Inventions, innovations and the origins of spelt wheat

Vegetation History and Archaeobotany https://doi.org/10.1007/s00334-023-00978-2 ORIGINAL ARTICLE Inventions, innovations and the origins of spelt wheat Jutta Lechterbeck1 · Tim Kerig2 Received: 26 April 2023 / Accepted: 17 August 2023 © The Author(s) 2023 Abstract What turns an invention into an innovation? How, if at all, might we observe this process archaeologically? Loosely put, new varieties of plants or animals might be considered as inventions (whether from deliberate breeding or by chance), but ones that are only taken up by humans more systematically as innovations when certain social, demographic, economic and environmental factors encourage such take-up. The archaebotanically-observed history of spelt wheat (Triticum spelta) is an interesting case in this respect. Prior to 3000 BCE, spelt is occasionally found in very small amounts at sites in eastern Europe and south-west Asia, but is usually considered to be a crop weed in such contexts, rather than a cultivar. However, rather suddenly across Central Europe ~ 3000−2500 BCE spelt appears more consistently at multiple Chalcolithic and especially Bell Beaker sites, in quantities which suggest a shift to its use as a deliberate crop. By the full-scale Bronze Age in this region, spelt becomes one of the major crops. This paper discusses this Central European process in greater detail via macro-botanical evidence. It argues that demographic factors during the Neolithic may have inhibited the spread of Asian spelt into central Europe, and that while small amounts of local European spelt were probably present earlier on, it was only at the very end of the Neolithic, in tandem with human population increases and major technological changes such as the introduction of the plough that spelt was taken up as a cultivar. In particular, a shift by some communities in the region ~ 3000−2500 BCEo more extensive (and sometimes plough-enabled) agricultural strategies may have favoured deliberate cultivation of spelt on less productive soils, given this variety’s relative robustness to harsher conditions. In other words, a combination of conditions was necessary for this innovation to really take hold. Keywords Triticum spelta · Innovation process · Neolithic · Demography · Evolutionary archaeology Introduction In the following, we make use of innovation theory to conceptualize the significance of spelt within the economic and social realm. The origin of spelt is an evolutionary biological process; its integration into agricultural practices and systems turns genetics into economics. The emergence of new varieties of plants and animals, under breeding conditions or by chance, can be considered from an economic point of view as an invention in the sense Communicated by F. Antolín. * Jutta Lechterbeck jutta.lechterbeck@uis.no 1 Museum of Archaeology, Universitet Stavanger, Peder Klows gate 30A, 4002 Stavanger, Norway 2 Institute for Prehistory and Early History, Christian-Albrechts-University Kiel, ChristianAlbrechts-Platz 4, 24118 Kiel, Germany that those varieties are new, and their purpose or usefulness is unknown. The success of such an invention is called innovation, which depends on many factors, societal as well as environmental. In this paper we would like to show which archaeologically comprehensible factors turn such an invention into an innovation; we use the origin, early spreading and final establishment of spelt as an example. Gabriel Tarde (1843–1904) and Joseph Alois Schumpeter (1883–1950) have each put forward the basics of a theory of innovation although neither of them uses the word itself. There is one feature that connects both the social and economic science concepts of innovation and that is the separation of invention from innovation. While the sociologist Tarde (1890) is interested in any novelty or improvement in any kind of social phenomenon, Schumpeter (1952) defines innovation neither psychologically nor sociologically but strictly economic. Schumpeter speaks of “Durchsetzung neuer Kombinationen“ (enforcement of new combinations) of the means of production. 13 Vol.:(0123456789) Vegetation History and Archaeobotany According to the widely accepted definition by Everett Rogers (1931–2004), innovation diffusion is “…the process by which (1) an innovation (2) is communicated through certain channels (3) over time (4) among the members of a social system” (Rogers 2003). Schumpeter (1934) defines innovations in the economic sphere as fresh combinations of new or existing knowledge, resources, equipment or other factors and he makes a distinction between, on the one hand, an invention, which might happen everywhere without any intent and, on the other hand, innovation as a process with an economic purpose. In fact, innovation is the process that carries inventions into society. Though invention and innovation are closely connected, both can stand alone: “Innovation is possible without anything we should identify as invention, and invention does not necessarily induce innovation” (Schumpeter 1939). If we see the invention as the origin of something new and the innovation as the disseminating process this concept can be successfully applied in archaeological contexts. The word “innovation” is often used in archaeological research (e.g. Müller 2013; O’Brien et al. 2014, for innovation theory in archaeology: Kerig 2016) but rarely clearly defined, with archaeologists tending to see it simply as the arrival of something new (e.g. Frieman 2021) and often used in connection with the notion of technological progress. In contrast, Shennan (2001) incorporated demography and cultural innovation where cultural innovations operate similar to genetic mutations, though unlike mutations, cultural traits can be transmitted by non-parents and horizontally by peers. In our understanding of innovation, the mutation would be analogous to the invention in the sense that some inventions might be attractive to imitate, others less so and some would not make any difference at all. The innovation process would then be the passing and spreading of the invention in the transmission process. Shennan’s model (2001) shows that larger populations where cultural traits are passed on by a combination of vertical and oblique (horizontal) transmission have a very major advantage to smaller ones. Important for the understanding are the two principal ways of increasing the population for transmission, one is demographic growth and the other the contact with another, formerly equally isolated population. Shennan uses the model to explain the emergence of novel traits in human culture. Though his model is able to show that transmission can only take place in phases of population growth it does not explain the unconnected occurrence of novel traits at certain times and places. The analogue of mutation and invention however is likewise not wholly convincing – that is because in small populations genetic mechanisms often prevent the phenotypic emergence of deleterious mutations (e.g. Robinson et al. 2018). It goes without saying that deleterious inventions have succeeded and continue to do so. 13 Inventions can come out of thin air or “from god”, which means they occur arbitrary and might or might not be of future use. Also, to continue the mutation analogue, they might be “pre-adaptations” and prove to be useful ex post. It seems likely that with the number of people also the number of ideas and the intellectual exchange increase and thus more inventions occur in larger populations. Population density and cultural transmission are two factors that seem closely connected. “Cultural transmission is the process through which cultural elements, in the form of attitudes, values, beliefs, and behavioural scripts, are passed onto and taught to individuals and groups” (Taylor and Thoth 2011). From an archaeological perspective, one might add that also the knowledge about the sources of raw materials and foreign goods, the mastering of technologies by crafts are passed through cultural transmission - cultural transmission is the inheritance of practices and knowledge. Necessary conditions for cultural transmission are contact and communication: Nothing can be passed on without people actually meeting each other. Population density becomes the crucial factor as innovations are easier passed on under a high encounter rate. Grove (2016) however proposes that individual encounter rates depend not only on population density but on the product of density and mobility: highly mobile but low-density populations might have the same encounter rates as low mobility, high density populations. Walker et al. (2021) carried this assumption to inter-community encounters but found only weak supporting evidence. Both these models are dealing with hunter-gatherers, which are thought to be per se more mobile than Neolithic farmers are. Thus, in farming communities, population density might play a more important role in cultural transmission processes than mobility. The invention of spelt ‑ where and when In Central Europe Triticum spelta L. (spelt) occurs as a new cultivar rather suddenly in disparate locations at the very end of the Neolithic, in Bell Beaker and contemporary contexts. In Switzerland (Akeret 2005; Jacomet 2008) and South–West Germany (Lechterbeck et al. 2014a), it is found in Bell-Beaker contexts, in Lauda-Königshofen (E. Fischer pers. comm.) in a corded ware context, but contemporary to the sites in South-West Germany and Switzerland and it was also found at Vojens, Brødrene Gram (Robinson 2003) in Denmark in Bell-Beaker contemporary contexts. The origins of T. spelta are somewhat mysterious. Two hypotheses as to its origin are discussed: in Southwest Asia, but then it is not known by which route it came to Central Europe, or spontaneously on site from local wheat species, which would explain its sparse occurrence at disparate locations. Newer Vegetation History and Archaeobotany analyses prove that T. spelta is polyphyletic (Dvorak et al. 2012), therefore both hypotheses might be true. In the beginning perhaps being a crop weed, T. spelta becomes later one of the main crops in the Early and Middle Bronze Age (EBA, MBA) in western Central Europe, the Alps and their foreland. From there it apparently spread to Scandinavia, to Southern France, to the Pannonian basin, and Greece (Stika and Heiss 2013). Though, according to Stika and Heiss (2013) it was not introduced to the Iberian Peninsula or to Italy south of the Po valley. But T. spelta has been cultivated later at least on the Northern Iberian peninsula in the Iron Age (Tereso et al. 2013; Seabra et al. 2018; Peña-Chocarro et al. 2019). T. spelta is a hexaploid hulled wheat. It is widely accepted that Asian spelt originated via hybridization of hulled emmer and goat grass (Aegilops tauschii Coss.). Newer genetic analyses prove however, that European T. spelta originated by hybridisation of a free threshing hexaploid wheat and hulled emmer (Dvorak et al. 2012) thus Asian and European spelt have different origins. However, it is not possible to prove genetically whether Asian spelt migrated to Europe during the Neolithic or whether European spelt originated on site. Up to now, aDNA analyses on prehistoric T. spelta are missing due to the lack of suitable material, therefore it is worthwhile to have a closer look at its early distribution and history. Triticum turgidum ssp. dicoccon (Schrank ex. Schübl) Thell., cultivated emmer, one of the parents of T. spelta, is one of the founder crops brought to Northwestern Europe with the first farmers (Bakels 2014). Triticum turgidum ssp. dicoccoides (Asch. & Graebn.) Thell., wild emmer, is distributed all over the fertile crescent (Fig. 1) but with a focus in the western part and it is thus not surprising that cultivated emmer occurs there very early, already around 10,500 BP (Zohary et al. 2012). Aegilops tauschii, the other parental species of Asian spelt, is almost exclusively distributed east of the 40° longitude (Fig. 1) and has its centre of distribution along the southern shores of the Caspian Sea and in Azerbaijan (Kole 2011). The area where both species occur is restricted to north-eastern Syria, South–Eastern Turkey and northern Iraq which is the area where T. spelta could have possibly originated. Asian spelt cannot occur spontaneously on location because A. tauschii is not present everywhere. It can only arise under cultivation of emmer wheat in the designated area of origin and then spread as a crop weed. The parent species for European spelt are a free threshing hexaploid wheat and cultivated emmer. Cultivated emmer occurs all over Central Europe since the beginning of the Neolithic. Confirming free threshing, hexaploid wheat in the archaeobotanical record however is problematic: the grains look similar to grains of tetraploid naked wheat; therefore, these are reasonably summarised as T. aestivum/ durum/turgidum in archaeobotanical reports. A safe differentiation is only possible if chaff is present (Jacomet 2006). Grains of free threshing wheat are reported all over Europe (Colledge 2016) during the Neolithic but sites with chaff from hexaploid free threshing wheat are quite rare (Fig. 2). Colledge’s dataset covers only Central- and NorthwestEurope, but evidence from other areas, especially the Near East, is likewise sparse: while tetraploid free threshing wheat derives directly from cultivated emmer, has hexaploid free threshing wheat the same ancestry as Asian spelt. It might have originated in the same area, but all of the classic sites only report tetraploid wheat (Maier 1996) in the Neolithic. More recently, hexaploid wheat, both naked and hulled, could be proven genetically in Çatalhöyük in Turkey Fig. 1 Distribution areas of Triticum turgidum ssp. diccocoides (wild emmer) and Aegilops tauschii (goat grass) redrawn from Zohary et al. 2012 and Kole 2011. Cultivated emmer occurs first in areas where wild emmer is present and hybridization of goat grass and cultivated emmer can occur in these areas 13 Vegetation History and Archaeobotany Fig. 2 Triticum turgidum ssp. diccocon (cultivated emmer, grains and chaff) and finds of free threshing hexaploid wheat rachis fragments (taxon code TRIFTHR) in the Middle and Late Neolithic of Central Europe (data from Colledge 2016; for details of the sites see ESM) around 8,400 BP (Bilgic et al. 2016). Unlike T. spelta, chaff of naked hexaploid wheat is found in Neolithic contexts of different ages in Central Europe. Important in this context is Erkelenz-Kückhoven (Knörzer 1995, 1998), where botanical remains from an old Neolithic well contained rachis fragments of hexaploid naked wheat (Maier 1996). This find shows that naked hexaploid wheat was present in European cereal spectra from the beginning of the Neolithic if only very sparse. Not morphologically but genetically detected was naked hexaploid wheat at the lake site Zürich/Mozartstrasse for around 3900 BC (Schlumbaum et al. 1998). Both Maier (1996) and Schlumbaum et al. (1998) refer to other sites in Southern Germany and in the Netherlands where Neolithic hexaploid naked wheat was found. It is however not present in Neolithic Eastern European contexts, though both T. spelta and T. aestivum grains are reported from Poland (Lityńska-Zając 2016) but due to the problem with the identification of grains that might be misidentifications. European spelt, unlike Asian spelt, can occur spontaneously through hybridisation on spot, as both parent species have been present throughout the Neolithic in Central Europe. T. spelta macroremains are difficult to identify in the archaeological record, the grains look very much like emmer grains, especially when charred in the spikelets (Jacomet 2006) and chaff can be confused with certain barrel-shaped 13 Aegilops species esp. A. cylindrica Host (Nesbitt and Samuel 1996). Reliable identification of T. spelta is only possible when spikelet forks and/or glume bases are present. For this paper, only sites with reliable T. spelta finds were taken into consideration. Sites where only grains are reported, are not discussed here. Early finds of T. spelta (Fig. 3), i.e. finds before the Bell Beaker period, are reported from the identified possible area of origin for the 7th-8th millennium BP (Zohary et al. 2012 and references therein). It has been questioned whether T. spelta spread from there to Central Europe and if so, by which route. The most obvious route would have been over Turkey and the Balkans, but there are no reliable finds of T. spelta from Turkey except from Erbaba (van Zeist and Buitenhuis 1983). Nesbitt (2001) doubts the presence of T. spelta in near eastern archaeobotanical assemblages at all and ascribes isolated occurrences to misidentifications. In the light of early T. spelta finds from Transcaucasia it was suggested that it originated in Transcaucasia or Iran and then spread to Europe to arrive several millennia after the introduction of agriculture (Andrews 1964; van Zeist 1976). However, T. spelta is also sparse, if not absent as Nesbitt (2001) states, in Transcaucasian Neolithic assemblages. Those occurrences which are reported in Zohary et al. (2012) and in Kohler-Schneider and Caneppele (2009) Vegetation History and Archaeobotany Fig. 3 Triticum spelta (spelt) finds before the End Neolithic (data from Zohary et al. 2012 and Kohler-Schneider and Caneppele 2009) suggest that T. spelta finds before 3000 BCE are recorded no more westerly than eastern Austria (Fig. 3). It is very difficult to draw conclusions as to its origin and spread from sparse and probably doubtful occurrences. The dates do not show a clear tendency for the spreading of T. spelta from the area of origin via either the Balkan or the Transcaucasian route. It has apparently never been a crop in the Neolithic therefore its occurrence is purely accidental. Radiocarbon dates for botanical macro remains from Hundsteig, Eastern Austria (Kohler-Schneider and Caneppele 2009) where T. spelta occurs in quite high quantities, date approximately between 3330 and 2900 BCE. The wide range of the date is due to a plateau in the calibration curve. The site of Hundsteig (Pieler 2001) belongs to the Jevišovice culture. In contrast to the roughly contemporaneous Baden culture, the Jevišovice culture features strikingly small settlements situated mainly on hilltops and above steep river valleys (Kohler-Schneider and Caneppele 2009). They were possibly part of a copper trading network. More westward finds of T. spelta are not known before the End-Neolithic, namely Bell-Beaker and contemporary contexts. In contrast to more easterly finds are those from Hundsteig quite substantial indicating that here T. spelta might have been a crop. The T. spelta finds from Bell Beaker and contemporary contexts (Fig. 4) come from disparate locations, though one might define two main regions, (i) lowland Switzerland and southwestern Germany and (ii) Denmark. The Bell Beaker T. spelta finds occur in regions where both parent species of European spelt are present in the Late Neolithic (Fig. 2) therefore it is highly probable that T. spelta originated onsite. In Eastern Austria however, no secure hexaploid free threshing wheat (i.e. rachis fragments) is recorded for the Neolithic – one of the parent species of European spelt seems to be missing from the area. It is therefore probable that the find from eastern Austria represents the westernmost find of Asian spelt in Europe so far and that the finds from the EndNeolithic are the first records of European spelt. Innovation processes behind spelt The appearance of T. spelta involves genetic mutation that can be seen as an invention occurring relatively suddenly. In contrast, its spread and establishment within the wider crop mix can be seen as an innovation process. The choice of which plant to cultivate is a cultural one, and the establishment of a crop as widely available, reified as a separate category in people’s taxonomies and routinely used depends on many factors, cultural as well as environmental. In nonliterate communities only such innovations can survive that are actually in use and/or passed on (Kerig 2016). In the last couple of decades, demographic reconstruction has returned to being a key agenda in archaeological research (e.g. Hassan 1981; Chamberlain 2006; Renfrew 2009; Zimmermann et al. 2009; Shennan 2013) and major 13 Vegetation History and Archaeobotany Fig. 4 Secure T. spelta finds from Bell Beaker and contemporary contexts (data from Akeret 2005; Lechterbeck et al. 2014a; Colledge 2016) efforts to assess population densities and demographic developments in the European Neolithic have been made. In the course of the EUROEVOL project, T. Kerig and A. Bevan took digitised versions of the Buchvaldek et al. (2007) culture area atlas as a rough proxy for settlement density (Fig. 5). These reflect which areas in Europe archaeologists assign to cultures. Figure 5a shows the situation for the time window 3400−3200 BCE. For this time window, 54.5% of the area have been assigned to a culture. Central Europe seems to have been quite densely populated, but there is almost a “culture free corridor” which separates Western and Eastern Europe. This might be due to research- or actual settlement gaps. Many of the empty spaces are at least mountainous regions or otherwise unfavourable areas like marshland, which were most probably not settled throughout the whole of the Neolithic (Fig. 6). Eastern Austria, where Neolithic T. spelta was found, is well connected to the east but there are unassigned regions to the west and north. That leads to the assumption that cultural contacts would have been preferably to the east. It does not mean, however, that there has not been any cultural exchange whatsoever to the west. Elements of the Baden culture for example are found in Horgen contexts at Lake Constance (Köninger et al. 2001; de Capitani 2002) thus evidencing cultural contact but they remain exceptions. T. spelta however was not one of those elements. There are 13 several well-analysed Horgen pile dwelling sites with excellent preservation conditions for botanical macro remains but T. spelta was never found in them. Figure 5b shows the situation in the time window 2700 −2450 BCE. Even fewer areas are attributed to archaeological cultures and many of them are isolated, indicating fewer cultural contacts between settled areas. Shortly after that period, T. spelta finds occur in disparate locations, and we would argue that the “invention” part of the T. spelta story took place at this time of assumed low population density. Two regional studies comparing the vegetational record of pollen analysis and demographic data derived from radiocarbon dates from the Neolithic to the BA were carried out in the areas where T. spelta occurs first in Bell Beaker and contemporary contexts (Lechterbeck et al. 2014b; Feeser et al. 2019). Lechterbeck et al. 2014b investigated the correlation of land use and demographic development in the Western Lake Constance area. Since 2014, the methodology for the reconstruction of population fluctuations was improved (Timpson et al. 2014; Crema 2022), therefore, the analysis was updated for the current paper. No additional radiocarbon dates were added. The radiocarbon density curve (SPD: sum probability distribution of 14C dates) for the region (Fig. 7) shows a rapid decline from the late Vegetation History and Archaeobotany Fig. 5 Maps of Central Europe, digitized from Buchvaldek et al. 2007. They depict the areas assigned to a culture in two different Neolithic time windows. The colours correspond to different Neolithic cultures in the area. Here only the assignment of a culture to a region is of interest not the culture as such Fig. 6 Overlay of all areas persistently attributed to Neolithic cultures. The colours display the number of times a region falls within a culture area, out of eight Neolithic phases Neolithic towards a minimum in the first part of the EBA and then a steep rise in the second part. This correlates with phases of abandonment and increased land use, respectively. Here indeed the “invention” of T. spelta happened in a phase of major demographic decline whereas its “innovation”, i.e. the spreading and establishment happened in a phase of demographic growth. Feeser et al. 2019 made a comparison of pollen data and radiocarbon derived demographic data for Northern Germany and Southern Jutland. Though the picture is not as 13 Vegetation History and Archaeobotany beaker do not occur in the same regions nor at the same time. Here, corded ware land use is forest based with a higher amount of wild food resources, evidence for open land is missing, it resembles the farming technique of the younger Neolithic. But there is a continuity between Bell Beaker and EBA lifestyles: the Bell Beaker phase resembles BA land use with indicators for open fields, grasslands and the cultivation of T. spelta. Lechterbeck et al. 2014a concluded that the Bell Beaker economy is the first stage of production typical for the EBA in the regions north of the Alps. It seems that T. spelta is only part of a whole package of inventions that became intrinsic in the BA. Conclusions Fig. 7 Radiocarbon density curves from the Western Lake Constance area. The black bar represents the radiocarbon dates for the Bell Beaker site Welschingen-Guuhaslen where T. spelta was found clear as in the Lake Constance area, also they record a demographic decline between 2400 and 2300 BCE. This coincides with the dating of house V on the site of Brødrene Gram, Vojens, Danmark to 2460 −2200 BCE where secure finds of T. spelta where made (Robinson 2003). Apparently, also in this area the first occurrence of T. spelta coincides with a demographic bust. The EBA is connected to a demographic boom in both regions. So far, the establishment of T. spelta fits Shennan’s model of transmission. Whether the “invention” of T. spelta also took place in a period of demographic bust in Cortaillod sur Les Rochettes cannot be decided as no analyses of SPD and pollen data for the region exist, but it is definitely an isolated occurrence. We cannot answer the question why T. spelta did not succeed earlier in the Neolithic, but some assumptions can be made. In a case study, Lechterbeck et al. (2014a) stated a discontinuity between corded ware and Bell Beaker lifestyles – in the Western Lake Constance area corded ware and bell 13 The evidence so far allows to assume that T. spelta originated in the Near-East sometime in the early Neolithic but did not migrate further west into Central Europe than Eastern Austria in the course of the Neolithic and it had never been a main crop. Only at the very end of the Neolithic at the transition to the BA it becomes a major crop in some disparate locations. In the EBA, secure T. spelta finds occur in sites close to those sites where it was first cultivated in the Neolithic (Fig. 8). T. spelta is a polyphyletic cereal and genetic analyses revealed that it originated twice in the course of prehistory, once in the Middle East as Asian spelt from cultivated emmer and Aegilops tauschii and a second time as European spelt from hexaploid free threshing wheat and emmer. Asian spelt cannot occur spontaneously, as one of the parent species is not present in European Neolithic contexts. Its sparse occurrence in Transcaucasia and the Balkan seems to indicate that it was a crop weed under emmer wheat cultivation and was not recognised as a cultivar. Early finds of T. spelta remain doubtful but it could be made probable that Asian spelt reached eastern Austria by the young Neolithic. The exceptional Jevišovice site of Hundsteig has substantial amounts of T. spelta finds, so here it might already have been recognised as a crop. The pattern of settled areas at that time suggests cultural contacts and exchange rather to the east. Between the early finds and the occurrences of T. spelta in Central Europe there is a time lag and, probably even more important, a large geographical distance. Also, the occurrences in Central Europe are substantial and suggest that T. spelta has been a crop rather than a weed. European spelt can occur spontaneously by hybridisation and it could be shown that both parent species were present in the Late Neolithic in the areas where it is later found in the Bell Beaker period. It is highly probable that T. spelta occurred during the whole of the Neolithic as a crop weed in Western and Central Europe but was not favoured as a cultivar. Under Vegetation History and Archaeobotany Fig. 8 Secure T. spelta occurrences in the Early Bronze Age (data from Colledge 2016; for details of the sites see ESM) the small scaled, intensive agricultural practices of the Neolithic T. spelta was not attractive. Only the agricultural changes at the end of the Neolithic and the beginning of the BA leading to more extensive arable farming made T. spelta an attractive cultivar as it gives good yields on less fertile soils and under a broad range of climatic conditions. This might also have levelled out the disadvantages of dehusking. It could be shown that the first cultivation of T. spelta in Central Europe occurred in a period of demographic bust. The occurrence of T. spelta can be seen as an “invention” (sensu Schumpeter) and the demographic boom at the beginning of the BA launched an innovation process that in the end established T. spelta as a crop. The establishment of T. spelta in Central Europe during the course of the BA coincides with a change in subsistence strategies: for the first time, large fields were worked with ploughs, the first meadows occur, the shift from two aisled to three aisled houses in the North European plain and parts of Scandinavia (Harding 2000; Armstrong Oma 2016) hints to the stabling of animals which made dung for manuring available, to name only a few examples. T. spelta is a relatively undemanding plant with respect to soil quality and gives good yields even under harsher conditions. This might have been an advantage under a more extensive land use strategy on less productive soils. Finally, it is worth briefly mentioning the later history of T. spelta, which continues to be innovative and impactful. A range of important changes occur to dominant mixes of cereal species present in Europe and the Mediterranean during the 1st millennium BCE which was at least in part due to the increasing scale of market economies, bulk grain shipment and growing taste in certain regions for breads over porridges (for a summary see Bevan 2019, pp 130–134). Amongst these changes, that occur several millennia after our period focus here, there is shift towards further increased use of T. spelta wheat in temperate Europe (from the Alps northwards and also the Atlantic fringe, but not further south, potentially linked to specific above-ground storage practices such as small poster granaries, see Sigaut 1989; Mills 2006). The twin attraction of T. spelta in this later phase is that its substantial glumes provide a certain hardiness in colder and wetter parts of Europe’s mid-latitudes, but also that its grain is glutinous enough to produce a bread, whilst also being good for other traditional foods. T. spelta continued to be a very important component in diets after the end of the Roman Empire and into the Early Medieval period (Devroey and van Mol 1989). 13 Vegetation History and Archaeobotany Supplementary information The online version contains supplementary material available at https://doi.org/10.1007/s00334-023-00978-2. Acknowledgements Our sincere thanks to Atakan Guven for help digitising the Buchvaldek et al. (2007) cultural atlas, to Andy Bevan, UCL, for suggestions, input and some remarks on the later history of spelt and to two anonymous reviewers whose comments and suggestions greatly improved the paper. The work of T. Kerig was funded by the Deutsche Forschungsgemeinschaft (DFG, German Research Foundation) – Project-ID 290391021 – SFB 1266”. Funding Open access funding provided by University of Stavanger & Stavanger University Hospital. Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit http://creativecommons.org/licenses/by/4.0/. References Akeret Ö (2005) Plant remains from a Bell Beaker site in Switzerland, and the beginnings of Triticum spelta (spelt) cultivation in Europe. Veget Hist Archaeobot 14:279–286. https://doi.org/10. 1007/s00334-005-0071-1 Andrews AC (1964) The genetic origin of spelt and related wheats. Der Züchter 34:17–22 Armstrong Oma K (2016) Long time – long house. In: Iversen F, Petersson H (eds) The Agrarian Life of the North 2000 BC–AD 1000: studies in rural settlement and farming in Norway. Portal Akademisk, Cappelen Damm, pp 11–25 Bakels C (2014) The first farmers of the Northwest European plain: some remarks on their crops, crop cultivation and impact on the environment. J Archaeol Sci 51:94–97 Bevan A (2019) A stored-products revolution in the 1st millennium BC. Archaeol Int 22:127–144. https://doi.org/10.5334/ai-404 Bilgic H, Hakki EE, Pandey A, Khan MK, Akkaya MS (2016) Ancient DNA from 8400 year-old Çatalhöyük wheat: implications for the origin of neolithic agriculture. PLoS ONE 11:e0151974. https:// doi.org/10.1371/journal.pone.0151974 Buchvaldek M, Lippert A, Košnar L (2007) Archeologický atlas pravěké Evropy (Archaeological Atlas of prehistoric Europe). Univerzita Karlova v Praze, Nakladatelství Karolinum, Praha Chamberlain AT (2006) Demography in archaeology. Cambridge University Press, Cambridge Colledge S (2016) The Cultural Evolution of Neolithic Europe. EUROEVOL Dataset 3: Archaeobotanical Data. J Open Archaeol Data. https://doi.org/10.5334/joad.42 Crema ER (2022) Statistical inference of prehistoric demography from frequency distributions of Radiocarbon dates: a review and a guide for the Perplexed. J Archaeol Method Theory 29:1387– 1418. https://doi.org/10.1007/s10816-022-09559-5 De Capitani A (2002) Gefässkeramik. In: de Capitani A, DeschlerErb S, Leuzinger U, Marti-Grädel E, Schibler J (eds) Die jungsteinzeitliche Siedlung Arbon | Bleiche 3: Funde. Archäologie Im 13 Thurgau 11. Departement für Erziehung und Kultur des Kantons Thurgau, Frauenfeld, pp 135–276 Devroey J-P, van Mol J-J (eds) (1989) L’Épeautre, Histoire Et Ethnologie. Université Libre de Bruxelles, Treignes Dvorak J, Deal KR, Luo M-C, You FM, von Borstel K, Dehghani H (2012) The origin of spelt and free–threshing hexaploid wheat. (Report). J Hered 103:426–441. https://doi.org/10.1093/jhered/ esr152 Feeser I, Dörfler W, Kneisel J, Hinz M, Dreibrodt S (2019) Human impact and population dynamics in the neolithic and bronze Age: multi-proxy evidence from north-western Central Europe. Holocene 29:1596–1606 Frieman CJ (2021) An archaeology of innovation: approaching social and technological change in human society. Manchester University Press, Manchester Grove M (2016) Population density, mobility, and cultural transmission. J Archaeol Sci 74:75–84 Harding AF (2000) European societies in the bronze age. Cambridge University Press, Cambridge Hassan FA (1981) Demographic archaeology. Academic Press, New York Jacomet S (2006) Plant economy of the northern Alpine lake dwellings — 3500–2400 cal BC. Environ Archaeol 11:65–85. https:// doi.org/10.1179/174963106x97061 Jacomet S (2008) Subsistenz Und Landnutzung während Des 3. Jahrtausends v. chr. Aufgrund Von archäobotanischen Daten aus dem südwestlichen mitteleuropa. In: Dörfler W, Müller J (eds) Umwelt – Wirtschaft – Siedlungen Im Dritten Vorchristlichen Jahrtausend Mitteleuropas Und Südskandinaviens. Internationale Tagung Kiel 4–6 November 2005. Wachholtz, Neumünster, pp 355–377. OffaBücher NF 84 Kerig T (2016) Einfache Und Komplexe Wirtschaften: Studien Zur Urgeschichte Des Faktors Arbeit Im mitteleuropäischen Neolithikum. Habilitationsschrift Universität Leipzig, Leipzig Knörzer K-H (1995) Pflanzenfunde aus dem bandkeramischen Brunnen von Kückhoven bei Erkelenz. Vorbericht. In: Kroll H, Pasternak R (eds) Res Archaeobotanicae. International Workgroup for Palaeobotany (IWGP). Proceedings of the 9th symposium Kiel 1992. Oetke-Voge, Kiel, pp 81–86 Knörzer K-H (1998) Botanische Untersuchungen am bandkeramischen Brunnen Von Erkelenz-Kückhoven. Rheinland-Verlag, Köln Kohler-Schneider M, Caneppele A (2009) Late neolithic agriculture in eastern Austria: archaeobotanical results from sites of the Baden and Jevišovice cultures (3600–2800 BC). Veget Hist Archaeobot 18:61–74. https://doi.org/10.1007/s00334-007-0129-3 Kole C (2011) Wild crop relatives: genomic and breeding resources: cereals. Springer, Berlin Köninger J, Kolb M, Schlichtherle H (2001) Elemente von Boleráz und Baden in den Feuchtbodensiedlungen des südwestdeutschen Alpenvorlandes und ihre mögliche Rolle im Transformationsprozess des lokalen Endneolithikums. In: Roman P, Diamandi S (eds) Cernavodă III – Boleráz: ein vorgeschichtliches Phänomen zwischen dem Oberrhein und der unteren Donau. Studia Danubiana. Pars Romaniae. Series Symposia 2. S. C. Vavila, Bucureşti, pp 641–672 Lechterbeck J, Edinborough K, Kerig T, Fyfe R, Roberts N, Shennan S (2014) Is neolithic land use correlated with demography? An evaluation of pollen-derived land cover and radiocarboninferred demographic change from Central Europe. Holocene 24:1297–1307 Lechterbeck J, Kerig T, Kleinmann A, Sillmann M, Wick L, Rösch M (2014) How was Bell Beaker economy related to Corded Ware and early bronze age lifestyles? Archaeological, botanical and palynological evidence from the Hegau, Western Lake Constance region. Environ Archaeol 19:95–113. https:// doi. org/ 10. 1179/ 1749631413y.0000000010 Vegetation History and Archaeobotany Lityńska-Zając M (2016) Early neolithic agriculture in south Poland as reconstructed from archaeobotanical plant remains. The origins and Spread of domestic plants in Southwest Asia and Europe. In: Colledge S, Conolly J (eds) . Routledge, Abingdon, pp 315–326 Maier U (1996) Morphological studies of free-threshing wheat ears from a neolithic site in Southwest Germany, and the history of the naked wheats. Veget Hist Archaeobot 5:39–55. https://doi. org/10.1007/BF00189434 Mills T (2006) A Study of European Cereal Frequency Change during the Iron Age and Roman Periods. PhD thesis, University of Sheffield, Sheffield Müller J (2013) Demographic traces of technological innovation, social change and mobility: from 1 to 8 million europeans (6000–2000 BCE). In: Kadrow S, Włodarczak P (eds) Environment and subsistence – forty years after Janusz Kruk’s settlement studies… Studien Zur Archäologie in Ostmitteleuropa 11. Habelt, Bonn, pp 493–506 Nesbitt M (2001) Wheat evolution: integrating archaeological and biological evidence. In: Caligari PDS, Brandham PE (eds) Wheat taxonomy: the legacy of John Percival. The Linnean Society, London, pp 37–59 Nesbitt M, Samuel D (1996) From staple crop to extinction? The archaeology and history of the hulled wheats. In: Padulosi S, Hammer K, Heller J (eds) Hulled Wheat: Proceedings of the First International Workshop on Hulled Wheats, 21–22 July 1995, Castelvecchio Pascoli, Tuscany, Italy. International Plant Genetic Resources Institute (IPGRI), Rome, pp 41–100 O’Brien MJ, Boulanger MT, Buchanan B, Collard M, Lyman RL, Darwent J (2014) Innovation and cultural transmission in the American paleolithic: phylogenetic analysis of eastern paleoindian projectile-point classes. J Anthropol Archaeol 34:100–119 Peña-Chocarro L, Pérez-Jordà G, Alonso N et al (2019) Roman and medieval crops in the Iberian Peninsula: a first overview of seeds and fruits from archaeological sites. Quat Int 499:49–66. https:// doi.org/10.1016/j.quaint.2017.09.037 Pieler F (2001) Die archäologischen Untersuchungen Der spätneolithischen Befestigungsanlage Von Krems – Hundssteig. In: Wewerka B (ed) Bericht zu den Ausgrabungen Des Vereins ASINOE Im Projektjahr 2001. Fundberichte aus Österreich 40. Bundesdenkmalamt, Wien, pp 503–513 Renfrew C (2009) Demography and Archaeology. Hum Biol 81:381–384 Robinson DE (2003) Neolithic and bronze age agriculture in Southern Scandinavia – recent archaeobotanical evidence from Denmark. Environ Archaeol 8:145–165. https://doi.org/10.1179/env.2003.8. 2.145 Robinson JA, Brown C, Kim BY, Lohmueller KE, Wayne RK (2018) Purging of strongly deleterious mutations explains long-term persistence and absence of Inbreeding depression in Island Foxes. Curr Biol 28:3487–3494 Rogers EM (2003) Diffusion of innovations, 5th edn. Free Press, New York Schlumbaum A, Neuhaus J-M, Jacomet S (1998) Coexistence of tetraploid and Hexaploid Naked Wheat in a Neolithic Lake Dwelling of Central Europe: evidence from morphology and ancient DNA. J Archaeol Sci 25:1111–1118. https://doi.org/10.1006/jasc.1998. 0338 Schumpeter JA (1934) The theory of Economic Development: an Inquiry into profits, Capital, Credit, Interest, and the business cycle. Transaction Publishers, New Jersey Schumpeter JA (1939) Business cycles: a theoretical, historical and statistical analysis of the capitalist process. McGraw-Hill, New York Schumpeter JA (1952) Theorie Der Wirtschaftlichen Entwicklung. Eine Untersuchung über Unternehmergewinn, Kapital, Kredit, zins und den Konjunkturzyklus, 5th edn. Duncker & Humblot, Berlin Seabra L, Tereso J, Bettencourt AMS, Dinis A (2018) Diversidad De cultivos y estructuras de almacenaje en El poblado de crastoeiro (Noroeste De La Península Ibérica): nuevos enfoques. Trab De Prehist 75:361–378. https://doi.org/10.3989/tp.2018.12221 Shennan S (2001) Demography and Cultural Innovation: a model and its implications for the emergence of Modern Human Culture. Camb Archaeol J 11:5–16 Shennan S (2013) Demographic continuities and discontinuities in Neolithic Europe: evidence, methods and implications. J Archaeol Method Theory 20:300–311. https://doi.org/10.1007/ s10816-012-9154-3 Sigaut F (1989) Les spécificités de l’épeautre et l’évolution des techniques. In: Devroey J-P, van Mol J-J (eds) L’Épeautre, Histoire Et Ethnologie. Université Libre de Bruxelles, Treignes, pp 29–49 Stika H-P, Heiss AG (2013) Bronzezeitliche Landwirtschaft in Europa – Der Versuch Einer Gesamtdarstellung Des Forschungsstandes. In: Willroth K-E (ed) Siedlungen Der älteren Bronzezeit: Beiträge Zur Siedlungsarchäologie Und Paläoökologie Des Zweiten Vorchristlichen Jahrtausends in Südskandinavien, Norddeutschland und den Niederlanden. Wachholtz, Neumünster, pp 189–222 Tarde G (1890) Les lois de l’imitation (translated by Elsie Clews Parsons in 1903 and published as The Laws of Imitation). Henry Holt and Company, New York. Taylor MJ, Thoth CA (2011) Cultural Transmission. In: Goldstein S, Naglieri JA (eds) Encyclopedia of child behavior and development. Springer, New York, pp 448–451 Tereso JP, Ramil-Rego P, Álvarez González Y, López González L, Almeida-da-Silva R (2013) Massive storage in as Laias/O Castelo (Ourense, NW Spain) from the late bronze age/iron age transition to the roman period: a palaeoethnobotanical approach. J Archaeol Sci 40:3865–3877. https://doi.org/10.1016/j.jas.2013.05.007 Timpson A, Colledge S, Crema E, Edinborough K, Kerig T, Manning K, Thomas MG, Shennan S (2014) Reconstructing regional population fluctuations in the European neolithic using radiocarbon dates: a new case-study using an improved method. J Archaeol Sci 52:549–557 Walker R, Eriksson A, Ruiz C, Newton TH, Casalegno F (2021) Stabilization of cultural innovations depends on population density: testing an epidemiological model of cultural evolution against a global dataset of rock art sites and climate-based estimates of ancient population densities. PLoS ONE 16:e0247973 Van Zeist W (1976) On macroscopic traces of food plants in southwestern Asia (with some reference to pollen data). Philos Trans R Soc Lond B 275:27–41 Van Zeist W, Buitenhuis H (1983) A palaeobotanical study of neolithic Erbaba, Turkey. Anatolica 10:47–65 Zimmermann A, Hilpert J, Wendt KP (2009) Estimations of population density for selected periods between the neolithic and AD 1800. Hum Biol 81:357–380 Zohary D, Hopf M, Weiss E (2012) Domestication of plants in the old world: the origin and spread of domesticated plants in Southwest Asia, Europe and the Mediterranean Basin, 4th edn. Oxford University Press, Oxford Publisher’s Note Springer Nature remains neutral with regard to jurisdictional claims in published maps and institutional affiliations. 13