Astyanax |
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Astyanax mexicanus |
Scientific classification |
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Genus: |
Astyanax
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Type species |
Astyanax argentatus
Baird & Girard, 1854
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Synonyms |
Anoptichthys Hubbs & Innes, 1936
Bertoniolus Fowler, 1918
Evenichthys Whitley, 1935
Poecilurichthys Gill, 1858
Psalidodon Eigenmann, 1911
Zygogaster Eigenmann, 1913
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Astyanax is a genus of freshwater fish in the family Characidae of the order Characiformes. Some of these fish, like many of their relatives, are kept as aquarium pets and known collectively as tetras. With around 140 described species and new ones being described every few months, this genus is among the largest of the entire order; Hyphessobrycon also has more than 130 species, and which one is larger at any one time depends on whether more species have been recently described in one or the other. The blind cave fish, a form of A. mexicanus, is a famous member of this genus. A. jordani a different but closely related blind fish, was formerly considered to form its own genus, Anoptichthys.
The type species is A. mexicanus, the Mexican tetra. The generic name comes from Astyanax, a character in Greek mythology, who was the son of Hector of Troy; in homage to this, several specific epithets also refer to the Iliad.[1]
Taxonomy and systematics
This genus is more or less defined as in 1917 by Carl H. Eigenmann. The lack of comprehensive phylogenetic analyses dealing with this genus is hindering a thorough review, and the relationships remain as indeterminate as the monophyly is doubtful. Although included in Astyanax by FishBase, phylogenetic evidence supports moving A. aurocaudatus back to the monotypic genus Carlastyanax.[2]
Species
There are currently 146 recognized species in this genus:
- Astyanax abramis (L. Jenyns, 1842)
- Astyanax aeneus (Günther, 1860) (Banded tetra)
- Astyanax ajuricaba Marinho & F. C. T. Lima, 2009
- Astyanax alburnus R. F. Hensel, 1870
- Astyanax altior C. L. Hubbs, 1936 (Yucatán tetra)
- Astyanax altiparanae Garutti & Britski, 2000
- Astyanax angustifrons (Regan, 1908)
- Astyanax anterior C. H. Eigenmann, 1908
- Astyanax aramburui Protogino, Miquelarena & H. L. López, 2006
- Astyanax argyrimarginatus Garutti, 1999
- Astyanax asuncionensis Géry, 1972
- Astyanax atratoensis C. H. Eigenmann, 1907
- Astyanax bagual Bertaco & A. C. Vigo, 2015 [3]
- Astyanax bifasciatus Garavello & F. A. A. Sampaio, 2010 [4]
- Astyanax bimaculatus (Linnaeus, 1758) (Two-spot tetra)
- Astyanax biotae R. M. C. Castro & Vari, 2004
- Astyanax bockmanni Vari & R. M. C. Castro 2007
- Astyanax bourgeti C. H. Eigenmann, 1908
- Astyanax brachypterygium Bertaco & L. R. Malabarba, 2001
- Astyanax brevirhinus C. H. Eigenmann, 1908
- Astyanax burgerai Zanata & Camelier, 2009
- Astyanax caucanus (Steindachner, 1879)
- Astyanax chaparae Fowler, 1943
- Astyanax chico Casciotta & Almirón, 2004
- Astyanax clavitaeniatus Garutti, 2003
- Astyanax cocibolca W. A. Bussing, 2008
- Astyanax cordovae (Günther, 1880)
- Astyanax correntinus (Holmberg, 1891)
- Astyanax courensis Bertaco, F. R. de Carvalho & Jerep, 2010
- Astyanax cremnobates Bertaco & L. R. Malabarba, 2001
- Astyanax daguae C. H. Eigenmann, 1913
- Astyanax depressirostris A. Miranda-Ribeiro, 1908
- Astyanax dissensus C. A. S. Lucena & Thofehrn, 2013 [5]
- Astyanax dissimilis Garavello & F. A. A. Sampaio, 2010 [4]
- Astyanax dnophos F. C. T. Lima & Zuanon, 2004
- Astyanax douradilho Bertaco, 2014 [6]
- Astyanax eigenmanniorum (Cope, 1894)
- Astyanax elachylepis Bertaco & P. H. F. Lucinda, 2005
- Astyanax endy Mirande, G. Aguilera & Azpelicueta, 2006
- Astyanax epiagos Zanata & Camelier, 2008
- Astyanax eremus Ingenito & Duboc, 2014 [7]
- Astyanax erythropterus (Holmberg, 1891)
- Astyanax fasciatus (G. Cuvier, 1819) (Banded astyanax)
- Astyanax fasslii (Steindachner, 1915)
- Astyanax festae (Boulenger, 1898)
- Astyanax filiferus (C. H. Eigenmann, 1913)
- Astyanax gisleni Dahl, 1943
- Astyanax giton C. H. Eigenmann, 1908
- Astyanax goyacensis C. H. Eigenmann, 1908
- Astyanax goyanensis (P. Miranda-Ribeiro, 1944)
- Astyanax gracilior C. H. Eigenmann, 1908
- Astyanax guaporensis C. H. Eigenmann, 1911
- Astyanax guaricana C. A. M. Oliveira, Abilhoa & Pavanelli, 2013 [8]
- Astyanax guianensis C. H. Eigenmann, 1909 [9]
- Astyanax gymnodontus (C. H. Eigenmann, 1911)
- Astyanax gymnogenys C. H. Eigenmann, 1911
- Astyanax hamatilis Camelier & Zanata, 2014 [10]
- Astyanax hastatus G. S. Myers, 1928
- Astyanax henseli F. A. G. Melo & Buckup, 2006
- Astyanax hermosus Miquelarena, Protogino & H. L. López, 2005
- Astyanax integer G. S. Myers, 1930
- Astyanax intermedius C. H. Eigenmann, 1908
- Astyanax ita Almirón, Azpelicueta & Casciotta, 2002
- Astyanax jacobinae Zanata & Camelier, 2008
- Astyanax jacuhiensis (Cope, 1894)
- Astyanax janeiroensis C. H. Eigenmann, 1908
- Astyanax jenynsii (C. H. Eigenmann, 1877)
- Astyanax jordanensis Vera Alcaraz, Pavanelli & Bertaco, 2009
- Astyanax jordani (C. L. Hubbs & Innes, 1936)
- Astyanax kennedyi Géry, 1964
- Astyanax kompi Hildebrand, 1938
- Astyanax kullanderi W. J. E. M. Costa, 1995
- Astyanax lacustris (Lütken, 1875)
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- Astyanax latens Mirande, G. Aguilera & Azpelicueta, 2004
- Astyanax laticeps (Cope, 1894)
- Astyanax leonidas Azpelicueta, Casciotta & Almirón, 2002
- Astyanax leopoldi Géry, Planquette & Le Bail, 1988
- Astyanax lineatus (Perugia, 1891)
- Astyanax longior (Cope, 1878)
- Astyanax longirhinus Garavello & F. A. A. Sampaio, 2010 [4]
- Astyanax maculisquamis Garutti & Britski, 1997
- Astyanax magdalenae C. H. Eigenmann & Henn, 1916
- Astyanax marionae C. H. Eigenmann, 1911
- Astyanax maximus (Steindachner, 1877)
- Astyanax megaspilura Fowler, 1944
- Astyanax mexicanus (De Filippi, 1853) (Mexican tetra)
- Astyanax microlepis C. H. Eigenmann, 1913
- Astyanax microschemos Bertaco & C. A. S. Lucena, 2006
- Astyanax minor Garavello & F. A. A. Sampaio, 2010 [4]
- Astyanax multidens C. H. Eigenmann, 1908
- Astyanax mutator C. H. Eigenmann, 1909
- Astyanax myersi (Fernández-Yépez, 1950)
- Astyanax nasutus Meek, 1907
- Astyanax nicaraguensis C. H. Eigenmann & Ogle, 1907
- Astyanax novae C. H. Eigenmann, 1911 [11]
- Astyanax obscurus (R. F. Hensel, 1870)
- Astyanax ojiara Azpelicueta & J. O. Garcia, 2000
- Astyanax orbignyanus (Valenciennes, 1850)
- Astyanax orthodus C. H. Eigenmann, 1907
- Astyanax pampa Casciotta, Almirón & Azpelicueta, 2005
- Astyanax paraguayensis (Fowler, 1918)
- Astyanax parahybae C. H. Eigenmann, 1908
- Astyanax paranae C. H. Eigenmann, 1914
- Astyanax paranahybae C. H. Eigenmann, 1911
- Astyanax paris Azpelicueta, Almirón & Casciotta, 2002
- Astyanax pedri (C. H. Eigenmann, 1908)
- Astyanax pelecus Bertaco & C. A. S. Lucena, 2006
- Astyanax pellegrini C. H. Eigenmann, 1907
- Astyanax pirabitira C. A. S. Lucena, Bertaco & Berbigier, 2013 [12]
- Astyanax pirapuan Tagliacollo, Britzke, G. S. C. Silva & Benine, 2011 [13]
- Astyanax poetzschkei C. G. E. Ahl, 1932
- Astyanax procerus C. A. S. Lucena, J. B. Castro & Bertaco, 2013 [5]
- Astyanax puka Mirande, G. Aguilera & Azpelicueta, 2007
- Astyanax pynandi Casciotta, Almirón, Bechara, J. P. Roux & Ruiz Díaz, 2003
- Astyanax ribeirae C. H. Eigenmann, 1911
- Astyanax rivularis (Lütken, 1875)
- Astyanax robustus Meek, 1912
- Astyanax ruberrimus C. H. Eigenmann, 1913
- Astyanax rupununi Fowler, 1914
- Astyanax saguazu Casciotta, Almirón & Azpelicueta, 2003
- Astyanax saltor Travassos, 1960
- Astyanax scabripinnis (L. Jenyns, 1842)
- Astyanax schubarti Britski, 1964
- Astyanax scintillans G. S. Myers, 1928
- Astyanax serratus Garavello & F. A. A. Sampaio, 2010 [4]
- Astyanax siapae Garutti, 2003
- Astyanax stenohalinus Messner, 1962
- Astyanax stilbe (Cope, 1870)
- Astyanax superbus G. S. Myers, 1942
- Astyanax symmetricus C. H. Eigenmann, 1908
- Astyanax taeniatus (L. Jenyns, 1842)
- Astyanax totae Ferreira Haluch & Abilhoa, 2005
- Astyanax trierythropterus Godoy (pt), 1970
- Astyanax troya Azpelicueta, Casciotta & Almirón, 2002
- Astyanax tumbayaensis Miquelarena & Menni, 2005
- Astyanax tupi Azpelicueta, Mirande, Almirón & Casciotta, 2003
- Astyanax turmalinensis Triques, Vono & Caiafa, 2003
- Astyanax unitaeniatus Garutti, 1998
- Astyanax utiariti Bertaco & Garutti, 2007
- Astyanax validus Géry, Planquette & Le Bail, 1991
- Astyanax varzeae Abilhoa & Duboc, 2007
- Astyanax venezuelae L. P. Schultz, 1944
- Astyanax vermilion Zanata & Camelier, 2009
- Astyanax villwocki Zarske & Géry, 1999
- Astyanax xavante Garutti & Venere, 2009
- Astyanax xiru C. A. S. Lucena, J. B. Castro & Bertaco, 2013 [5]
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References
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- ↑ Froese, Rainer, and Daniel Pauly, eds. (2015). Species of Astyanax in FishBase. October 2015 version.
- ↑ Mirande, J.M., Jerep, F.C. & Vanegas-Ríos, J.A. (2013): Phylogenetic relationships of the enigmatic Carlastyanax aurocaudatus (Eigenmann) with remarks on the phylogeny of the Stevardiinae (Teleostei: Characidae). Neotropical Ichthyology, 11 (4): 747-766.
- ↑ Lua error in package.lua at line 80: module 'strict' not found.
- ↑ 4.0 4.1 4.2 4.3 4.4 Lua error in package.lua at line 80: module 'strict' not found.
- ↑ 5.0 5.1 5.2 Lucena, C.A.S.d., Castro, J.B. & Bertaco, V.A. (2013): Three new species of Astyanax from drainages of southern Brazil (Characiformes: Characidae). Neotropical Ichthyology, 11 (3): 537-552.
- ↑ Bertaco, V.A. (2014): Astyanax douradilho, a new characid fish from the rio Tramandaí system, southern Brazil (Characiformes: Characidae). Zootaxa, 3794 (3): 492–500.
- ↑ Ingenito, L.F.S. & Duboc, L.F. (2014): A new species of Astyanax (Ostariophysi: Characiformes: Characidae) from the upper rio Iguaçu basin, southern Brazil. Neotropical Ichthyology, 12 (2): 281-290.
- ↑ Oliveira, C.A.M., Abilhoa, V. & Pavanelli, C.S. (2013): Astyanax guaricana (Ostariophysi: Characidae), a new species from the rio Cubatão drainage, Paraná State, Southern Brazil. Neotropical Ichthyology, 11 (2): 291-296.
- ↑ Marinho, M.M.F., Camelier, P. & Birindelli, J.L.O. (2015): Redescription of Astyanax guianensis Eigenmann 1909 (Characiformes: Characidae), a poorly known and widespread fish from the Amazon, Orinoco and Guiana Shield drainages. Zootaxa, 3931 (4): 568–578.
- ↑ Camelier, P. & Zanata, A.M. (2014): A new species of Astyanax Baird & Girard (Characiformes: Characidae) from the Rio Paraguaçu basin, Chapada Diamantina, Bahia, Brazil, with comments on bony hooks on all fins. Journal of Fish Biology, 84 (2): 475–490
- ↑ Freitas, M.R.M., Lucinda, P.H.F. & Lucena, C.A.S. (2015): Redescription of Astyanax novae Eigenmann, 1911 (Teleostei: Characidae). Ichthyological Exploration of Freshwaters, 26 (2): 183-192.
- ↑ Lucena, C.A.S.d., Bertaco, V.A. & Berbigier, G. (2013): A new species of Astyanax from headwater streams of southern Brazil (Characiformes: Characidae). Zootaxa, 3700 (2): 226–236.
- ↑ Lua error in package.lua at line 80: module 'strict' not found.