Technische Universität Bergakademie Freiberg
Palaeontology and Stratigraphy
During the mid-1980s, structural geological field work in the up to 300 m thick red-bed sequence of the Permian Tiddas Basin, central Morocco, revealed the first occurrence of Palaeozoic tetrapod footprints in NW Africa. Preservational... more
During the mid-1980s, structural geological field work in the up to 300 m thick red-bed sequence of the Permian Tiddas Basin, central Morocco, revealed the first occurrence of Palaeozoic tetrapod footprints in NW Africa. Preservational ambiguity of part of the few published trace fossils and resulting taxonomic inconsistency necessitate a long-needed reanalysis of the type locality. Based on 110 recently collected specimens the revised tetrapod ichnofauna of the study area comprises tracks of Batrachichnus Woodworth, -Limnopus Marsh, 1894, Amphisauropus Haubold, 1970, Dimetropus Romer and Price, 1940, Varanopus Moodie, 1929 -Hyloidichnus Gilmore, 1927, and Dromopus Marsh, 1894 which can be referred to small and medium-sized temnospondyl, seymouriamorph, basal synapsid ('pelycosaur'), captorhinomorph, and araeoscelid trackmakers. This assemblage is interpreted to represent a late Early Permian (Artinskian-Kungurian) tetrapod ichnofauna. Due to the relative abundance of footprints assigned to semiaquatic tetrapods and sedimentological evidence for pronounced humid palaeoclimatic conditions we propose correlation of the track-bearing strata with the late Artinskian/early Kungurian wet phase. Regarding the quality, quantity, and peculiar diversity of its fossils, including the earliest record of seymouriamorph and advanced captorhinomorph tracks from the African continent, the Tiddas Basin could be of great importance for the reconstruction of late Early Permian terrestrial ecosystems in proximity to the interface of Euramerican and Gondwanan biotic provinces.
- by Hafid Saber and +2
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- Paleontology, Biostratigraphy
Oribatid mites (Acari: Oribatida) are very diverse and important detritivorous and fungivorous micro-arthropods in modern forest ecosystems. Although the fossil record of oribatid mites can be traced to the Early Devonian, the... more
Oribatid mites (Acari: Oribatida) are very diverse and important detritivorous and fungivorous micro-arthropods
in modern forest ecosystems. Although the fossil record of oribatid mites can be traced to the Early Devonian, the
paleoecology of oribatid mites during the deep geological past remains poorly understood. Remarkably good
preservation of tunnel networks in a permineralized coniferwood specimen is described fromthe Early Permian
of Germany. This fossil provides evidence for four aspects of oribatid mite feeding habits. First, there is preferred
consumption of the more indurated tissues from growth-ring cycles. Second, tracheids were targeted for
consumption. Third, feeding on tissues resulted in fecal pellet accumulations at the bottoms of tunnels. And
fourth, the absence of feeding on ambient decomposing fungi such as necroses and rots, but rather the processing
of pristine plant tissues, indicate the presence of a self-contained, microorganismic gut biota. These rather
specialized feeding habits allowed oribatid mites a prominent role in the decomposition of digestively refractory
plant tissues in Early Permian ecosystems.
in modern forest ecosystems. Although the fossil record of oribatid mites can be traced to the Early Devonian, the
paleoecology of oribatid mites during the deep geological past remains poorly understood. Remarkably good
preservation of tunnel networks in a permineralized coniferwood specimen is described fromthe Early Permian
of Germany. This fossil provides evidence for four aspects of oribatid mite feeding habits. First, there is preferred
consumption of the more indurated tissues from growth-ring cycles. Second, tracheids were targeted for
consumption. Third, feeding on tissues resulted in fecal pellet accumulations at the bottoms of tunnels. And
fourth, the absence of feeding on ambient decomposing fungi such as necroses and rots, but rather the processing
of pristine plant tissues, indicate the presence of a self-contained, microorganismic gut biota. These rather
specialized feeding habits allowed oribatid mites a prominent role in the decomposition of digestively refractory
plant tissues in Early Permian ecosystems.
- by Zhuo Feng and +2
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- Plant-Animal Interactions, Fossil Wood, Permian
Two poorly known trigonotarbids (Arachnida: Trigonotarbida), Eophrynus scharfi SCHARF, 1924 and Eophrynus ilfeldicus SCHARF, 1924 from the Lower Permian (Asselian) Ilfeld Basin in the Harz Region of Germany are redescribed. The latter is... more
Two poorly known trigonotarbids (Arachnida: Trigonotarbida), Eophrynus scharfi SCHARF, 1924 and Eophrynus ilfeldicus SCHARF, 1924 from the Lower Permian
(Asselian) Ilfeld Basin in the Harz Region of Germany are redescribed. The latter is transferred to the genus Aphantomartus POCOCK, 1911. The former resembles members of Lissomartidae, but given its incompleteness it is assigned here to Trigonotarbida incertae sedis. A phalangiotarbid (Arachnida: Phalangiotarbida) from Ilfeld described as Opiliotarbus elongatus (SCUDDER, 1890) represents the youngest record of this group. With respect to the genus Opiliotarbus POCOCK, 1910, Architarbus
hoffmanni GUTHORL, 1934 - one of three names erected
for a lost specimen from the Upper Carboniferous of the
Saar region of Germany - represents the senior synonym of
Opiliotarbus kliveri WATERLOT, 1934 and Goniotarbus sarana
GUTHORL, 1965. The Lower Permian fossil Rhabdotarachnoides simoni Haupt, 1957 from the Rotliegend of the Thuringian Forest Basin, Germany was described as an arachnid and tentatively referred to Opiliones. Since it is identified here as a coalified plant remain, this species is regarded as nomen dubiurn.
(Asselian) Ilfeld Basin in the Harz Region of Germany are redescribed. The latter is transferred to the genus Aphantomartus POCOCK, 1911. The former resembles members of Lissomartidae, but given its incompleteness it is assigned here to Trigonotarbida incertae sedis. A phalangiotarbid (Arachnida: Phalangiotarbida) from Ilfeld described as Opiliotarbus elongatus (SCUDDER, 1890) represents the youngest record of this group. With respect to the genus Opiliotarbus POCOCK, 1910, Architarbus
hoffmanni GUTHORL, 1934 - one of three names erected
for a lost specimen from the Upper Carboniferous of the
Saar region of Germany - represents the senior synonym of
Opiliotarbus kliveri WATERLOT, 1934 and Goniotarbus sarana
GUTHORL, 1965. The Lower Permian fossil Rhabdotarachnoides simoni Haupt, 1957 from the Rotliegend of the Thuringian Forest Basin, Germany was described as an arachnid and tentatively referred to Opiliones. Since it is identified here as a coalified plant remain, this species is regarded as nomen dubiurn.
- by Jason A. Dunlop and +2
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- Systematics (Taxonomy), Arachnida, Permian, Trigonotarbida
The Kinney Brick Quarry is a world famous Late Pennsylvanian fossil Lagerstätte in central New Mexico, USA. The age assigned to the Kinney Brick Quarry (early-middle Virgilian) has long been based more on its inferred lithostratigraphic... more
The Kinney Brick Quarry is a world famous Late Pennsylvanian fossil Lagerstätte in central New Mexico, USA. The age assigned to the Kinney Brick Quarry (early-middle Virgilian) has long been based more on its inferred lithostratigraphic position than on biostratigraphic indicators at the quarry. We have developed three datasets --stratigraphic position, fusulinids and conodonts-that indicate the Kinney Brick Quarry is older, of middle Missourian (Kasimovian) age. Our detailed local lithostratigraphic studies coupled with regional stratigraphic investigations indicate the Kinney Brick Quarry is in the Tinajas Member of the Atrasado Formation, so it is stratigraphically lower than suggested by previously published maps. A laterally extensive fusulinid-bearing limestone a few meters below the level of the Kinney Brick Quarry yields an early-middle Missourian fusulinid assemblage consisting of Tumulotriticites cf. T. tumidus and species of Triticites: T. cf. T. planus, T. cf. T. myersi and T. ex gr. T. ohioensis. The Kinney conodont fauna is characterized by Idiognathodus corrugatus and I. cherryvalensis, which suggest an assignment to the Idiognathodus confragus Zone of the North America Midcontinent region (Dennis cyclothem; middle Missourian). Nonmarine biostratigraphic indicators at the Kinney Brick Quarry indicate either an imprecise age (Late Pennsylvanian: megaflora) or a slightly younger age (late Kasimovian-early Gzhelian: blattids) than do stratigraphic position and marine microfossils. The well-established age of Kinney on the marine timescale thus can be used to better calibrate the nonmarine biostratigraphy. So, the insect biozonation of the Euramerican continental basins, which was calibrated to the so-called regional West European Carboniferous stages by macrofloras and to the global marine scale by sparse, ambiguous isotopic ages, can now be linked directly to the marine conodont zonation.
- by Joerg W. Schneider and +1
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At Carrizo Arroyo southwest of Albuquerque, New Mexico, USA, an approximately 105-m-thick section of upper Paleozoic clastic and carbonate rocks yields extensive fossil assemblages of marine and nonmarine origin. Most of the section at... more
At Carrizo Arroyo southwest of Albuquerque, New Mexico, USA, an approximately 105-m-thick section of upper Paleozoic clastic and carbonate rocks yields extensive fossil assemblages of marine and nonmarine origin. Most of the section at Carrizo Arroyo belongs to the Red Tanks Member of the Bursum Formation,~100m thick and mostly variegated shale, mudstone and siltstone of nonmarine origin, intercalated with some beds of limestone and shale of marine origin. Red Tanks Member fossils include palynomorphs, charophytes, plant megafossils, non-fusulinid foraminifers, fusulinids, bryozoans, brachiopods, gastropods, bivalves, nautiloids, eurypterids, ostracods, syncarid crustaceans, conchostracans, insects and some other arthropods, echinoids, crinoids, conodonts, fish ichthyoliths and bones of amphibians and reptiles. At stratigraphic levels 43m and 68m above the base of the section are Lagerstätten of plants, insects, crustaceans, eurypterids and other fossils that form unique Late Paleozoic nearshore arthropod assemblages. Most of the fossil groups from the Red Tanks Member have been used to support diverse placements of the Pennsylvanian-Permian boundary at Carrizo Arroyo. We review previous placement of the Pennsylvanian-Permian boundary in the Carrizo Arroyo section and present newly collected insect and conodont data. The insects indicate that the two Lagerstätten in the Red Tanks Member are of early Asselian age. The new conodont data include the presence of Streptognathodus virgilicus in the uppermost part of the Atrasado Formation, which constrains its age to the middle to upper part of the Virgilian and to a comparable position in the Gzhelian. The only biostratigraphically-significant conodont assemblage in the Red Tanks Member comes from a marine horizon near the middle of the member, and the assemblage is probably equivalent in age to the Midcontinent Streptognathodus nevaensis Zone, of early to middle Asselian in age. A significant amount of latest Pennsylvanian to earliest Permian time apparently is not represented by rock record at the Carrizo Arroyo section, most likely at a major disconformity at the top of the Atrasado Formation and smaller ones at the bases of depositional sequences in the lower part of the Red Tanks Member. Conodont biostratigraphy provides compelling evidence that Bursum Formation deposition was not simply driven by glacio-eustatic cyclicity, but in this area it was partly overprinted by local tectonics..
Upper Pennsylvanian (Missourian) strata of the Tinajas Member of the Atrasado Formation in the Cerros de Amado, Socorro County, include an unusual lacustrine deposit, the Tinajas locality. This locality yields a diverse fossil assemblage... more
Upper Pennsylvanian (Missourian) strata of the Tinajas Member of the Atrasado Formation in the Cerros de Amado, Socorro County, include an unusual lacustrine deposit, the Tinajas locality. This locality yields a diverse fossil assemblage of plants, crustaceans, insects, a cnidarian, molluscs and fishes from laminar black shale that contains a prolific conchostracan assemblage. A lacustrine origin of the black shale is supported by the abundance of conchostracans; presence of darwinuloid (freshwater) ostracods, land plants and insect wings; lack of marine indicators; and the lithology and geometry of the deposit. Overlying gray shales are full of Dunbarella, which indicates the onset of nearshore, marginal marine conditions. Indeed, the stratigraphic succession at the Tinajas locality is analogous to a classical cyclothem. Thus, fluvial/deltaic deposits grade upward into a coastal lake that is then transgressively overlain by marginal marine and fully marine sediments. The preservation of the lacustrine sediments at the Tinajas locality was a local, unique event on the Late Pennsylvanian coastline in central New Mexico, as no other occurrence of a similar paleoenvironment is known. FIGURE 1. Measured stratigraphic section at the Tinajas locality. See text for description of lithographic units.
An insect zonation with a time resolution of 1.5-2 Ma for Late Pennsylvanian to Early Permian (Kasimovian to Artinskian) non-marine deposits is presented. The zonation is based on the directed morphogenetic evolution of colour pattern in... more
An insect zonation with a time resolution of 1.5-2 Ma for Late Pennsylvanian to Early Permian (Kasimovian to Artinskian) non-marine deposits is presented. The zonation is based on the directed morphogenetic evolution of colour pattern in the forewings of the blattid (cockroach) family Spiloblattinidae. This evolution is observed in lineages of succeeding species of three genera. All three genera are widely distributed in the palaeo-equatorial zone from Europe to North America, that is, in the Euramerican biota province. Increasing reports of spiloblattinid zone species in conodont-bearing, interfingered marine/continental strata of North American Appalachian, Mid-Continent and West Texas basins could be the key to direct biostratigraphical correlations of pure continental profiles, as are present in the most parts of the Hercynides, to the global marine scale.
- by Ralf Werneburg and +1
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- Earth Sciences, North America, North American
A well-preserved vertebral column from the Late Permian of Southern France (Lopingian, La Lieude Formation, Lodève Basin) is described. It is composed of diplospondylous vertebrae and is most comparable with the temnospondyl... more
A well-preserved vertebral column from the Late Permian of Southern France (Lopingian, La Lieude Formation, Lodève Basin) is described. It is composed of diplospondylous vertebrae and is most comparable with the temnospondyl Tupilakosaurus previously known from the Early Triassic of Greenland and Russia. This new specimen therefore represents the earliest occurrence of a diplospondylous tupilakosaur, and extends the geographic range of the group to Western Europe. It is an aquatic temnospondyl that used the anguilliform undulatory mode of swimming.
- by Ralf Werneburg and +1
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- Evolutionary Biology, Geology, Western Europe, Ecology
The late Paleozoic/early Mesozoic encompasses the formation and beginning of the breakup of Pangea, the transition from the Hercynian to the Alpidic geotectonic supercycle, as well as the transition from an icehouse to a greenhouse... more
The late Paleozoic/early Mesozoic encompasses the formation and beginning of the breakup of Pangea, the transition from the Hercynian to the Alpidic geotectonic supercycle, as well as the transition from an icehouse to a greenhouse climate. It covers the gradual crossover from the Palaeozoic biota and ecosystems to those of the Mesozoic as well as the P/T mass extinction. Late Paleozoic/early Mesozoic sediments are important source and reservoir rocks of fuels. For exploration tasks as well as for the understanding of global environmental changes and of biotic and geotectonic events too, an exact time control is essential.
- by Ralf Werneburg and +3
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The giant arthropod Arthropleura was a common member of the late Paleozoic continental biota of paleo-equatorial biomes for more than 35 million years, from the Early Carboniferous late Visean (FOD; Middle Mississippian,... more
The giant arthropod Arthropleura was a common member of the late Paleozoic continental biota of paleo-equatorial biomes for more than 35 million years, from the Early Carboniferous late Visean (FOD; Middle Mississippian, Asbian/Brigantian) up to the Early Permian lower Rotliegend (LOD = ?LAD; Asselian). In Upper Pennsylvanian red beds in Cañon del Cobre of northern New Mexico, trackways of Arthropleura are present in strata that also yield body fossils of the amphibian Eryops. We review the Arthropleura tracksite from Cañon del Cobre, New Mexico, as well as other tracksites of this animal and arthropleurid/eryopid associations in order to better interpret the paleoenvironmental preference and the paleobiology of Arthropleura. This review supports the conclusion that Arthropleura was well adapted to alluvial environments of ever wet humid to seasonally dry and semihumid climates. Preferred habitats of semi-adult and adult Arthropleura were open, vegetated, river landscapes. They co-occurred in these habitats with semi-aquatic eryopid amphibians and terrestrial pelycosaurs.
- by Ralf Werneburg and +2
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An insect zonation with a time resolution of 1.5-2 Ma for Late Pennsylvanian to Early Permian (Kasimovian to Artinskian) non-marine deposits is presented. The zonation is based on the directed morphogenetic evolution of colour pattern in... more
An insect zonation with a time resolution of 1.5-2 Ma for Late Pennsylvanian to Early Permian (Kasimovian to Artinskian) non-marine deposits is presented. The zonation is based on the directed morphogenetic evolution of colour pattern in the forewings of the blattid (cockroach) family Spiloblattinidae. This evolution is observed in lineages of succeeding species of three genera. All three genera are widely distributed in the palaeo-equatorial zone from Europe to North America, that is, in the Euramerican biota province. Increasing reports of spiloblattinid zone species in conodont-bearing, interfingered marine/continental strata of North American Appalachian, Mid-Continent and West Texas basins could be the key to direct biostratigraphical correlations of pure continental profiles, as are present in the most parts of the Hercynides, to the global marine scale.
- by Ralf Werneburg and +3
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2010. New hybondontoid shark from the Permocarboniferous (Gzhelian-Asselian) of Guardia Pisano (Sardinia, Italy). Acta Palaeontologica Polonica 55 (2): 241-264.
Rend. Soc. Paleont. hai, 1 (2002): 169-176 Early Permian continental biota from Southeastern Sardinia (Ogliastra and Gerrei) Pierre Freytet, Jean Galtier, Ausonio Ronchi, Jörg W. Schneider, Andrea Tintori and Ralf Werneburg P. Freytet -... more
Rend. Soc. Paleont. hai, 1 (2002): 169-176 Early Permian continental biota from Southeastern Sardinia (Ogliastra and Gerrei) Pierre Freytet, Jean Galtier, Ausonio Ronchi, Jörg W. Schneider, Andrea Tintori and Ralf Werneburg P. Freytet - Laboratoire de ...
- by Joerg W. Schneider and +1
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