16 - Ecosystem Energetics
16 - Ecosystem Energetics
16 - Ecosystem Energetics
2 Laws of Thermodynamics 1. Energy can neither be created nor destroyed, only transformed from one type to another
conservation of energy
Primary production capturing light energy and storing it in chemical bonds of carbon compounds
primary productivity the rate of primary production primary producers photosynthetic autotrophs
Assimilated Carbon
Gross primary production (GPP) total amount of energy assimilated by photosynthesis Net primary production (NPP) energy actually stored as biomass GPP NPP = respiration
GPP
NPP
Measuring NPP in nature Units: energy per unit area per year
kJ per m2 per yr, or W per m2
1 g C assimilated = 39 kJ energy
can use plant biomass or CO2 uptake as an estimate of energy
2. Use radioactive isotopes to measure total uptake of C 3. In aquatic systems, measure changes in O2 concentration
Light
Dark
Temperature
most plants have an optimum temperature respiration increases with temperature
Water
transpiration (or water use) efficiency amount of plant tissue produced per kilogram of water transpired
2-4 g tissue/kg water
Nutrients
Liebigs law of the minimum most important in deserts, open ocean, agriculture
Ecological efficiency proportion of the biomass of one trophic levels transformed into biomass at the next higher trophic level
Exploitation efficiency proportion of production on one trophic level consumed by the next higher level
usually less than 100%
Ingestion by herbivores
Amount of energy actually absorbed from food is assimilated energy Assimilation efficiency proportion of ingested energy actually absorbed by the body
seeds 80% young vegetation 60-70% grazing/browsing 30-40% wood 15% animals 60-90%
Ingestion by herbivores
Indigestible
Assimilation
Ingestion by herbivores
Indigestible
Assimilation
Growth
Ingestion by herbivores
Indigestible
detritivores
terrestrial communities
Ingestion by herbivores
Indigestible
Ingestion by herbivores
Indigestible
Ingestion by herbivores
Indigestible
Growth
Residence time average time that energy spends on one trophic level
= (energy stored in biomass)/(net productivity)
Biomass accumulation ratio residence time based on biomass rather than energy
= (biomass)/(rate of biomass production)