The Human Eye

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The Human Eye

STRUCTURE AND ROLES

Sclera: the tough and fibrous outer layer of the eye.

Choroid layer: the middle layer of the eye. It contains blood vessels and melanin pigment.

Retina: the inner layer of the eye. It contains light-sensitive rod and cone cells.

Cornea: the front, transparent part of the sclera that refracts light rays entering the eye.

Conjunctiva: a membrane surrounding the cornea, lubricated by watery tear fluid.

Iris: a modification of the choroid layer at the front of the eye. It contains smooth radial muscle and
circular muscle. Contraction and relaxation of the iris muscles change the change the size of the pupil
and allow variable amounts of light to enter the eye.

Pupil: a hole in the iris that becomes wider in dim light.

Optic nerve: conveys nerve signals to the brain. (The light-sensitive rod and cone cells in the retina
connect to sensory neurones that leave the retina at the blind spot and form the optic nerve).

Lens: a transparent disc of tissue that changes shape for near or far vision.

Suspensory ligaments: holds lens within the ring of ciliary muscle.

Ciliary muscle: a ring of muscle that alters the shape of the lens.

Aqueous humour: a clear, watery fluid in front of the lens.

Vitreous humour: a clear, jelly-like fluid behind the lens.

(http://blogs.hightechhigh.org/mhansen2014/files/2014/05/pic-eye-anatomy.jpg)

FOCUSING LIGHT

When light rays enter the eye they are refracted (bent). First, the light is refracted by the cornea at the
front of the eye. Most of the refraction occurs here because it is curved. The light is then refracted
further by the lens, which focuses (converges) the light so that it forms a clear image on the retina.

There are three refracting surfaces: the cornea, the front surface of the lens, and the rear surface of
the lens.

ACCOMMODATION

Definition: the flattening or thickening of the lens by the ciliary muscles depending on the distance
of an object from the eye.
Accommodation causes light to be refracted to a greater or lesser extent depending on how far the
object is from the eye.

DISTANT OBJECTS

Light rays from a distant object are almost parallel when they enter the eye and these need a thin lens
to focus them on the retina. When you look at a distant object the ciliary muscles relax, pulling the
suspensory ligaments taut. The lens, which is made of flexible proteins, is pulled into a thin shape.

NEAR OBJECTS

Light rays from a near object enter the eye at a more acute angle and these need a thicker lens to focus
them on the retina. When you look at a near object the ciliary muscles contract, causing the suspensory
ligaments to become looser. This allows the lens to return to its natural round, fat shape. This refracts
the light rays more so that they can be focused on the retina.

RECEPTOR CELLS: RODS & CONES

Definition: a receptor cell is able to detect changes in the environment, converting sensory
information into a form that can be transmitted to the central nervous system. In the eye, these
are the rod and cone cells.

RODS

Definition: photoreceptors that are responsible for vision at low light intensities. They are mostly
found in the retina, spread evenly across it.

ACTION POTENTIALS

Rod cells contain the light-sensitive pigment rhodopsin. When light hits a rod cell it causes the
rhodopsin to split into opsin and retinal. This process is called bleaching. As a result of bleaching, the
permeability of the rod cell to sodium ions is altered. The rod cell membrane becomes depolarised (a
generator potential).

If this potential is enough to overcome the threshold value, an action potential is generated in the
sensory neurone which carries the nerve impulse along the optic nerve to the brain. After this, the
rhodopsin is regenerated - this process requires ATP.

MONOCHROMATIC VISION

Rod cells are extremely sensitive to low light intensities, so they are most useful for vision in dim light.
However, they cannot distinguish between different wavelengths, meaning they only produce images in
black and white. This is called monochromatic vision.

LOW VISUAL ACUITY

Several rod cells synapse with just one sensory neurone. Therefore, rod cells have low visual
acuity, e.g. they cannot discriminate very well.
However, this grouping of red cells together also increases their sensitivity, which is why they can detect
low levels of light. If each of several rod cells results in a tiny amount of neurotransmitter being released
into the synaptic cleft, these small amounts 'add together' so that there is enough to set up an action
potential in the sensory neurone. This is called summation.

CONES

Definition: photoreceptors that are responsible for colour vision and ones which function best in
bright light conditions. They are mostly concentrated at the fovea.

TRICHROMATIC COLOUR THEORY

Cone cells have a similar structure to rod cells. However, they contain a pigment called iodopsin instead.
There are three different kinds of iodopsin, each sensitive to red, green or blue light.

Different colours are perceived depending on the proportion of the different types of cone cell that are
stimulated. E.g. if red and green cones are stimulated equally, the brain interprets the colour as yellow.
This is called the trichromatic theory of colour vision.

HIGH VISUAL ACUITY

Most cones have an individual connection to a sensory neurone, meaning that they have a much
greater visual acuity than rods. They are therefore responsible for all high-resolution vision.

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