Second Edition

Field Guide to

Stink Bugs
of Agricultural Importance
in the United States

Field Guide to Stink Bugs of Agricultural

Importance in the United States

Authors:
D. Ames Herbert Jr., Virginia Tech
Katherine Kamminga, Virginia Tech
Sean Malone, Virginia Tech
Thomas P. Kuhar, Virginia Tech
Eric Day, Virginia Tech
Jeremy Greene, Clemson University
C. Scott Bundy, New Mexico State University
Lydia Brown, University of Arizona
Peter Ellsworth, University of Arizona

Produced by:
Thea Glidden, Director, Communications and Marketing,
College of Agriculture and Life Sciences, Virginia Tech
Lori Greiner, Communications Manager, Virginia Cooperative
Extension, Virginia Tech
Tim FisherPoff, Art Director, Communications and Marketing,
College of Agriculture and Life Sciences, Virginia Tech
Bobbi A. Hoffman, Copy Editor, Communications and
Marketing, College of Agriculture and Life Sciences,
Virginia Tech

Supported by:
Virginia Integrated Pest Management Program
Virginia Cooperative Extension
Virginia Tech and Virginia State University
StopBMSB.org
USDA-NIFA: Regional IPM Centers

Table of Contents
Photo Credits................................................................................... iii
Stink Bug Diagram......................................................................... iv
Beaks of Predators versus Pests................................................. vi
Economically Important Species................................................. 1
Less Common Species.................................................................. 16
Beneficial Species......................................................................... 21
Stink Bug-Like Species.................................................................25
Feeding Injury...............................................................................29
Helpful Identification Features..................................................36
Index................................................................................................43

Field Guide to Stink Bugs of Agricultural

Importance in the United States

ii

iii

Field Guide to Stink Bugs of Agricultural

Importance in the United States

Photo Credits
Ralph Bagwell, Bayer
CropScience

Henry Hogmire, West Virginia

University

Bob Barber

Katherine Kamminga, Virginia

Tech (back cover)

Thomas Bentley
Joseph Berger, Bugwood.org
Lydia Brown, University of
Arizona
Mark Brown, USDA, ARS
Margarethe Brummermann
Valeria Bugh
C. Scott Bundy, New Mexico
State University

Thomas Kuhar, Virginia Tech

David R. Lance, USDA, APHIS
PPQ, www.insectimages.org
Deepak Matadha
Ron Melder
Graham Montgomery
Kim Moore, kimssight.zenfolio.
com
Russ Ottens

Byrain, Bugguide.net

David Owens, Virginia Tech

Jack Kelly Clark, courtesy

University of California
Statewide IPM Program

Herbert A. Joe Pase III,

Bugwood.org

Patrick Coin
Stephen Cresswell

Herb Pilcher, USDA, ARS,

Bugwood.org

H. Doughty, Virginia Tech

Robert Pitman, Virginia Tech

Susan Ellis, Bugwood.org

Phillip Roberts, University of

Georgia

Tim FisherPoff, Virginia Tech

Jim Gilbert (front cover)
Jeremy Greene, Clemson
University
Joyce Gross

Naomi Pier, University of Arizona

Scott Stewart, University of

Tennessee
John Van Duyn, North Carolina
State University
James Wilson, Virginia Tech

Stink Bug Diagram

Antennae

Head
Thorax

Abdomen

Tim FisherPoff

Wings

Stink Bug Diagram

iv

Stink Bug Diagram cont.

Jugum
Tylus

Margins

Pronotum

Shoulders

Scutellum

1
2

Tim FisherPoff

Tarsi

Beaks of Predators versus Pests

Predator: Broad proboscis

(beak) twice the width of the

antennae; first segment is
not attached to head except
at base.

Tim FisherPoff

Tim FisherPoff

Pest: Narrow proboscis (beak),

width of the proboscis and
antennae are similar; first segment
attached to head along its length.

Beaks of Predators versus Pests

vi

Economically Important Species

Unhatched

David Owens

Hatched

Katherine Kamminga

AD

Green stink bug, Acrosternum hilare (Say)

ULT SIZE

Eggs: One of the largest stink

bug eggs. Deposited in clusters of
approximately 20-50. Micropylar
processes (little nubs at the crown
of the egg) are numerous and bend
distinctly at the end; distinctively
pure white until ready to hatch;
egg shell is smooth.
Nymphs: Early instars are mostly
black with orange markings. Later
David Owens
instars have a mostly black
or green head and thorax;
abdomen is mostly green
with dark spots down the
center. Darker color forms
are common.
Adults: Green with black
bands on antennae;
pointed spine on the
underside of the abdomen
between where the hind
legs attach to the body.

Katherine Kamminga
Katherine Kamminga

LT SIZE

ADU

Southern green stink bug, Nezara viridula (L.)

Eggs: White changing to pink

as they mature; deposited in
hexagon-shaped clusters of rows
with approximately 150 eggs
per cluster.
Nymphs: Early instar is
dark brown. Later instars are
green with white spots on
the abdomen with pink-red
markings around the outer
edge.
Adults: Solid mottled brown
usually with rounded shoulders.
Green with red bands on
antennae; rounded spine on
the underside of the abdomen
between where the hind legs
attach to the body.

Herb Pilcher

Scott Stewart

Jeremy Greene

Scott Stewart
Jeremy Greene

Economically Important Species

Economically Important Species

AD

Brown stink bug, Euschistus servus (Say)

U LT SIZE

Herb Pilcher

Eggs: Manila-colored (less

white than green stink bug
eggs) and deposited in loosely
bound clusters; rough-looking
egg shell.
Nymphs: Head and pronotum
of early instar is dark brown;
abdomen is light brown with
slightly darker spots. Later
instar is green-brown to
yellow-brown with light brown
spots down the middle of the
abdomen.
Adults: Solid mottled
brown usually with rounded
shoulders.
Katherine Kamminga

Katherine Kamminga

Katherine Kamminga

U LT SIZE

AD

Dusky stink bug, Euschistus tristigmus (Say)

Herb Pilcher

Eggs and nymphs:

Similar to the brown

stink bug.
Adults: Similar to
the brown stink bug
except with pointed
shoulders; underside
of the abdomen is
light-colored with
three spots.

Herb Pilcher

Katherine Kamminga
Patrick Coin

Economically Important Species

U LT SIZE

Economically Important Species

Euschistus quadrator (Rolston)

AD

Eggs and nymphs: Similar to the

brown stink bug.

Adults: Similar to the dusky stink

bug, but has a convex pronotum

(not apparent in image) and lacks
abdominal spots.

Herb Pilcher

Herb Pilcher

Herb Pilcher

Herb Pilcher

Jack Kelly Clark

ADU

LT SIZE

Consperse stink bug,

Euschistus conspersus Uhler

Eggs: Similar to the brown

stink bug; eggs are pearly
white when first laid, and
eggs are laid in batches.
Nymphs: Similar to the
brown stink bug.
Adults: Similar in general
appearance to the brown
stink bug, but with much
larger black spots on the
legs. Other than genitalia,
the relative size of leg spots is
the best way to separate the
consperse stink bug from the
brown stink bug.

Jack Kelly Clark

Jack Kelly Clark

Economically Important Species

Brown marmorated stink bug,

Halyomorpha halys (Stl)

Deepak Matadha

Deepak Matadha

LT SIZE

Economically Important Species

ADU

Eggs: White to pale green

and deposited in clusters of
approximately 25; appear
somewhat shiny.
Nymphs: Early instars have
a dark head and pronotum;
abdomen is orange and red with
black stripes on the outer edges
and down the center. Later
instars have a mostly black head
and pronotum; abdomen is
rust-colored with black spots on
the outer edges and down the
center; antennae and legs have
white bands.
Adults: Speckled brown-gray;
antennae have alternating dark and white
bands; dark and white bands around the
outer edges of the abdomen; small round
coppery patches on or near the head;
the underside is light gray, brown, or tan
(not green or yellow).
David R. Lance

Deepak Matadha

U LT SI ZE
AD

Rice stink bug, Oebalus pugnax (F.)

Eggs: Deposited in two rows of up to

45 eggs that change from green to red
as they mature.
Nymphs: White to brown head and
thorax; light-colored abdomen with red
speckles and dark spots in the center.
Adults: Tan, narrow body with
forward-pointing spines on the
shoulders; scutellum is yellow.
C. Scott Bundy

C. Scott Bundy

Patrick Coin

Economically Important Species

Redshouldered stink bug,

Thyanta custator accerra McAtee

C. Scott Bundy

Eggs: Loosely aligned in

large clusters that change
from white to green as they
mature.
Nymphs: Mostly white with
dark red to black spots on
the thorax and black and
red-orange stripes on the
abdomen.
Adults: Green, often with
a red-pink stripe across the
pronotum; scutellum with
a red-pink tip; no spine
on the underside of the
abdomen between where
the hind legs attach to the
body.

Jeremy Greene

Jeremy Greene

U LT SI ZE

Economically Important Species

AD

U LT SIZE

AD

Thyanta custator custator (F.)

Nymphs: Dark red to black

thorax with an outer white

margin; abdomen is white
with black stripes.
Adults: Green, often with a
broad single red-pink stripe
across the pronotum;
black markings along
the outer edge of the
pronotum.

Herb Pilcher

Herb Pilcher

Herb Pilcher

Economically Important Species

10

Redbanded stink bug,

Piezodorus guildinii (Westwood)
C. Scott Bundy

Russ Otttens

U LT SI ZE

Economically Important Species

AD

11

Eggs: Usually dark red to

brown with a white band
around the outer edge and
deposited in two rows with
approximately 30 eggs.
Nymphs: Later instars are
mostly green and have
brown to red stripes on
the sides and top of the
abdomen.
Adults: Shiny green to
yellow-green with two
stripes across the pronotum
(the front is yellow and the
back is dark red); long spine
on the underside of the
abdomen between where the
hind legs attach to the body
(circled).

Russ Otttens
Ralph Bagwell

Thomas Kuhar

Thomas Kuhar

AD

U LT SIZ

Harlequin bug,
Murgantia histrionica (Hahn)

Eggs: White with a

small black stripe at
the base, a larger black
stripe near the top, and
a black crescent on the
top. Deposited in small
clusters of two rows.
Nymphs: Shiny,
mostly black with
orange-red markings
on the pronotum and
outlining the middle of
the abdomen; abdomen
also has white stripes.
Adults: Black with
distinct orange or red
markings; head slopes
downward.

Thomas Kuhar

Economically Important Species

12

13 Economically Important Species

U LT S

C. Scott Bundy

Eggs: White when deposited, turning pink as they mature; typically

deposited singly beneath the soil, occasionally deposited in small
clusters on or near the host plant.
Nymphs: First four instars
with head, thorax, and legs
reddish-brown; abdomen red
with reddish-brown plates.
Fifth instar typically with
white markings on thorax and
abdomen.
Adults: Body black with white
and orange markings on head,
C. Scott Bundy
thorax, and abdomen; head
strongly angled downward
(ventrally). This invasive
species looks somewhat
similar to the harlequin bug,
but is significantly smaller.
Note: This invasive Old
World pest first was detected
in California in 2008 and
has since spread eastward to
Texas.
C. Scott Bundy

IZ E

AD

Painted bug (bagrada bug),

Bagrada hilaris (Burmeister)

LT SIZE
ADU

Say stink bug, Chlorochroa sayi (Stl)

and Conchuela, Chlorochroa ligata (Say)

Because they are similar in

appearance, descriptions of
Say stink bug and Conchuela
are combined.

Lydia Brown

Naomi Pier

Say stink bug color variants

Eggs: Cream-colored with two

tan rings dorsally with a central
cream (C. ligata) or tan (C.
sayi) spot. Eggs are deposited
in clusters of 20-50 in multiple
rows.
Nymphs: Head and thorax
usually black in early instars,
turning dark green (or
remaining black) in late instars;
margins of thorax (and often
abdomen) tinged white, yellow,
or orange. Abdomen black to
violet, often turning dark green
in late instars.

continued on page 15

Lydia Brown

Photos on this page are of Say stink bug.

Economically Important Species

14

Say stink bug, Chlorochroa sayi (Stl)

and Conchuela, Chlorochroa ligata (Say)

LT SIZE

Economically Important Species

ADU

15

continued from page 14

Adults: Body green or black* with

orange or white margins on the

thorax and abdomen and orange
markings on head, thorax, and
abdomen. Scutellum with a white
or red spot at the tip, often with
three white spots along upper
margin (C. sayi).
Valerie Bugh

*Note: In much of the Southwest

(western Texas, New Mexico, Arizona,
and California) adults of C. ligata are
present in their black form and are easily
distinguished from C. sayi, which are
usually green. Even with dark variants of
C. sayi, the adults still are shades of green
rather than black seen in C. ligata. In
other regions, adults of both species may
be green and require examination of other
features to separate species.

Kim Moore

Joyce Gross

Photos on this page are of the Conchuela

A D ULT SIZE

Less Common Species

Edessa bifida (Say)
Ron Melder

Ron Melder

Adults: Large, oval, green-brown

with a broad convex pronotum;

scutellum with a white tip.

AD

U LT S I

Twice-stabbed stink bug,

Cosmopepla lintneriana Kirkaldy

Adults: Small and black with a

red stripe across the pronotum

and along the outer edge of the
pronotum and abdomen; scutellum
with two red
spots at the tip.

Thomas Bentley

Scott Stewart

Less Common Species

16

ZE

A D ULT SIZE

Less Common Species

17

Menecles insertus (Say)

Adults: Large, oval, brown

and appears flattened;

pronotum is broad and
protrudes forward almost
level with the eyes; a
thin white stripe cuts the
pronotum lengthwise.

Patrick Coin

AD
ULT SIZE

Proxys punctulatus (Palisot de Beauvois)

Adults: Black with

sharp shoulders;
scutellum with a
white spot at the
tip; legs are ivorycolored near the
body, changing to
black.

David Owens

E
LT SIZ
ADU

Jade stink bug, Banasa euchlora (Say)

Adults: Small and

green with white to

yellow markings at
the three corners of
the scutellum.

Patrick Coin

DU
LT SI Z E

Banasa dimidiata (Say)

Adults: Small, green-

brown; front portion

of the pronotum
is green and back
portion is red-brown;
scutellum may be
either green-brown or
red-brown.

Patrick Coin

Less Common Species

18

ULT SIZE

Less Common Species

19

AD

Mormidea lugens (F.)

Adults: Small and

black-brown; scutellum
and pronotum with
white to yellow outer
edges. Antennae are
black with white bands.

Thomas Bentley

ADU

LT SIZE

Rough stink bug,

Brochymena quadripustulata (F.)

Adults: Large and brown

with small orange spots;

pronotum toothed on the
outer edge; legs have
white bands.

Patrick Coin

LT SIZE
ADU

Onespotted stink bug,

Euschistus variolarius (Palisot de Beauvois)

Eggs and nymphs (not

shown): Similar to

brown stink bug.

Adults: Similar to
brown stink bug
but has one spot on
the underside of the
abdomen. Similar to
spined soldier bug but
lacks pointed shoulders
and pointed abdominal
spine.

Graham Montgomery

Graham Montgomery

Less Common Species

20

Eggs: Silvery to metalliclooking with spiny nubs and

deposited in loose clusters.
Nymphs: Early instars are
dark red with black heads
and black spots on the top
side of the abdomen. Later
instars are tan to orange with
red and white stripes on the
abdomen.
Adults: Brown with pointed
shoulders; underside of
the abdomen is light with
a single black spot in the
center toward the rear and
a long, pointed abdominal
spine.

AD

Spined soldier bug,

Podisus maculiventris (Say)

U LT SIZ E

Beneficial Species

21

Jeremy Greene

Katherine Kamminga

Katherine Kamminga

Katherine Kamminga

ADU

LT SI Z

Florida predatory stink bug,

Euthyrhynchus floridanus (L.)

Adults: Mostly black with

red, yellow, or orange

spots at each corner of
the scutellum; however,
spots can merge; shoulders
pointed.

Patrick Coin
A
DU

..

LT SI Z E

Perillus strigipes (Herrich-Schaffer)

Adults: Black with an

orange-red-shaped
V-pattern and rounded
shoulders.

Ron Melder

Beneficial Species

22

Beneficial Species
ZE

AD

Anchor stink bug, Stiretrus anchorago (F.)

U LT S I

23

Adults: Unusual in that they

can be all black, black and

orange, or black and white
with different color patterns.
Body is convex with a large
scutellum.

Patrick Coin

Patrick Coin

Ron Melder
Stephen Cresswell

AD

U LT SIZ E

Twospotted stink bug,

Perillus bioculatus (F.)
Adults: Mostly

black; pronotum
red- or ivorycolored with two
black spots; red- or
ivory-colored
triangle on the
scutellum.

Margarethe Brummerman

Bob Barber

Beneficial Species

24

Squash bug, Anasa tristis (De Geer)

James Wilson

LT SIZE

Stink Bug-Like Species

ADU

25

Eggs: Newly laid eggs are typically

light in color and become coppery,
then dark maroon as they develop.
Eggs are laid in masses often in
a diamond-shaped pattern along
leaf veins.
Nymphs: Are light green upon
hatching and light to dark gray with
each successive molt. All nymphs
have black legs and antennae.
Adults: Are cryptic and like to hide
in ground litter. They range from
2
/511/5 inches long and are dark
gray, brown, or black in color with
orange or brown markings on the
sides of the abdomen.

H. Doughty

H. Doughty

LT SIZE
AD U

Leaffooted bug, Leptoglossus

phyllopus (L.) and L. oppositus (Say)

Leaffooted bugs are distinguished by the unusual Thomas Kuhar

flattening of the hind tibia that resembles a
leaf, hence the common name. There are two
common species of agricultural importance.
Eggs: The eggs are brown and are laid in a single
row along a stem or leaf midrib. They
are somewhat cylindrical, flattened on
the undersides and at the ends, and are
laid closely, end to end.
Nymphs: The nymphs have black legs
while their bodies range in color from
orange to reddish brown. The leaf-like
flattening on the hind tibia does not
appear until later instars.
Adults: The adults of both species have
a flat, dark brown body with gray to
medium brown sides. The body is
V-shaped in cross section. L. phyllopus
H. Doughty
has a white to pale yellow straight,
broad band across the wing covers.
Thomas Kuhar
While L. oppositus
lacks the band,
H. Doughty
small white spots
may appear on
some individuals.

Adult L. oppositus

Adult L. phyllopus

Stink Bug-Like Species

26

Kudzu bug, Megacopta cribraria (F.)

Jeremy Greene

LT SIZE

Stink Bug-Like Species

ADU

27

The species has a preference for

leguminous hosts, such as kudzu,
wisteria, soybeans, and others, but it
has been reported on fruit trees and
various other hosts. Loss of soybean
yield can result from extended
exposure to these insects.
Eggs: Eggs are light tan and laid in a
slanting position in two-row masses
about 3/8 inch long.
Nymphs: Young nymphs are
small and orange, and older
nymphs are very hairy but
resemble adults in body shape.
When disturbed, the insects
produce a foul odor similar to
that produced by stink bugs.
Adults: Adults are about 1/5 inch

Jeremy Greene

long, olive-green colored with

dark brown speckles, and are
almost square-shaped but taper
near the head region.

Jeremy Greene

Jeremy Greene

ADULT SIZE

Wheel bug,
Arilus cristatus (L.)
Joseph Burger

Eggs: Eggs are laid tightly in

large clusters of 50182 eggs.

Individual eggs resemble
small brown bottles with
white stoppers.
Nymphs: Nymphs are bright

red with black marks that

become darker and more
gray with each molt.

Nymphs emerging from egg mass

Adults: This large dark gray

assassin bug ranges from 1

11/4 inches long with long legs
and antennae, a stout beak,
large eyes on a small head,
and has a prominent dorsal
semicircular crest shaped like a
cogwheel on
its thorax.

Susan Ellis

Herbert A. Joe Pase III

Stink Bug-Like Species

28

29

Feeding Injury

Injury to cotton
External boll injury is
characterized by small,
round, shallow purpleblack depressions,
usually 1/32 1/16 inch in
diameter. These spots
tend to be larger than
the tiny spots usually
seen on maturing bolls.
Phillip Roberts

Phillip Roberts

Injured bolls will have yellow-, tan-,

or brown-stained lint in the seed
areas, often, but not always, under
the external feeding spots. Other
injury symptoms include small wartlike growths and/or dark pin prick
spots on the inside of the boll wall.
Internal boll injury may be present
without obvious external evidence.
Phillip Roberts

Injury to cabbage

Thomas Kuhar

The harlequin bug sucks sap

from the cabbage plant, causing
yellow or white blotches to appear on
leaves. This feeding can cause plants to wilt, turn brown, or die and
affects commercial value.

Feeding Injury

30

Feeding Injury

31
Injury to corn

Stink bug feeding on ears

results in loss of kernels
and cob deformation.

John Van Duyn

John Van Duyn

Feeding on stalks appears as

brown spots and may include
a white stylet sheath in the
center of the feeding spot.

John Van Duyn

David Owens

Injury to peaches
The most
common form
of injury is the
characteristic
catfacing. The
fruit growth stage
determines the
severity of the
damage.

Henry Hogmire

Henry Hogmire

Henry Hogmire

Feeding Injury

32

33

Feeding Injury

Injury to apples

Mark Brown

Early season feeding results in dimples or depressed areas.

Mark Brown

Internal injury appears as white, pithy areas that turn brown.

Injury to tomatoes
Feeding on
tomatoes results
in minute
puncture marks
on the fruit
surrounded by a
yellow halo.

Katherine Kamminga

Injury to ripe tomatoes appears as a pithy

or white to yellow corky area.

Feeding Injury

Katherine Kamminga

34

Feeding Injury

35
Injury to soybean

Feeding
punctures
appear as small
brown or black
spots on seed
and result in
deformed,
undersized,
discolored, or
shriveled seeds.

Scott Stewart

Feeding can result in flattened pods.

Robert Pitman

Helpful Identification Features

Features of green-colored stink bugs

Jeremy Greene

Southern green stink bug has a

rounded spine (called the abdominal
spine) on the underside of the
abdomen between where the legs
attach to the body, and antennae
have red bands.

Tim FisherPoff

Tim FisherPoff

Katherine Kamminga

Green stink bug has a pointed

abdominal spine, and antennae have
black bands.

Helpful Identification Features

36

37

Helpful Identification Features

Features of green-colored stink bugs cont.

Say stink bug

Lydia Brown

Say stink bug and Conchuela can vary from black to green, but
always have orange or white margins on the thorax and abdomen
Conchuela

Joyce Gross

Features of small, green-colored stink bugs

Thyanta custator custator
occurs only in the Costal
Plain of the U.S. and has
narrow black markings
along the outer edge of the
pronotum.

Herb Pilcher

Redshouldered stink bug lacks

the black markings on the outer
edge of the pronotum.

Jeremy Greene

Redbanded stink bug has a long

abdominal spine.

Russ Ottens
Ralph Bagwell

Helpful Identification Features

38

39

Helpful Identification Features

Features of brown-colored stink bugs

Brown stink bug usually
has rounded shoulders
and lacks an abdominal
spine.

Katherine Kamminga

Dusky stink bug has pointed

shoulders, lacks an abdominal
spine, and has three black
spots on the underside of the
abdomen.

Katherine Kamminga

Patrick Coin

Spined soldier bug has

pointed shoulders, a long
abdominal spine, and a
black spot on the underside
of the abdomen. Differs
from other brown species by
having predaceous mouthparts (see figures, p. vi).

Katherine Kamminga

Katherine Kamminga

Euschistus quadrator
has pointed
shoulders, lacks
an abdominal
spine, and lacks
abdominal spots.
Herb Pilcher

Helpful Identification Features

40

41

Helpful Identification Features

Features of brown-colored stink bugs cont.

Byrain

Consperse stink bug looks similar to brown stink bug, but with large
brown spots on legs.

Graham Montgomery

Graham Montgomery

Onespotted stink bug looks similar

to the spined soldier bug but lacks
pointed shoulders, and similar
to the brown stink bug but has a
single spot on the underside of the
abdomen.

Rough stink
bug, Brochymena
quadripustulata, has
a pronotum that is
toothed on the outer
edge.

Patrick Coin

David R. Lance

Brown marmorated stink bug, Halyomorpha halys, has a pronotum

that is smooth on the outer edge; antennae have alternating dark and
white bands.

Helpful Identification Features

42

43

Index
Page

Acrosternum hilare, Chinavia hilaris, Green stink bug................... 1, 36

Anasa tristis, Squash bug.......................................................................25
Arilus cristatus, Wheel bug....................................................................28
Bagrada hilaris, Painted bug.................................................................13
Banasa dimidiata...............................................................................18
Banasa euchlora, Jade stink bug*.........................................................18
Brochymena quadripustulata, Rough stink bug.............................19, 42
Chlorochroa ligata, Conchuela...................................................... 15, 37
Chlorochroa sayi, Say stink bug..................................................... 14, 37
Cosmopepla lintneriana, Twice-stabbed stink bug*.............................16
Edessa bifida......................................................................................16
Euschistus conspersus, Consperse stink bug*................................... 6, 41
Euschistus quadrator...................................................................... 5, 40
Euschistus servus, Brown stink bug................................................... 3, 39
Euschistus tristigmus, Dusky stink bug............................................... 4, 39
Euschistus variolarius, Onespotted stink bug.................................20, 41
Euthyrhynchus floridanus, Florida predatory stink bug*......................22
*Not approved Entomological Society of America common name.

Page

Halyomorpha halys, Brown marmorated stink bug.........................7, 42

Leptoglossus phyllopus, Leaffooted bug...............................................26
Megacopta cribraria, Kudzu bug*.......................................................27
Menecles insertus...............................................................................17
Mormidea lugens...............................................................................19
Murgantia histrionica, Harlequin bug...........................................12, 30
Nezara viridula, Southern green stink bug..................................... 2, 36
Oebalus pugnax, Rice stink bug............................................................. 8
Perillus bioculatus, Twospotted stink bug..............................................24
Perillus strigipes..................................................................................22
Piezodorus guildinii, Redbanded stink bug................................... 11, 38
Podisus maculiventris, Spined soldier bug..................................... 21, 40
Proxys punctulatus..............................................................................17
Stiretrus anchorago, Anchor stink bug*................................................23
Thyanta custator accerra, Redshouldered stink bug....................... 9, 38
Thyanta custator custator............................................................. 10, 38
*Not approved Entomological Society of America common name.

Index

44

Visit StopBMSB.org to learn more about the biology, ecology, and

management of brown marmorated stink bug in specialty crops.
This guide is supported by SCRI Coordinated Agricultural Project grant
#2011-51181-30937.

Support comes from USDA-NIFA:

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