from 2 June to 18 July. It may be theatened because of its restricted range and the considerable human alteration of rivers in the Columbia Basin. 60 D.R. Paulson I examined specimens of all the North American species of this subgenus except ozarkensis Westfall and ventricosus Walsh, which are well described in the literature (WESTFALL, 1975; WALKER, 1958). I have discounted 'Gomphurus' adelphus Selys entirely in all the following discussions, as it is now known that it is not a valid species (M.J. Westfall, Jr., pers. comm.), and it and all Old World species are excluded when 1 refer to 'all Gomphurus.' GOMPHUS (GOMPHURUS) LYNNAE SP.N. Figures 1 a-f Material (all from Benton County, Washington).
Leptobasis guanacaste is described from seasonal wetlands in dry forest in Guanacaste, Costa Rica... more Leptobasis guanacaste is described from seasonal wetlands in dry forest in Guanacaste, Costa Rica. It is unique among the five species of the genus in thoracic color pattern and the structure of the male terminal appendages and female mesostigmal laminae and appears to be closest to L. candelaria through similarities in genital ligula, male metafemur, and female ovipositor.
Seventeen species of Odonata were collected in 1994-1996 from 21 localities in the Kuril Islands.... more Seventeen species of Odonata were collected in 1994-1996 from 21 localities in the Kuril Islands. Mnais pruinosa, Aeshna nigroflava, Cordulia aenea, and Pseudothemis zonata are first records from the archipelago, and the last species represents a significant range extension from Honshu. Enallagma belyshevi is synonymized with E. circulatum, which is considered a valid species rather than a subspecies of E. boreale.
... 101 ISCHNURA PERPARVA MCLACHLAN (ZYGOPTERA: COENAGRIONIDAE) HAS AN ANDROMORPHIC FEMALE, AND A... more ... 101 ISCHNURA PERPARVA MCLACHLAN (ZYGOPTERA: COENAGRIONIDAE) HAS AN ANDROMORPHIC FEMALE, AND ANOTHER SUGGESTION TO MODIFY THE TERMINOLOGY OF ... females were collected in Washington state, from a pond 0.5 mi E Beverly, Grant Co., 5 ...
The sibling species Orthemis discolor and 0. ferruginea are distinguished by coloration, although... more The sibling species Orthemis discolor and 0. ferruginea are distinguished by coloration, although one structural characteristic can be used to differentiate many females. Further information on the distribution and relative abundance of the two species in North and Middle America is presented.
ABSTRACT Herein I respond to a critique of my paper on wing positions in Zygoptera. The author of... more ABSTRACT Herein I respond to a critique of my paper on wing positions in Zygoptera. The author of that critique suggested that most of the hypotheses presented in that paper were flawed and questioned some of the facts brought to bear on them. In addition, he presented his own ideas in support of hypotheses I had rejected. I take this opportunity to clarify my reasoning further. Although I did not elaborate sufficiently in some cases, no statement made in my paper was incorrect. My critic and I are in agreement that this is a complicated matter, and all hypotheses continue to be worth further testing.
ABSTRACT This report summarizes progress that has been made during the past five years toward the... more ABSTRACT This report summarizes progress that has been made during the past five years toward the understanding of Neotropical Odonata. It also presents a list of critical species and sites, threats to Odonata conservation in the region, and priorities for further research. This region, the richest in the world for Odonata, must be a focus of intense research and conservation efforts.
Abstract Zygoptera show two perching modes, one with wings closed and one with wings open. These ... more Abstract Zygoptera show two perching modes, one with wings closed and one with wings open. These perching modes are distributed unequally through the suborder; most Zygoptera perch with closed wings, but species in 43 genera of eight families at least occasionally - in most cases usually - perch with open wings. Alternative hypotheses to explain this dichotomy are assessed. The dichotomy does not seem to be explicable by the Phylogenetic Inertia Hypothesis (PIH), the Wing Display Hypothesis (WDH), or the Thermoregulation Hypothesis (TH). I propose a hypothesis that the openwing position used by some zygopterans facilitates either more rapid takeoff or quicker orientation toward flying prey: the Quick Takeoff Hypothesis (QTH). That openwing species usually take flying prey furnishes support for the QTH, although many closedwing species also take flying prey. However, as most zygopterans perch with closed wings, that behavior needs explanation too, and I propose a hypothesis that perching with wings spread may make a zygopteran more conspicuous to predators and thus may be disadvantageous: the Shiny Wing Hypothesis (SWH). Larger species are less at risk of predation than smaller species, open wings in shade should be less conspicuous than in sunlight, and the majority of zygopterans with open wings are large tropical shade perchers, furnishing support for the SWH.
The accuracy of the Content should not be relied upon and should be independently verified with p... more The accuracy of the Content should not be relied upon and should be independently verified with primary sources of information. Taylor and Francis shall not be liable for any losses, actions, claims, proceedings, demands, costs, expenses, damages, and other liabilities whatsoever or howsoever caused arising directly or indirectly in connection with, in relation to or arising out of the use of the Content. This article may be used for research, teaching, and private study purposes. Any substantial or systematic reproduction, redistribution, reselling, loan, sub-licensing, systematic supply, or distribution in any form to anyone is expressly forbidden. Terms &
A recent collection of 69 specimens together with survey counts and incidental observations durin... more A recent collection of 69 specimens together with survey counts and incidental observations during June-July 2002 provide new information on the odonate fauna of Kosrae, Micronesia. The fauna comprises one zygopteran (Isehnura aurora) and six anisopterans. It appears to have remained stable with no known extinctions or colonizations over the past half century. The fauna is nearly a subset of that of Pohnpei and the islands to the west, and it comprises six widespread weedy species and one endemic, Hemieordulia erieo. Upland aquatic habitats appear largely unexploited or underutilized by odonates, and the absence of any Teinobasis species on Kosrae is in marked contrast to the presence of six species on the nearest high island, Pohnpei.
Fifteen species of Odonata are recorded from Yap, Micronesia-two Zygoptera and 13 Anisoptera. Non... more Fifteen species of Odonata are recorded from Yap, Micronesia-two Zygoptera and 13 Anisoptera. None is endemic to Yap. Hemicordulia lulico occurs elsewhere only in Palau, whereas most of the other species are widespread in the western Pacific and Indo-Australian regions. Macrodiplax cora and Tramea loewi, both recorded by Lieftinck in 1962, were the only species not encountered during this study; Tramea loewi remains known in Micronesia only from a single male collected in Yap by R. J. Goss in 1950. Six of the breeding species in Yap that are widespread in Indo-Australia occur no farther east in the Caroline Islands except possibly as unusual extralimital records in the cases of Agriocnemis femina and Neurothemis terminata; the four other species reaching only as far east as Yap are Anaciaeschna jaspidea, Agrionoptera insignis, Orthetrum serapia, and Rhyothemis phyllis. Orthetrum serapia is reported from Micronesia for the first time, although a very old single specimen record of O. sabina from Tobi Island may possibly pertain to O. serapia. The odonate fauna of the outer islands of Yap State is poorly known; only six species have been recorded from among four of the 15 island groups. In addition, Tramea transmarina euryale rather than T. t. propinqua was found to be the subspecies occurring in the Chuuk Islands, contrary to earlier publications.
... m. We were surprised by the proximity of the sandpipers to the plovers, because there seemed ... more ... m. We were surprised by the proximity of the sandpipers to the plovers, because there seemed to be many similar areas of tundra that could ... Lesser Golden-Plovers (Pluvi-alis dominica) were less likely to attack avian predators; from the five pairs in our study area, we observed ...
Page 1. SHORT COMMUNICATIONS 99 Condor 85:99-101 ? The Cooper Ornithological Society 1983 FLEDGIN... more Page 1. SHORT COMMUNICATIONS 99 Condor 85:99-101 ? The Cooper Ornithological Society 1983 FLEDGING DATES AND SOUTHWARD MIGRATION OF JUVENILES OF SOME CALIDRIS SANDPIPERS DENNIS R. PAULSON ...
This article appeared in a journal published by Elsevier. The attached copy is furnished to the a... more This article appeared in a journal published by Elsevier. The attached copy is furnished to the author for internal non-commercial research and education use, including for instruction at the authors institution and sharing with colleagues. Other uses, including reproduction and distribution, or selling or licensing copies, or posting to personal, institutional or third party websites are prohibited. In most cases authors are permitted to post their version of the article (e.g. in Word or Tex form) to their personal website or institutional repository. Authors requiring further information regarding Elsevier's archiving and manuscript policies are encouraged to visit: http://www.elsevier.com/copyright
BioOne sees sustainable scholarly publishing as an inherently collaborative enterprise connecting... more BioOne sees sustainable scholarly publishing as an inherently collaborative enterprise connecting authors, nonprofit publishers, academic institutions, research libraries, and research funders in the common goal of maximizing access to critical research.
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Papers by Dennis Paulson