Pond and Lake Ecosystems: Littoral Macrophytes Euphotic Zone Photosynthesizers Epilimnion
Pond and Lake Ecosystems: Littoral Macrophytes Euphotic Zone Photosynthesizers Epilimnion
Pond and Lake Ecosystems: Littoral Macrophytes Euphotic Zone Photosynthesizers Epilimnion
A pond or lake ecosystem includes biotic (living) plants, animals and micro-organisms, as well as abiotic
(nonliving) physical and chemical interactions. Pond and lake ecosystems are a prime example of lentic
ecosystems. Lentic refers to stationary or relatively still water, from the Latin lentus, which means
sluggish.
A typical lake has distinct zones of biological communities linked to the physical structure of the lake.
(Figure below) The littoral zone is the near shore area where sunlight penetrates all the way to the
sediment and allows aquatic plants (macrophytes) to grow. Light levels of about 1% or less of surface
values usually define this depth. The 1% light level also defines the euphotic zone of the lake, which is
the layer from the surface down to the depth where light levels become too low for photosynthesizers.
In most lakes, the sunlit euphotic zone occurs within the epilimnion.
However, in unusually transparent lakes, photosynthesis may occur well below the thermocline into the
perennially cold hypolimnion. For example, in western Lake Superior near Duluth, MN, summertime
algal photosynthesis and growth can persist to depths of at least 25 meters, while the mixed layer, or
epilimnion, only extends down to about 10 meters. Ultra-oligotrophic Lake Tahoe, CA/NV, is so
transparent that algal growth historically extended to over 100 meters, though its mixed layer only
extends to about 10 meters in summer. Unfortunately, inadequate management of the Lake Tahoe
basin since about 1960 has led to a significant loss of transparency due to increased algal growth and
increased sediment inputs from stream and shoreline erosion.
The higher plants in the littoral zone, in addition to being a food source and a substrate for algae and
invertebrates, provide a habitat for fish and other organisms that is very different from the open water
environment.
Pond and Lake Ecosystems
The productivity of this zone largely depends upon the organic content of the sediment, the amount of
physical structure, and in some cases upon the rate of fish predation. Sandy substrates contain relatively
little organic matter (food) for organisms and poor protection from predatory fish. Higher plant growth
is typically sparse in sandy sediment, because the sand is unstable and nutrient deficient. A rocky
bottom has a high diversity of potential habitats offering protection (refuge) from predators, substrate
for attached algae (periphyton on rocks), and pockets of organic "ooze" (food). A flat mucky bottom
offers abundant food for benthic organisms but is less protected and may have a lower diversity of
structural habitats, unless it is colonized by higher plants.
Lake Organisms