Hardwoods of The Pacific Northwest

Research Contribution 8 March 1995
HARDWOODS OF THE PACIFIC NORTHWEST

by
Stanley S. Niemiec
Glenn R. Ahrens
Susan Willits
David E. Hibbs
College of Forest Research Laboratory

Forestry Oregon State University
The Forest Research Laboratory of Oregon State University was established by
the Oregon Legislature to conduct research leading to expanded forest yields,
increased use of forest products, and accelerated economic development of
the State. Its scientists conduct this research in laboratories and forests
administered by the University and cooperating agencies and industries
throughout Oregon. Research results are made available to potential users
through the University’s educational programs and through Laboratory pub-
lications such as this, which are directed as appropriate to forest landowners
and managers, manufacturers and users of forest products, leaders of govern-
ment and industry, the scientific community, and the general public.
The Authors
Stanley S. Niemiec is former Senior Research Assistant, Department of Forest
Products, Oregon State University (OSU), Corvallis, Oregon; Glenn R. Ahrens
is Research Assistant, Department of Forest Science, OSU; Susan Willits is
Research Forest Products Technologist, USDA Forest Service, Pacific Northwest
Research Station, Portland, Oregon; and David E. Hibbs is Associate Professor
and Extension Hardwood Management Specialist, Department of Forest
Science, OSU.
Acknowledgments
Supported by the Red Alder/Value-Added Research and Demonstration Pro-
gram of USDA Forest Service through Pacific Northwest Research Station
Cooperative Agreement PNW 92-0285 with Oregon State University.
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Hardwoods of the Pacific Northwest
by
Stanley S. Niemiec
Glenn R. Ahrens
Susan Willits
David E. Hibbs
Table of Contents
Introduction ............................................................................... 3
Pacific Northwest Hardwoods ................................................... 4
Bigleaf maple (Acer macrophyllum) .......................................... 14
Black cottonwood (Populus trichocarpa) .................................. 24
California black oak (Quercus kelloggii ) ................................... 33
California-laurel (Umbellularia californica ) .............................. 43
Giant chinkapin (Castanopsis chrysophylla) .............................. 51
Oregon ash (Fraxinus latifolia) .................................................. 60
Oregon white oak (Quercus garryana) ...................................... 68
Pacific madrone (Arbutus menziesii) .......................................... 77
Red alder (Alnus rubra) .............................................................. 87
Tanoak (Lithocarpus densiflorus) ............................................. 101
Appendix 1 ............................................................................. 111
Appendix 2 ............................................................................. 114
Appendix 3 ............................................................................. 115
Appendix 4 ............................................................................. 117
Introduction
The history of hardwoods in the Pacific Northwest is sometimes char-

acterized by contradictions. Hardwoods have always been valued as use-
ful trees: they have been a raw material resource for homes, furniture,
and implements; a source of food for people and game animals; and a
source of beauty in a largely evergreen landscape. Hardwoods have also
been weeds, undesirable competitors with regenerative conifers. This
contradiction continues today.
The current economic condition of the forest products industry and
the increased emphasis in forest management on diversity and ecosys-
tem processes continue to stress the importance of Pacific Northwest
hardwoods. More people are seeking a variety of information on hard-
woods, information that is scattered among many, sometimes obscure,
publications or found in the experience of many individuals.
To make this diverse information more readily available, this publi-
cation brings together in one place information on the biology, man-
agement, and processing of the Pacific Northwest (western Washington
and Oregon, and northwestern California) hardwood species. While we
cover a long list of species and a wide range of subjects, we have also
tried to be reasonably comprehensive. We have included ten species:
Common Name Scientific Name
Bigleaf maple Acer macrophyllum Pursh

Black cottonwood Populus trichocarpa Torr. & Gray
California black oak Quercus kelloggii Newb.
California-laurel Umbellularia californica (Hook. & Arn.) Nutt.
Giant chinkapin Castanopsis chrysophylla (Dougl.) A. DC.
Oregon ash Fraxinus latifolia Benth.
Oregon white oak Quercus garryana Dougl. ex Hook.
Pacific madrone Arbutus menziesii Pursh
Red alder Alnus rubra Bong.
Tanoak Lithocarpus densiflorus (Hook. & Arn.) Rehd.
In each chapter, what is and is not known about a given spe-

cies is described according to the same list of topics (see box for
Principal topics covered in principal topics). In many cases, very little is known. Although
each chapter of Hardwoods most of the book is written for people with some background in
of the Pacific Northwest: forestry and wood products, the first chapter reviews basic prin-
ciples and terminology and discusses characteristics common to all
• General characteristics
hardwood species.
• Biology and management The purpose of this document is to increase the knowledge of
• Harvesting and utilization indigenous hardwood species among forest managers and wood
product users, and thus promote and maintain ecosystem diversity
• Wood characteristics and sustainable supplies of these species. We also hope to increase
• Related literature utilization options related to the economic development of hard-
woods.
3
PACIFIC NORTHWEST HARDWOODS
General Characteristics
This chapter provides information that is general to most species

and an explanation of common terminology.
Size, Longevity, and Form

In this book, a hardwood tree is considered mature when it reaches
90 percent or more of its maximum height on a given site. A clear bole
refers to the portion of the tree trunk that has no branches (living or
dead) and is the primary section of the tree used in wood products.
Geographic Range
No species grows well everywhere; each is adapted to a particular
set of circumstances. Most of the western hardwood species of Califor-
nia, Oregon, and Washington have a range that is longer north-south
than east-west. Some have a fairly limited range, but are quite numer-
ous within that range. The proportion of higher-density hardwoods in-
creases from north to south within the region.
Timber Inventory
The most recent available estimates of growing stock inventory for
the major hardwood species were compiled from results of surveys made
by the USDA Forest Service, Forest Inventory and Analysis unit (see Ap-
pendix 1, Table 1). This information is incomplete, however, because
many agencies do not collect or report data on hardwoods, which are
often grouped under the heading “lesser species.”
Biology and Management

Tolerance, Crown Position
Tolerance refers to the ability of the tree to survive and grow under
the cover of the forest canopy. Shading, root competition, and other
factors may be involved in determining tolerance. Species are described
as either intolerant, intermediate, or tolerant; these terms may be modi-
fied by “moderately” or “very.”
4
Crown position or canopy position refers to the relative position of the
crown (foliage) with respect to neighboring trees. This can be described as
suppressed, intermediate, codominant, dominant, or emergent.
Ecological Role
Ecological role refers to the response of the tree to forest distur-
bances (fire, wind, logging, landslides, flooding) and the tree’s require-
ments for such disturbance in order to maintain itself as a viable com-
ponent in the forest. Ecological succession proceeds in the absence of
major disturbances; our terminology refers to the period in the succes-
sional sequence wherein the species is most prominent (pioneer, seral,
or early-successional versus climax or late-successional).
Associated Vegetation
Tree, shrub, and, in some cases, herb or grass species commonly
found with each hardwood species are listed by common name. Appen-
dices 2, 3, and 4 contain lists of common and scientific names of these
plants. Common and scientific names for other organisms are listed
within the text for each hardwood species.
Suitability and Productivity of Sites

For many western hardwoods, there are no established site-index
curves or other guides for estimating the productivity of a site. Gener-
ally, the potential growth of a species on a given site can be evaluated
by examining height growth, diameter growth, and stem form on exist-
ing trees.
The crown position and microsite location of site trees can be repre-
sentative of conditions on the prospective management site. Care must
be taken here, since many hardwoods may be sensitive to slight changes
in topographic position or microclimate.
In the absence of representative trees, site capability can be as-
sessed by evaluating site factors relative to known requirements for the
species in terms of climate, topographic position, soils, elevation, and
disturbance, as discussed in other sections.
Climate
The climate of the Pacific Northwest forests where most of the hard-
woods grow is quite mild. The winters are moist and temperatures
rarely are below 0 ∞C. The summers are dry. Annual precipitation de-
creases and moisture demand increases from north to south and east to
west through this geographic range.
Elevation
The wide range of elevation in the Pacific Northwest creates a wide
gradient of climates. Hardwoods tend to be found at the low and middle
elevations; conifers are found over the whole range below the treeline.
5
Soils
The soils of the region are diverse. Many soils in Washington are de-
rived from glacial debris. Farther south, the soils are generally older, with
parent rock of sedimentary, metamorphic, and volcanic origins. Certain
combinations of soils and topography can produce very droughty condi-
tions.
Flowering and Fruiting

Most hardwoods have inconspicuous, insect-pollinated flowers. Red
alder, however, is wind-pollinated.
Seed
The seeds of hardwoods tend to be large. Many species rely on
animals for seed distribution; most hardwood seeds are an important
source of food for wildlife. Only the pioneer species have small, wind-
blown seed. Seed crops for some species are annual; others produce
large seed crops at intervals of two or more years.
Regeneration from Seed

Trees have regenerated from seed for millennia. Strategies for ac-
complishing this vary by species, but all basically involve the production
of lots of seed over long time periods. Natural regeneration is a slow
process, especially compared to that obtained by planting seedlings.
Many species require a special type of seed bed. Generally, the very
small-seeded species reproduce better on bare mineral soil. Competing
vegetation can limit the success of seedling establishment by reducing
the availability of water, light, or both below levels necessary for hard-
wood survival.
Regeneration from Vegetative Sprouts

Most hardwood species discussed here sprout from the stump after
cutting or fire, providing immediate and vigorous regeneration. Gener-
ally, many more sprouts are produced than can survive over the long
term. Thinning sprouts can increase the growth of residual stems.
Regeneration from Planting

Commercially available supplies of seed, seedlings, or cuttings vary
among species; some species are available, but many are not. Advance
inquiries and arrangements for acquisition of seed and production of
planting stock are often necessary. We know very little about genetic
adaptation for hardwood species. Generally, seed or vegetative propagules
should be collected locally to ensure that trees will be adapted to local
conditions.
The quality of hardwood planting stock can be extremely variable.
In many cases, poor performance of plantings is due to poor quality
stock. Generally, it is not worthwhile to plant stock of marginal quality.
6
Site Preparation and Vegetation Management
As with the culture of many plants, preparation of the growing site
and control of competing vegetation (weeding) will allow hardwood
seedlings to achieve their best potential. In forest settings, site prepara-
tion offers the best opportunity to control competing vegetation; weed-
ing in established hardwood stands is difficult and/or expensive. With
faster growing species, effective management of competing vegetation
is achieved with good site preparation followed by rapid establishment
of hardwood dominance. In many cases, moderate levels of competing
vegetation, debris, or both will require little or no site preparation
beyond the logging operation. Site preparation needs must be carefully
evaluated, however. Hardwoods should not be selected for heavy brush
sites because of brush control costs.
Stand Management
Natural establishment of many hardwood species often occurs via
high initial density and immediate dominance over competing plants.
Management of stand density is the primary avenue for reducing rota-
tion length and improving quantity and quality of commercial yields.
Diameter growth is very responsive to increased growing space. The
strategy in density management is to find the spacing regime that pro-
vides optimal growth while maintaining stem quality. Moderate crowd-
ing is often necessary to maintain stem form, reduce branching and
forking, and induce self-pruning. Once form and self-pruning goals are
met, thinning can maintain or improve diameter growth rate.
Wide initial spacings may be appropriate with intensive culture of
selected genetic stock of high-value hardwoods, or with species that
produce both fruit or nut and timber crops.
Mixed-species Stands
Mixed-species stands, particularly conifer/hardwood mixes, are ap-
pealing for many reasons. Hardwoods can maintain or improve desir-
able soil characteristics via input of nutrients and organic matter. Mixed
stands provide forest products (both special forest products and wood
products) and diversity for wildlife; they mitigate fire damage and are
aesthetically attractive. The challenge with management of mixed stands
is the integration of differing growth patterns, sensitivities to competi-
tion, and rotation ages. The simplest approach may be to manage mixed
species in monospecific patches.
Fast-growing species and stump sprouts may suppress establishing
seedlings of slow-growing species. When conifers reach maximum growth
potential later in a rotation, it can become difficult to maintain codominance
and growth of intermixed hardwoods. Strategies for management in-
clude delaying establishment or maintaining low proportions of fast-
growing species among slow growers. Later, the conifers must be thinned
to provide growing space for hardwoods.
Growth and Yield

Of the Pacific Northwest hardwoods, the growth and yield of red alder
is, by far, the best. For red alder, almost all of the available information
7
comes from natural, unmanaged stands. Although there have been few
research studies and thinning demonstrations of red alder and several other
hardwoods, we are forced to rely heavily on this limited data base.
An important characteristic of hardwood growth is that hardwood
stands do not and cannot achieve the stocking levels common for coni-
fers. For example, a basal area of only 150 ft 2 per acre is considered
high for red alder. This stocking level, however, can be achieved in a
relatively short time, compared to conifers. Thus, management strate-
gies of hardwoods tend to be based on either short rotations or high-
value products.
Interactions with Wildlife

The interactions of hardwood trees with wildlife vary with species.
Many hardwoods provide habitat characteristics that conifers do not;
some wildlife species are dependent on hardwoods. Inclusions of hard-
woods, therefore, in a landscape or forest stand increases wildlife diver-
sity and abundance.
Insects and Diseases

All plants have their own particular insect and disease problems. In
hardwoods, these problems generally do not become epidemics.
Genetics
Little is known about the levels of genetic variability of the region’s
hardwoods. Red alder has received the most attention; one study of
Oregon white oak suggested that there is limited variability among the
white oak populations in Washington.
Because little is known of the levels of genetic variability, a conser-
vative, careful approach is required in retaining and selecting seed sources
for regeneration. Seed should be collected from stands of similar cli-
mate and soils to the regeneration site. We recommend simply staying
within a locally defined conifer seed zone and limiting seed movement
to within a 500-ft elevation band. Within the seed-collecting area, select
stands and trees within stands of better growth and form.
Harvesting and Utilization

Cruising and Harvesting
Two types of cruises are currently being used to estimate tree vol-
umes. Both use standard sampling design and cruising techniques, in-
cluding the measurement of DBH, form class, merchantable height, and
top diameter, but they differ in volume estimation. One method uses
standard 16-ft log lengths, and deducts for sweep, crook, rot, and other
defects. It will generate consistent volume estimates without regard to
manufacturing lengths and processing technology. The other method
8
uses preferred milling lengths (usually 8, 10, or 12 ft) and attempts to
fit segments between defects. Although this method may result in esti-
mates closer to final utilization by a particular mill, the results will be
highly variable and will depend on cruisers’ judgement and harvesting
and processing technology. We strongly recommend that volume esti-
mates be based on cubic feet rather than board feet in order to elimi-
nate biases within the Scribner system. Tools such as stand-volume
tables or tree-volume equations are lacking or poorly developed for
many hardwoods and may not yield accurate results for specific stands.
Estimates of tree quality and value are typically based on estimates
of log grades and sizes. Although log grades have been developed and
tested for most hardwood species, they are not often used by log buy-
ers in the Pacific Coast region. The use of standard log grades would
assist in the development of the hardwood industry, however. Purchas-
ers often determine prices and specifications for log diameters and lengths
for individual sales based on their knowledge of local timber and on the
current needs of the mill.
In the past, there have been problems in adapting softwood technol-
ogy to the harvesting of hardwood timber. The current emphasis on small-
log harvesting is providing new technology and better trained loggers for
the hardwood industry. Typically, hardwood manufacturers process shorter
logs (8, 10, and 12 ft) than do softwood manufacturers; this allows much
more flexibility in the log bucking process. It is important to consider log
grade as well as volume during bucking. Bucking for grade makes a critical
difference in the net value of the trees; bucking to variable lengths to
maximize the grade and reduce the impact of crook and sweep is recom-
mended. Proper felling and bucking techniques are essential for making a
profit in the hardwood industry. Harvesting and transportation costs for
hardwoods may be slightly higher than for softwoods because there are
generally fewer stems per acre to be harvested, species have a higher
density (heavier logs), logs tend to be more crooked, and, because of tree
form, there may be more short logs.
As mentioned previously, traditional softwood log scaling rules do
not necessarily provide accurate estimates of usable volumes because
the hardwood manufacturers can use shorter log lengths and different
lumber sizes to produce end products. In some cases, logs are sold on a
weight basis to avoid inaccuracies in the log measurement systems.
Product Recovery
Logs are generally bucked at the sawmill into preferred milling lengths,
which circumvents some problems with misshapen logs. Sawlogs have a
minimum diameter of 6 to 10 in., depending on the species and the
mill. Hardwood mills are typically smaller than softwood mills and manu-
facture lumber at lower production rates. Mills normally produce ran-
dom-width lumber in 4/4, 5/4, 6/4, and 8/4 thicknesses, with 4/4 be-
ing predominate.
The standard grades are based on the percentage of surface mea-
sure with clear cuttings on the poorest side of the board. These clear
cuttings are located between knots and other types of defects. Select
grade is the exception because it is based on the best side of the board.
Lumber grades are designed to be used on green lumber; in practice
9
most of the lumber is dried and surfaced before it is graded. At the
sawmill, care should be taken so that over-edging does not significantly
reduce grade and recovery volume.
Average rates of lumber grade recovery from northwestern hard-
wood logs are generally similar to rates for related eastern hardwood
species within comparable log grades (Appendix 1, Table 2).
Wood Properties
Characteristics
These properties describe the texture, color (for both heartwood and
sapwood), odor or taste, and the cellular structure of the wood. In the
latter group, consideration is given to the pore patterns, description of the
annual growth increments, and prominence of various cell types.
The pore structure of end-grain surfaces is identified as either dif-
fuse porous, ring porous, or a combination of both, which then is termed
either semi-diffuse or semi-ring porous. Diffuse porous woods have little
or no variation in pore size within an annual growth increment. Ring
porous woods have a few large pores at the start of each growth incre-
ment that change to small, more numerous pores for the remainder of
the annual growth. If the transition from large to small pores is gradual
or if the initial pores are not extremely large, then the wood is de-
scribed as semi-ring or semi-diffuse porous. The visibility of the start of
each growth increment also helps to identify woods. For many diffuse
porous woods, that beginning is difficult to discern without magnifica-
tion. For ring porous woods, the start of each growth increment is very
prominent. Diffuse porous woods are red alder, bigleaf maple, and Pa-
cific madrone; ring porous woods are the oaks, tanoak, and ash. Ring
porous woods are also more coarse and grainy when viewed on tangen-
tial (flat-sawn) surfaces. The size and volume of the wood rays—cells
that appear as ribbon-like strands extending across the grain in the
radial direction, which transport fluids and nutrients laterally—also help
to identify woods. Rays can be very fine and visible only with magnifica-
tion, or very large and prominent. When radial surfaces are exposed
during sawing, their structure can enhance the surface appearance and
create unique patterns.
Weight
Specific gravity or density is an excellent descriptor of wood and is
correlated to many important wood attributes such as mechanical strength,
shrinkage, and cutting forces associated with machining. Specific gravity
often receives first attention when the potential of a species is assessed
(Appendix 1, Table 3).
Basic specific gravity is the weight divided by the volume; it directly
relates the wood weight to the weight of water (considering equal vol-
umes). Water has a specific gravity of 1.00, so a wood with a specific
10
gravity of 0.50 weighs half as much as water. The specific gravity is
typically given for wood that is green (freshly cut), ovendry (0 percent
moisture content), and at 12 percent moisture content, which approxi-
mates many in-service conditions. For wood, specific gravity is calcu-
lated on the ovendry weight and the green-wood volume.
It is important to recognize that wood is a natural material that results
from considerably diverse origins, life histories, and growth conditions. As
such, the values given for a specific wood represent average values and do
not indicate the variability or the range of values possible.
Mechanical Properties
It must be emphasized that the mechanical properties we present
have been summarized from several sources (Appendix 1, Table 3). Sampling
and testing procedures may have varied; therefore, the values should be
considered only for comparison purposes between species and for de-
termination of possible appropriate end uses. Test results reported here
are based on the ASTM procedures of D 143. As with specific gravity
information, the values given are averages and do not indicate the pos-
sible range and variability of the material.
Drying and Shrinkage

We have noted the response of individual woods to air- and kiln-
drying, and the types of degrade likely to occur. Shrinkage data from
green to ovendry is given to provide information about changes in size
and in-service stability.
Kiln schedules are provided for each species, as kiln efficiencies vary
considerably; however, these schedules should be considered as conserva-
tive starting points. These schedules are also based on specific stacking and
stickering procedures. Any changes in kiln design or function, stacking,
stickering, or pre-drying/air-drying should be noted. These changes may
require schedule alteration to prevent excessive degrade of material.
Some pre-drying treatments and post-drying reconditioning may help
minimize degrade (as with Pacific madrone) but the exact procedures
are for the most part either proprietary or experimental.
All moisture content (MC) data are based on an ovendry basis using
the following formula:
MC percent = [(wet weight - dry weight)/dry weight] ¥ 100
Machining
Much of the information on the machining, adhesive bonding, and
finishing of hardwoods is subjective. Judgement, rather than test data
has determined some of the evaluations, and the use of different sources
has further complicated the situation.
We have described the machinability of wood, such as its tendency
to chip and tear during planing and shaping; the smoothness of the
final machined surface; and the dulling of tooling. We have included
recommendations for tooling geometry as well.
11
Adhesives
Glue-joint quality for side grain joints is given. Quality and in-ser-
vice performance of glue joints depend on the density of the wood
(lower density woods generally bond better than higher density woods),
the quality and newness of the machined surfaces, the type and quality
of the adhesive, and, for long-term performance, the effectiveness of
the finish in excluding moisture and protecting the wood.
Finishing
In this section, we have considered the application and appearance
of clear interior coating and associated coloring systems used for furni-
ture, cabinets, or flooring. Finish quality depends on surface prepara-
tion and quality, the grain/pore structure of the wood, and the unifor-
mity of color between heartwood and sapwood.
Durability
Information in this section describes the natural resistance of the
heartwood to decay fungi during ground-contact exposure. If the wood
is particularly susceptible to insect attack, we have noted the specific
types of insects and damage. We have also indicated whether a wood is
specifically susceptible to staining during handling, storage, or expo-
sure to certain metals.
Uses
The uses of hardwood lumber are many and varied. Higher quality
material is utilized in products such as veneers for paneling and hard-
wood plywood, cabinets (both as solid wood and from hardwood ply-
wood), furniture, doors, and moulding and millwork. Intermediate grades
tend to be used for furniture, solid paneling, and flooring. Lower grades
serve utilitarian applications such as pallets, boxes and crating, and
small cut-stock applications that avoid the defects. Very low-grade ma-
terial and residues can be used for particle-board-type composites, pulp
for paper and container board, or fuel or agricultural mulches. Care
should be taken so that the higher value products are considered before
these resources are processed for high-volume, low-value end uses, as is
often the case.
The development of products from indigenous hardwoods species
has not been fully explored. As a result, many opportunities exist for
creating employment and for supporting economic diversification with
entrepreneurial innovation, increased entrepreneurial, corporate, and
governmental sector research and development, and market opportu-
nity analysis.
Related Literature
BOLSINGER, C.L. 1987. Major findings of a statewide resource assess-

ment in California. P. 291–297 in Proceedings of the Symposium on
12
Multiple-Use Management of California’s Hardwood Resources. T.R.
Plumb and N.H. Pillsbury, tech. coords. USDA Forest Service, Pacific
Southwest Forest and Range Experiment Station, Berkeley, Califor-
nia. General Technical Report PSW-100.
BOONE, R.S. 1988. Dry Kiln Schedules for Commercial Woods—Temper-
ate and Tropical. USDA Forest Service, Forest Products Laboratory,
Madison, Wisconsin. General Technical Report FPL-GTR-57. 158 p.
GRATKOWSKI, H. 1961. Brush problems in southwest Oregon. USDA
Forest Service, Pacific Northwest Forest and Range Experiment Sta-
tion, Portland, Oregon. 53 p.
HARLOW, W.M., E.S. HARRAR, and F.M. WHITE. 1979. Textbook of Den-
drology. McGraw-Hill, New York. 510 p.
LITTLE, E.L., Jr. 1979. Checklist of United States Trees (Native and Natu-
ralized). USDA Forest Service, Washington D.C. Agriculture Hand-
book 541. 375 p.
McDONALD, P.M. 1983. Local volume tables for Pacific madrone, tanoak,
and California black oak in north-central California. USDA Forest
Service, Pacific Southwest Forest and Range Experiment Station, Ber-
keley, California. Research Note PSW-362. 6 p.
McDONALD, P.M., D. MINORE, and T. ATZET. 1983. Southwestern Or-
egon-northern California hardwoods. P. 29-32 in Silvicultural Sys-
tems for the Major Forest Types of the United States. R.M. Burns,
tech. compil. USDA Forest Service, Washington, D.C. Agriculture
Handbook 445.
NATIONAL HARDWOOD LUMBER ASSOCIATION. 1982. Rules for the mea-
surement and inspection of hardwood and cypress. Memphis, Ten-
nessee.
OVERHOLSER, J.L. 1977. Oregon Hardwood Timber. Forest Research Labo-
ratory, Oregon State University, Corvallis. Research Bulletin 16. 42 p.
PANSHIN, A.J., and C. de ZEEUW. 1980. Textbook of Wood Technology.
4th edition. McGraw-Hill Book Co., New York. 722 p.
PILLSBURY, N.H., and M.L. KIRKLEY. 1984. Equations for total, wood,
and saw-log volume for thirteen California hardwoods. USDA Forest
Service, Pacific Northwest Forest and Range Experiment Station, Portland,
Oregon. Research Note PNW-414. 52 p.
RAETTIG, T., G.R. AHRENS, and K. CONNAUGHTON. 199?. Hardwood
supply in the Pacific Northwest: a policy perspective. USDA Forest
Service, Pacific Northwest Research Station, Portland, Oregon. In
preparation.
SNELL, J.A.K., and S.N. LITTLE. 1983. Predicting crown weight and bole
volume of five western hardwoods. USDA Forest Service, Pacific Northwest
Forest and Range Experiment Station, Portland, Oregon. General Technical
Report PNW-151. 37 p.
STROEMPL, G. 1983. Thinning clumps of northern hardwood stump
sprouts to produce high quality timber. Ontario Ministry of Natural
Resources, Toronto, Ontario. Forest Resource Information Paper 104.
27 p.
USDA FOREST SERVICE. 1987. Wood Handbook: Wood as an Engineer-
ing Material. Forest Products Laboratory, Madison, Wisconsin. Agri-
culture Handbook 72. 466 p.
13
BIGLEAF MAPLE
ACER MACROPHYLLUM
OREGON MAPLE
LARGE-LEAF MAPLE
BROADLEAF MAPLE
Bigleaf maple is second to red alder among native hardwood species

in abundance and in commercial importance in the Pacific North-
west. It is the only western maple that reaches commercial size, yet
Bigleaf Maple its potential as a commercial species has not been fully recognized.
YUKON
Range

BRITISH
Mature bigleaf maple trees range from 50 to 100 ft in height (101 ft
COLUMBIA maximum) and 12 to 36 in. in DBH (133 in. maximum). Bigleaf maple
is moderately long-lived; some individuals may reach 300 years of age.
Height growth becomes negligible after 50 to 70 years. In forest stands,
maples often develop clear (50 to 70 percent of total height), well-
formed stems with narrow crowns. Open-grown trees have broad, rounded
crowns on short, branching boles. The root system of bigleaf maple is
shallow and spreading on wet or shallow soils.
WA Geographic Range
Bigleaf maple is native to the Pacific Northwest at low to middle
OR elevations from San Diego to Vancouver Island (lat 33 to 51∞N). It
ID usually grows from near the Pacific Ocean to a maximum of 186 miles
CA inland.
Timber Inventory
Substantial volumes of bigleaf maple occur in every subregion of
the Northwest (Appendix 1, Table 1). Maple is most abundant in the
Puget Sound region of Washington and the Northwest region of Or-
14
egon. About 18 percent of the total hardwood volume in the Pacific
Northwest is bigleaf maple.

Bigleaf maple occurs as a dominant, codominant, or intermediate
tree in even- or uneven-aged stands. It is quite tolerant and commonly
establishes in forest understories, where it can persist for long periods.
Maple in the understory can quickly respond to release when openings
are created in the overstory. It also has the capacity to grow rapidly and
maintain dominance under favorable conditions in the open sun.
Ecological Role
Bigleaf maple increases in abundance during intermediate to late
stages of succession. It can follow willow and alder in riparian succes-
sion. On drier upland sites, bigleaf maple can replace oaks and Pacific
madrone in the absence of significant disturbance.
The abundance of bigleaf maple also increases after disturbance in
stands with an established maple component. Basal sprouts stimulated
by cutting or burning can dominate other vegetation in the new stand.
Removal of overstory trees can stimulate rapid growth on bigleaf maple
in the understory.
Bigleaf maple is commonly found in association with almost all other
tree species in its range. The greatest abundance of bigleaf maple is in
the foothills of the Cascade and Coast ranges. Its most common associ-
ates are Douglas-fir, grand fir, red alder, Sitka spruce, western redcedar,
western hemlock, Pacific dogwood, and Pacific madrone. Common shrub
and herb associates include serviceberry, salal, red huckleberry, devil’s-
club, Pacific rhododendron, thimbleberry, salmonberry, vine maple, and
hazel. Herbaceous associates include maidenhair fern, western swordfern,
ladyfern, red woodsorrel, false lily-of-the-valley, prince’s-pine, spread-
ing sweetroot, and twinflower.
Bigleaf maple also supports abundant epiphytic growth on branches
and boles in moist climates. Common epiphytes are licorice fern, club
moss (Selaginella oregana), and other mosses (Hylocomium splendens,
Leucolepis menziesii, Isothecium stoloniferum, and Neckera menziesii) and
lichens (Cladonia, Nephroma, and Crocynia spp.).
15
Bigleaf maple grows best on river terraces, flood plains, and seepage
areas. Good performance is also common on upland sites if soils are
deep and moisture is abundant. Bigleaf maple will establish and grow
on a wide variety of sites, including relatively harsh, dry areas in south-
western Oregon; however, growth and stem form may be poor. The
suitability of prospective sites should be carefully evaluated before man-
agement of bigleaf maple is attempted.
There are no established guides or site-index curves for estimating
the productivity of a site for bigleaf maple. Good potential for growth
of bigleaf maple on a site is indicated by the following:
• Top height of existing mature trees is 80 to 100 ft

• Rapid juvenile height growth of >3 ft per year
• Sustained height growth from age 15 to 30 of 1 to 2 ft per year
• Continuing diameter growth on mature trees.
Climate
Bigleaf maple is commonly found across a wide range of climates,
from the cool, moist (temperate rain forest) conditions of the Olympic
Peninsula to the warm, dry (Mediterranean) conditions of southwestern
Oregon. Within its range, precipitation varies from 22 to 260 in. (560 to
6600 mm) annually, and from 2 to 46 in. (50 to 1170 mm) during the
growing season. Average temperatures range from a minimum of 28 ∞F
(January) to a maximum of 81∞F (July). Where maple is most prolific, the
climate is mild and humid, with moderate precipitation during the growing
season.
Bigleaf maple can grow on relatively hot, dry areas, such as upland
sites in southwestern Oregon. Maple tolerates moisture stress as low
as -20 bars (-2 MPa, nocturnal), which is a low value for broadleaved
deciduous trees. Planted maple seedlings are susceptible to heat-gir-
dling and sunscald on the lower stem, however.
We have little information concerning susceptibility of bigleaf maple
to cold, ice, and snow. A low tolerance to cold and snow is indicated by
the absence of maple at higher elevations and by its narrow coastal
range. Cold temperatures probably limit the northern occurrence of
bigleaf maple.
Elevation
Bigleaf maple usually grows at low to middle elevations (near sea
level to 3000 ft) in the central part of its range. In southern California,
it may be found from 3000 to 7000 ft; at the northern end of its range
it seldom occurs above 1000 ft in elevation.
Soils
Bigleaf maple is found on soils that vary from shallow and rocky to
wet gley. Although it may tolerate drought or poor soil conditions, its
16
growth will be poor at those extremes. It grows best on deep, well-
drained soils with abundant moisture, conditions that occur most com-
monly on river terraces, flood plains, and seepage areas.
Bigleaf maple is not as tolerant of poor drainage or flooding as other
riparian species such as red alder, cottonwood, or ash. Bigleaf maple
appears to be somewhat tolerant of wet conditions, as indicated by its
shallow, spreading root system and its common presence in wet areas.
Flooding for more than 2 months during the growing season will kill
maples of all ages.
Bigleaf maple does not seem to require high soil fertility, based on
its competitive success over a range of soil nutrient conditions. One
study indicates a high sensitivity to toxic boron in the soil. Maple is
considered to be a soil-building species; leaves and litter contain rela-
tively high concentrations of macro- and micro-nutrients.

Bigleaf maple begins to produce seed at about 10 years of age.
Maple is polygamous, bearing both male flowers and perfect flowers in
one cylindrical raceme. The flowers appear before the leaves in early
spring. The greenish-yellow flowers are pollinated by insects within 2 to
4 weeks after bud-burst.
Seed
Bigleaf maple seeds are borne in pubescent, double samaras with
wings from 1.4 to 2 in. long. Seeds are triangular or oval in shape and
0.16 to 0.47 in. long. There are from 2700 to 4000 seeds/lb. Seeds
ripen early in September and October, and are dispersed by the wind
from October through January. Many seeds may remain on trees during
this period.
Seed should be collected from healthy, well-formed trees. If the
outplanting site is known, parent trees should be selected from a nearby
locale that has conditions similar to the outplanting site. Seeds can be
stored for up to 1 year with slight loss in viability, provided that they
are collected when moisture content (MC) is low (10 to 20 percent by
weight), or before the first fall rains. If seed are collected later, at higher
MC, they should not be dried. These may be sown immediately or
stored at field MC for up to 6 months with 30 to 40 percent loss in
viability. Seeds should be stored in airtight containers at 34 ∞F.
Bigleaf maple seeds are typically sown in the fall, soon after collec-
tion. Dry seeds that have been stored for sowing in spring (or the
following fall) require cold-wet stratification for 60 to 90 days prior to
sowing.
Natural Regeneration from Seed

Most bigleaf trees produce seed every year, although the amount
may vary greatly from year to year. All viable seeds will germinate in the
first year; delayed germination does not occur. Seeds germinate well on
17
both mineral and organic substrates. Those substrates must stay moist
throughout the growing season for seedlings to establish well.
Germination and establishment rates are best under partial shade.
Natural rates of establishment are low under dense shade or in clearcuts.
Good conditions for natural regeneration occur when overstory conifers
are thinned out by natural self-thinning or silvicultural thinning. Dense
brush and understory vegetation inhibit natural regeneration.

Maple has a prodigious capability to sprout from cut stems of any
size, from seedlings to large trees. Sprouts provide a reliable means of
regeneration from existing trees. Unmanaged sprout clumps produce
too many stems and poor stem quality. Management may greatly im-
prove the quality of stems from sprouts of bigleaf maple.

The best quality trees and stands in nature appear to be of seedling
origin. There have been very few efforts to plant bigleaf maple in the
wild. Although some forest nurseries are producing bigleaf maple on a
trial basis, commercial supplies of maple seedlings are not consistently
available. To ensure a supply of maple seedlings, arrangements must
first be made for collection of seed and for production of seedlings at a
qualified nursery. Very large (3 to 6 ft tall) nursery seedlings may be
produced in one year.
Observations made on field plantings to date indicate that plantings
are very susceptible to deer browse. Planted seedlings are sometimes
prone to forking and poor form, particularly after they have been browsed
or physically damaged. To maintain stem quality in plantations, seed-
lings may need to be protected from deer, planted in high densities,
and correctively pruned.

There are no specific studies of site preparation and vegetation man-
agement practices for bigleaf maple. The outstanding performance of
bigleaf maple cultivated in nurseries and back yards indicates great po-
tential for management of seedlings in the field. Control of competing
vegetation should improve on the low rates of establishment and slow
growth observed for maple in the presence of dense understory vegeta-
tion. Little or no site preparation is required with maple regenerating
from stump sprouts.
Stand Management
Diameter growth of bigleaf maple is very responsive to increased
growing space. Management should target a spacing regime that pro-
duces optimal growth while maintaining the benefits of crowding in
young stands or clumps. Moderate crowding is necessary to reduce
18
branching and forking and induce self-pruning. A dense canopy of bigleaf
maple will also suppress competing understory vegetation, reducing the
need for other vegetation management treatments.
Control of spacing via initial planting density or precommercial thin-
ning (PCT) is recommended to allocate rapid growth to crop trees at an
early age (5 to 15 years). To maintain moderate crowding, stands should
not be opened too much initially. Intermediate thinning (for pulpwood,
firewood, PCT) may be needed to maintain diameter growth. Bigleaf
maple can continue to grow in diameter and respond to released grow-
ing space for at least 30 to 40 years.
The shade-tolerant bigleaf maple can be grown in any crown posi-
tion in stands with mixed species or age classes. Management of mixed
stands is complex; stands may require periodic treatments to maintain
the growth of diverse components. Bigleaf maple stump sprouts must
be controlled or thinned to prevent the early suppression of associated
seedlings. Later treatments may be needed to maintain growth of bigleaf
maple when intermixed conifers ultimately reach their superior height.
Delayed establishment of bigleaf maple seedlings in conifer planta-
tions is a sensible strategy, since maple naturally establishes and grows
under partial cover. Thinning and vegetation management may be needed
to maintain adequate space for the growth of bigleaf maple. With even-
aged mixtures or short (<10 years) delays in bigleaf maple establish-
ment, maple sawtimber can be harvested at the same time as associated
conifers.
Because this species tends to establish in patches or clumps, it may
be sensible to manage bigleaf maple in monospecific patches in mixture
with other trees. This management in patches approach may be applied
to groups of dominant maple sprout clumps or to patches of seedlings
establishing in openings.
Growth and Yield

Juvenile growth rates of dominant maple stands or patches are quite
rapid, matching or exceeding those of red alder. Yields of about 4500
ft 3 per acre were estimated for 70-year-old stands of pure bigleaf maple
in British Columbia. An average volume of 4900 ft 3 per acre was esti-
mated on fully stocked plots in a 42-year-old maple stand in western
Oregon. Gross annual volume growth on these plots was about 140 ft 3
per acre. On these plots, height growth after 45 years already appeared
to be very slow (no measurable growth in 3 years). A rotation age of 40
to 50 years may be feasible in managed stands.

Damage caused by deer and elk is probably the most important
factor affecting the height and form of bigleaf maple seedlings and
sprouts. The foliage and young stems of bigleaf maple are preferred
browse for elk and deer, and these animals often use saplings for rub-
19
bing antlers. Seedlings and saplings are often clipped by mountain bea-
ver. Birds and rodents feed on bigleaf maple seeds. Predation by rodents
and invertebrates is a major cause of seedling mortality. Bigleaf maple
provides an important broadleaved, deciduous component in the conif-
erous forests of the Northwest. The leaves are rich in bases and provide
for a diversity of insects and other microfauna. The understory vegeta-
tion associated with maple can be quite different from that in adjacent
conifer forests. A variety of birds and mammals may benefit from these
distinct attributes of food and habitat.

Young, undamaged trees are generally free of serious disease or de-
cay. Old or damaged trees commonly have serious defects caused by
wood-rotting fungi, which invade through stem and branch wounds.
Root pathogens (Armillaria spp.) and butt rots also attack older trees.
The verticillium wilt (Verticillium albo-atrum) can be a serious problem
for ornamental trees and it sometimes kills bigleaf maple in the forest.
Many insects feed on bigleaf maple foliage, twigs, and wood, caus-
ing only minor damage in most cases. The carpenter worm (Prionoxystus
robinae) can cause serious damage in living trees of all sizes. Roundheaded
borers often damage the wood of dead and dying trees. Powderpost
beetles (Ptilinus basalis) may rapidly infest dead trees or lumber that is
improperly stored.
Genetics
In genetics research, the major interest is in cultivars for ornamental
uses. A red-leaved variety of bigleaf maple (Acer macrophyllum Pursh
forma rubrum) has been found in northern California. Another variety
with triple samaras (Acer macrophyllum Pursh var. kimballi Harrar) is oc-
casionally found in Washington.

Total tree volume in cubic feet and sawlog volume can be estimated
from DBH and total height with tables or equations. Log grades devel-
oped by the Forest Service for eastern hardwoods and modified by Or-
egon State University appear to separate log values to the point that it
may be worth grading logs for marketing. Generally, logs are priced on
the basis of diameter and length specifications developed by local log
buyers.
Product Recovery
Sawlogs generally have a minimum diameter of 7 to 10 in. Lumber is
graded with the special NHLA rules for bigleaf maple; grades include
20
Selects and Better, No. 1 Shop, No. 2 Shop, No. 3 Shop, and Frame.
Unlike the standard NHLA grading rules, these grades are generally based
on the best face of the piece. Grades can be applied to rough, surfaced,
green, or dry lumber; in practice, lumber is usually dried and surfaced
before grading. One mill study conducted with NHLA standard grades
rather than the modified red alder and maple grades found a percentage
recovery of No. 1 Common or Better green lumber from bigleaf maple
logs (80 percent and 58 percent for grade 1 and 2 logs, respectively)
that is quite good compared to other hardwoods (Appendix 1, Table 2).
There are also specialty markets for figured wood from bigleaf maple,
although there are no standard grades. Craftsmen, both local and dis-
tant, may pay premium prices for burls and wavy, quilted, fiddle-back,
or bird’s-eye grain patterns. This figured material is utilized for thin-
sliced, decorative veneers for furniture and architectural paneling.
Wood Properties
Characteristics
The wood from bigleaf maple is fine-grained and of moderate weight
and hardness. The sapwood is reddish-white, sometimes with a grayish
cast; the heartwood is light pinkish-brown. The wood is without any
characteristic odor or taste. The growth rings on the end grain are not
very distinct; on the radial and tangential surfaces, however, the growth
rings are generally delineated by a narrow, dark brown line, which shows
the figure of the wood. The pores are evenly distributed in the growth
rings (diffuse porous) and are moderately small to medium in size. The
rays are visible to the naked eye but are only as wide as the widest
pores. Although much of the wood is straight-grained, some highly fig-
ured wood that includes wavy, quilted, fiddle-back, or bird’s-eye grain
patterns is also produced.
Weight
Bigleaf maple weighs about 47 lb/ft 3 when green, and 34 lb/ft 3 when
dried to 12 percent MC. The average specific gravity is 0.44 (green) or
0.51 (ovendry).
The strength properties of bigleaf maple are exceptionally good, con-
sidering its intermediate specific gravity. While it is not as strong as the
eastern hard maples, it performs better in most tests than the soft maples.
Bigleaf maple is suitable for most furniture design applications, and the
lower grades perform well as pallet stock. It holds nails well and is not
likely to split with nailing. See Appendix 1, Table 3, for average me-
chanical properties for small clear specimens.
21
Under properly controlled conditions, bigleaf maple can be success-
fully kiln-dried in a short time with a minimum of degrade. The most
prevalent drying defects are end checks and collapse, or mold growth
that causes stained wood. The average green MC is 72 percent (ovendry
basis).
The radial shrinkage (green to ovendry) is 3.7 percent and the tan-
gential shrinkage value is 7.1 percent. These values are the same as
those of black cherry, and are better than those of red alder (radial 4.2
percent, tangential 7.3 percent) and Oregon white oak (radial 4.4 per-
cent, tangential 9.0 percent). See Table 1 for an appropriate kiln sched-
ule for 4/4, 5/4, and 6/4 lumber.
Table 1. Kiln schedule—Bigleaf maple 4/4, 5/4, 6/4.

Equilibrium
Moisture Temperature ∞F moisture Relative Temperature ∞C
content Dry- Wet- content humidity Dry- Wet-
Step (%) bulb bulb (%) (%) bulb bulb
1 Above 50 130 123 14.3 81 54.5 50.5

2 50 to 40 130 120 12.2 74 54.5 49.0
3 40 to 35 130 115 9.6 62 54.5 46.0
4 35 to 30 130 105 6.7 43 54.5 40.5
5 30 to 25 140 100 4.2 25 60.0 37.5
6 25 to 20 150 100 3.2 18 65.5 37.5
7 20 to 15 160 110 3.4 21 71.0 43.5
8 15 to final 180 130 3.5 26 82.0 54.5
Equalize and condition as necessary.
Machining
Bigleaf maple wood retains many of the favorable machining (plan-
ing, shaping, boring, and turning) characteristics of the eastern hard
maples, while allowing for greater production feed rates because of its
lower density. Best results for planing were obtained with hook angles
of 20∞. As with other fine-grain, hard woods, surface scratching associ-
ated with sanding (swirls and cross-grain) can be a problem, although
not to the same degree as with the hard maples.
Adhesives
There appear to be no reliable test results available that deal directly
with the newer synthetic adhesives, but communication with local users
indicates that bigleaf maple performs satisfactorily with good quality
joints if conditions are well controlled. The glue line can be visible when
darker resins are used because of the wood’s light color.
22
Finishing
Bigleaf maple finishes well and there is no need to fill the grain. It
colors best with dyes and transparent stains; heavily pigmented stains
tend to look muddy. With careful color selection, the straight-grained
wood can be stained to resemble cherry. Figured bigleaf maple is ex-
ceptionally beautiful when clear-coated.
Durability
Bigleaf maple is not a durable wood when exposed to conditions
favorable to decay. When properly seasoned, the wood is relatively stable
and is not apt to split or surface check in use.
Uses
Bigleaf maple is used for furniture, veneer, paneling, hardwood ply-
wood, musical instruments, moulding, pallets, turnery, pulpwood, and
firewood.
Related Literature
BASTENDORFF, K.M., and A. POLENSEK. 1984. Strength and stiffness of

red alder and bigleaf maple pallet materials. Forest Products Journal
34(7/8):51-56.
FRIED, J.S., J.R. BOYLE, J.C. TAPPEINER II, and K. CROMACK Jr. 1990.
Effects of bigleaf maple on soils in Douglas-fir forests. Canadian
Journal of Forest Research 20:259-266.
FRIED, J.S., J.C. TAPPEINER II, and D.E. HIBBS. 1988. Bigleaf maple seed-
ling establishment and early growth in Douglas-fir forests. Canadian
Journal of Forest Research 18:1226-1233.
MINORE, D., and J.C. ZASADA. 1990. Bigleaf maple. P. 33-40 in Silvics
of North America, Volume 2, Hardwoods. R.M. Burns and B.H. Honkala,
coords. USDA Forest Service, Washington D.C. Agriculture Hand-
book 654.
NADKARNI, N.M. 1984. Biomass and mineral capital of epiphytes in an
Acer macrophyllum community of a temperate moist coniferous for-
est, Olympic Peninsula, Washington State. Canadian Journal of Botany
62:2223-2228.
WOLLIN, A.C., and J.R. PFEIFFER. 1955. Oregon maple log and lumber
grading. Oregon Forest Products Laboratory, State Board of Forestry
and School of Forestry, Corvallis. Report No. G-4. 21 p.
23
BLACK COTTONWOOD
POPULUS TRICHOCARPA
WESTERN COTTONWOOD
BALSAM POPLAR
BALSAM COTTONWOOD
CALIFORNIA POPLAR
Black cottonwood is a large deciduous tree belonging to the willow

family (Salicaceae). It is one of the largest of some 40 species of Populus
and is the tallest, fastest-growing hardwood in the western United States.
Cottonwood is a well-known, common tree along rivers and streams
throughout the West.
Cultivation of hybrid poplars (Populus trichocarpa x. P. deltoides)
can produce very high yields of fiber or fuel in 2-to-8-year rota-
AK tions. Refer to the publication “High yield hybrid poplar planta-
Black Cottonwood
tions in the Pacific Northwest” for detailed information on plan-
YUKON
Range tation culture.

BRITISH
COLUMBIA Mature cottonwoods attain heights of 125 to 150 ft (225 ft
maximum) and diameters of 48 to 60 in. (108 in. maximum). Cot-
ALBERTA
tonwoods mature as early as 60 years and live at least 200 years. In
forest stands, cottonwoods develop narrow, cylindrical crowns and
long clear boles. Open-grown trees develop deep crowns with large
branches, often on single, massive stems. Natural rooting habits
WA are not well known; planted cuttings develop deep, spreading roots.
MT
OR
NV
ID
WY Geographic Range
Black cottonwood grows from Kodiak Island in Alaska (lat 62∞N)
CA
UT
to northern Baja California (lat 31∞N), and eastward to the Rocky
Mountains in Idaho, Montana, and Canada. In the Pacific North-
west, black cottonwood is most abundant in the Puget Sound basin
and in the Columbia and Willamette river basins.
BAJA
24
Timber Inventory
The largest inventory of black cottonwood is in Washington, where
it is third in total volume after red alder and bigleaf maple (Appendix 1,
Table 1). Sixty percent of the total volume (456 MMCF) of black cotton-
wood is found in the Puget Sound subregion. Most of the remaining
volume is evenly distributed between the other regions in Washington
and the Northwest subregion of Oregon.

Black cottonwood is intolerant of shade. It typically maintains a dominant
position in the canopy via superior growth rates. In pure stands, domi-
nance is rapidly expressed and inferior trees die out quickly. Cotton-
woods commonly occur in mixed stands as scattered, emergent trees,
with crowns far above associated species.
Ecological Role
Black cottonwood is a pioneer species specifically adapted to colo-
nizing and dominating areas disturbed by floodwaters. It is generally
perpetuated in such environments by recurring floods. It also colonizes
disturbed upland sites with adequate moisture. Without disturbance,
cottonwood is replaced by other species on upland sites.
Several species of willow are the major associates of black cotton-
wood on recent alluvium along most major rivers in the Northwest.
These species include Pacific, river, northwest, and Scouler willows. Other
common associates are red alder, Oregon ash, bigleaf maple, Douglas-
fir, western redcedar, western hemlock, Sitka spruce, grand fir, birch,
cherry, and hawthorn. Common shrubs include red osier dogwood, vine
maple, hazel, salmonberry, elderberry, thimbleberry, honeysuckle, spirea,
and snowberry. Herbaceous associates include western swordfern, ladyfern,
horsetail, stinging nettle, hedge nettle, false Solomon's-seal, Canada violet
buttercup, bittercress, angelica, enchanter’s-nightshade, golden-saxifrage,
and bedstraw.

There are no guides for estimating site productivity for black cotton-
wood in Oregon and Washington. The British Columbia Forest Service
has identified site-quality classes for natural cottonwood. Most of the
interest in the productivity of managed stands is focused on the inten-
25
sive cultivation of selected clones or hybrids, in which case productivity
is quite dependent on the clone under specific conditions.
Climate
Throughout its wide range, black cottonwood grows in climates that
vary from humid to arid. Cottonwood thrives in the humid climate of
the coastal Northwest, although it grows along rivers in arid regions as
well. In its range, annual precipitation can vary from 10 to 120 in., with
about one-third falling during the growing season. Much of the precipi-
tation may fall as snow. Temperatures range from a maximum of 60 to
117 ∞F to a minimum of -53 to 32 ∞F.
Young trees are susceptible to mortality or damage from early or late
frosts. Frost cracks are common on larger trees. Stems and branches may
bend or break under heavy snow or ice. Wind breakage is also common,
and although their root systems are windfirm, erosion of river banks
commonly topples trees.
Elevation
Black cottonwood grows from sea level to 6000 ft at the northern
end of its range, to 5000 ft in the Cascade Range, and up to 7000 ft in
interior mountains.
Soils
Black cottonwood grows best on deep alluvial soils with good aera-
tion and abundant moisture. It is limited by high soil acidity, inadequate
aeration, and low nutrient supplies. Although it is tolerant of winter
flooding, it requires adequate drainage during the growing season. Some
upland sites with deep, moist, and fertile soils are also productive for
black cottonwood.

Black cottonwood begins to produce seed at about 10 years of age.
Male and female flowers (catkins) are borne on separate trees beginning
in early March (or as late as mid-June in the north). Deciduous male
catkins are 0.8 to 1.2 in. long; female catkins are 3 to 8 in. long when
mature, and bear seed in capsulate fruits 0.2 to 0.32 in. long.
Seed
Black cottonwood seeds are minute, and are tufted with cottony
hairs. Seed generally ripens in late May and June in the Northwest, and
it is rapidly disseminated by wind and water. Viability is usually high,
although it is often short-lived. Seed may remain viable for up to 1 year
when dried and stored at cold temperatures.
26
Black cottonwood generally produces abundant crops of seed every
year. Seedbeds must remain moist for up to 1 month for good germina-
tion and establishment. In much of the Northwest, these conditions are
restricted to wet bottomlands. Cottonwood regeneration is common
after logging on more upland sites in northern Washington.

Black cottonwood sprouts readily from stumps or from buried frag-
ments of branches. Repeated coppicing is quite successful. Regeneration
also occurs from abcised shoots with green leaves, which root where
they fall or are deposited by water.

Plantations of black cottonwood are easily established from rooted
or unrooted cuttings, which allows complete control over genetic origin
of stock. Successful plantings are established from unrooted cuttings of
1- to 2-year-old-wood that are 12 to 24 in. long and 0.4 to 1.2 in. in
diameter. Branchwood cuttings from trees as old as 30 years have also
done well. Cuttings are planted in spring to a depth of 12 to 16 in.,
with at least one leaf bud aboveground. Long cuttings of up to 10 ft
have been used successfully in some cases; problems with inadequate
rooting and poor stem form are common with long cuttings, however.

Newly planted or establishing black cottonwoods are extremely sen-
sitive to vegetative competition, particularly from grasses and herbs.
Cottonwood often avoids competition from other species by establish-
ing at high densities on new substrates and overwhelming any compet-
ing species. Complete site preparation and continued control of com-
peting vegetation is essential for good performance in plantations. This
is often accomplished with repeated mechanical cultivation and applica-
tions of herbicide.
Stand Management
Diameter growth of black cottonwood is very responsive to increased
growing space. Management should target a spacing regime that produces
optimal growth, while maintaining benefits of crowding in young cotton-
wood stands. Moderate crowding produces rapid crown closure (2 to 4
years), which suppresses competing vegetation and eliminates the need for
further vegetation management treatments. For sawlog or veneer produc-
tion, moderate crowding is necessary to maintain stem form, reduce branching
and forking, and induce self-pruning. Typical spacings for pulpwood rota-
tions are 6 X 6 ft to 7 X 10 ft, which accommodate tractors. Longer
rotations require thinning to maintain diameter growth.
27
Black cottonwood must maintain a dominant crown position to sur-
vive and grow. To maintain other species with cottonwood, spacing and
proportion of the super-dominant cottonwood must be controlled. A
mixture of red alder and black cottonwood can improve the growth of
cottonwood via nitrogen fixed by the red alder.
Growth and Yield

Height growth of black cottonwood often exceeds 5 ft per year for
at least 10 years, and trees may reach diameters of 6 to 8 in. in that
time. Various trial plantings indicate that annual yield rates from man-
aged cottonwood can average 150 to 300 ft 3 per acre for up to 24
years. Pulpwood can be grown in 10 to 15 years and sawlogs in 20 to
25 years.

Because of its large size and relatively long life, black cottonwood is
a superior wildlife tree in riparian areas. Fungal decay is common in
wood after weather damage to tops, producing good conditions for
cavity-nesting birds. Raptors such as osprey frequently nest in cotton-
wood. Large trees that have toppled into streams provide structure for
aquatic habitat.
Animals significantly hinder establishment of black cottonwood plan-
tations. Voles and mice cause serious losses on grassy or herbaceous
sites. Beaver are often within range, and remove many trees for dams
and food. Elk and deer can devastate plantings with browsing and antler
rubbing. Protection or prevention of damage may be necessary to avoid
severe losses.

Many insects feed on black cottonwood, but none have caused seri-
ous problems as yet. Fungal diseases are a significant threat. Decay
fungi are common; two species are noted for significant damage in
British Columbia, Spongipellis delectans and Pholiota destruens. A leaf rust
(Melampsora sp.) causes significant problems with clones from dry areas
that are planted in western Washington. Cytospora canker (C. chrysosperma)
is common in natural forests and may be a threat to regeneration from
cuttings.
Genetics
Black cottonwood has outstanding potential for genetic improve-
ment and manipulation with superior clones and hybrids. Substantial
natural genetic variation within the species facilitates the development
of hybrids with a variety of characteristics. Currently, the best hybrids
are those between selected strains of P. trichocarpa and P. deltoides, or
strains of P. maximawitzii and P. deltoides.
28

Diameter at breast height and total height can be used with tables
or equations to estimate total tree volumes in cubic feet and sawlog
volume. Mechanized harvesting is quite suitable for typical plantations
on agricultural lands (fiber farms). Harvesting of native black cotton-
wood may often be subject to restrictions for the protection of riparian
zones.
Logs must be manufactured and delivered very soon after trees are
felled. Black cottonwood is one of the least durable species when placed
in contact with the soil or exposed to the weather. Logs left in damp
conditions show signs of decay within a week or two.
Product Recovery
Most cottonwood is used for pulp and paper because of its softness,
light color, and ease of bleaching. Black cottonwood has been success-
fully sawn into lumber and peeled for veneer. Demand for black cotton-
wood lumber is occasionally good. Lumber and veneer are often consid-
ered to be of relatively low quality and are used primarily for cores in
plywood or interior furniture parts.
Wood Properties
Characteristics
Black cottonwood is a soft, light wood of uniform texture. The sap-
wood is almost white, and merges into the light grey or greyish-brown
heartwood, which may contain dark streaks. The wood is considered
semi-ring to diffuse porous. Vessels are moderately few to very numer-
ous; pores are numerous, small, and barely visible to the naked eye. The
growth rings are inconspicuous, and are differentiated by a thin group-
ing of pores, which gradually increase in density at the end of the
growth increment. The rays are very fine and scarcely visible with a
hand lens. The wood is straight-grained and odorless and tasteless when
dry. When the wood is wet, it has a characteristic disagreeable odor.
Weight
Black cottonwood weighs about 46 lb/ft 3 when green and 24/ft 3 at
12 percent moisture content (MC). The average specific gravity is 0.32
(green) or 0.37 (ovendry).
29
Black cottonwood is a weak wood that is rarely used in applications
where high strength is required. Bending strength is classed as low,
while in stiffness and toughness, black cottonwood is rated as interme-
diate. Because of its low density, cottonwood does not readily split
during nailing. The nail-holding strength of the wood is low. In bending
applications, almost 70 percent of the pieces failed. Despite these ap-
parent shortcomings, cottonwoods show some potential as framing ma-
terials because their strength properties are comparable to those of
currently used softwood species of similar densities. Appendix 1, Table
3 provides information on clear strength values, as well as comparative
values for similar species.

The most important consideration in drying cottonwood is recogniz-
ing its extremely high MC. Heartwood MC averages 162 percent, sap-
wood MC averages 146 percent. The high MC, however, does not cause
an unnecessarily elongated schedule. Green 4/4 lumber can be dried in
8 to 12 days, and air-dried material can be dried in 4 to 8 days. Cotton-
woods have a high tendency to warp during drying because of tension
wood and wet pockets. Average shrinkage from green to ovendry is 3.6
percent radially and 8.6 percent tangentially. Table 2 provides a dry-kiln
schedule appropriate for the indicated thicknesses.
Table 2. Kiln schedule—Black cottonwood* 4/4, 5/4, 6/4.

Equilibrium
1 above 70 140 130 12.0 75 60.0 54.5

2 70 to 60 140 126 10.0 66 60.0 52.0
3 60 to 50 140 120 8.0 55 60.0 49.0
4 50 to 40 140 105 4.9 31 60.0 40.5
5 40 to 30 140 90 2.9 15 60.0 32.0
6 30 to 25 150 100 3.2 18 65.5 37.5
7 25 to 20 160 110 3.4 21 71.0 43.5
8 20 to 15 170 120 3.5 24 76.5 49.0
9 15 to final 180 130 3.5 26 82.0 54.5
*For wood with wet streaks, additional schedules are available.
Machining
Cottonwoods generally machine poorly. Surface roughness and torn
grain result from planing and shaping; bore holes in cottonwood show
considerable roughness and variation in size. The wood is subject to
30
fuzzing and scratching when sanded, which contributes to problems in
finishing. Tool wear and dulling is low because of the wood’s low den-
sity. Best results in planing were obtained with a hook angle of 15∞.
Adhesives
Cottonwood bonds easily with adhesives of a wide range of proper-
ties and under a wide range of gluing conditions. Weak joints are some-
times created if low-viscosity adhesives are used, or if glue spread are
too low. This is because cottonwood absorbs the adhesive more readily
than other hardwoods.
Finishing
Cottonwood is generally painted rather than clear-coated or stained
in applications that require finishes. Cottonwood’s paintability rating
falls in the middle range when compared to other hardwoods. Its resis-
tance to cupping and checking associated with outdoor exposure and
weathering is very poor.
Durability
Black cottonwood is a nondurable species that will degrade rapidly
if exposed to conditions favorable to decay organisms. Average service
life of untreated cottonwood posts in ground contact is 4 to 5 years.
Heartwood is classed as moderately difficult to penetrate with wood
preservatives. The wood is moderately susceptible to sap stains and very
susceptible to molds. Oxidative stains are sometimes present in cotton-
wood.
Uses
Cottonwood is used for veneer, pulp, pallets, interior case good
parts, boxes, crates, moulding (painted), and studs.
Related Literature
BASSMAN, J.H., and J.C. ZWIER. 1993. Effect of partial defoliation on

growth and carbon exchange of two clones of young Populus trichocarpa
Torr. & Gray. Forest Science 39:419-431.
BOWERSOX, T.W., L.R. STOVER, C.H. STRAUSS, and P.R. BLANKENHORN.
1992. Advantages of an effective weed control program for Populus
hybrids. Tree Planters Notes 43(3):81-86.
BRAATNE, J.H., T.M. HINCKLEY, and R.F. STETTLER. 1992. Influence of
soil water on the physiological and morphological components of
31
plant water balance in Populus trichocarpa, Populus deltoides and their
F1 hybrids. Tree Physiology 11:325-339.
DeBELL, D.S. 1990. Black cottonwood. P. 570-576 in Silvics of North
America. Volume 2, Hardwoods. R.M. Burns and B.H. Honkala, coords.
USDA Forest Service, Washington D.C. Agriculture Handbook 654.
HEILMAN, P.E., R.F. STETTLER, D.P. HANLEY, and R.W. CARKNER. 1990.
High yield hybrid poplar plantations in the Pacific Northwest. Wash-
ington State Cooperative Extension; Oregon State University Exten-
sion Service; University of Idaho Cooperative Extension System; USDA,
[Olympia, Washington]. Pacific Northwest Bulletin PNW 356.
HEILMAN, P.E., and F.G. XIE. 1993. Influence of nitrogen on growth
and productivity of short-rotation Populus trichocarpa X Populus deltoides
hybrids. Canadian Journal of Forest Research 23:1863-1869.
HSIANG, T., and G.A. CHASTAGNER. 1993. Variation in Melampsora
occidentalis rust on poplars in the Pacific Northwest. Canadian Jour-
nal of Plant Pathology 15:175-181.
KELLOGG, R.M., and E.P. SWAN. 1986. Physical properties of black cot-
tonwood and balsam poplar. Canadian Journal of Forest Research
16:491-496.
McLENNAN, D.S. 1990. Spatial variation in black cottonwood (Populus
trichocarpa) foliar nutrient concentrations at seven alluvial sites in
coastal British Columbia. Canadian Journal of Forest Research 20:1089-
1097.
ROGERS, D.L., STETTLER, R.F., and P.E. HEILMAN. 1989. Genetic varia-
tion and productivity of Populus trichocarpa and its hybrids. III. Structure
and pattern of variation in a 3-year field test. Canadian Journal of
Forest Research 19:372-377.
SACHSSE, H. 1975. A comparative of important wood properties of the
Populus trichocarpa cultivar “Senior.” Mitteilungen. Verein fuer Forstliche
Standortskunde und Forstpflanzenzuchtung 24:68-77.
32
CALIFORNIA BLACK OAK
QUERCUS KELLOGGII
BLACK OAK
KELLOGG OAK
California black oak is a deciduous member of the beech family

(Fagaceae). It is the most abundant oak on the West Coast and is most
similar in form and wood quality to eastern red oaks.

Mature California black oaks (>90 years old) attain heights of 50 to
110 ft (130 ft maximum) and diameters of 14 to 40 in. (108 in. maxi-
mum), and may live 500 years. In closed stands on good sites, Califor-
California nia black oaks develop narrow, thin crowns on straight, clear boles.
Black Oak Open-grown trees have broad crowns with multiple stems that fork
Range
repeatedly. A scrubby form of California black oak is common on mar-
BRITISH ginal sites. The root system is usually composed of a surface root system
COLUMBIA and several deep vertical roots, which may spread laterally over bed-
rock; some roots penetrate cracks in the rock. Seedlings have a tap root.
WA
Geographic Range
OR
The native range of California black oak extends from Eugene, Or-
egon (lat 44∞N), to the San Pedro de Martin Mountains of Baja Califor-
nia (lat 32∞N). It grows in the valleys of southwestern Oregon and is
abundant on the west side of the Sierra Nevada and in the northern and
central Coast Range of California.
Timber Inventory
CA
Among hardwoods on the West Coast, the inventory of California
black oak (2662 MMCF) is second only to that of red alder. In the
33
Pacific Northwest, it is restricted to southwestern Oregon, with a total
volume of 131 MMCF (Appendix 1, Table 1).

California black oak is intolerant of shade for most of its life. Young
seedlings can persist in the shade; saplings can survive as intermediate
trees, growing tall and thin towards the light. California black oak will
grow towards openings, leaning as much as 15 to 20 degrees. Older
trees cannot survive continued overtopping.
Ecological Role
California black oak may function as a climax species in transitional
environments between conifer forest and chaparral. Over much of the
range, it is probably a persistent subclimax species maintained by resprouts
after periodic fire. On better sites in the absence of disturbance, Califor-
nia black oak is eventually replaced by more shade-tolerant or competi-
tive associates (tanoak, Douglas-fir, California white fir, pines). Under
harsh conditions, conifer regeneration is often restricted to sheltered
areas under black oak; the black oak serve as nurse trees.
Although fire kills trees of all ages, periodic fires probably have maintained
California black oak populations in many areas. Populations appear to be
declining after decades of fire suppression. Prescribed burns of moderate
to low intensity are recommended to improve regeneration of California
black oak from seed.
California black oak is the dominant tree over large areas classified as
the black oak forest type, and it is a major component in other forest
types dominated by conifers. The most common tree species associated
with California black oak are ponderosa pine, Douglas-fir, tanoak, Pacific
madrone, and Oregon white oak. Common shrub associates include greenleaf
manzanita, whiteleaf manzanita, deerbrush, bear-clover, oceanspray, and
poison-oak. Understory vegetation is generally sparse under California
black oak, although shrubs may become abundant and competitive after
fire or cutting.

To evaluate site productivity for California black oak, site index should
be estimated where possible, using methods described by Powers (1972).
In general, good ponderosa pine sites in the range of California black
oak are also good black oak sites. The presence of tanoak with California
34
black oak is also an indicator of good productivity for black oak. Early
growth of stump sprouts is often independent of site quality; thus, older
trees should be used to assess productivity.
Climate
California black oak is adapted to a climate characterized by hot, dry
summers and cool, moist winters. Its climate has an average annual
precipitation range of 30 to 70 in., with extremes of 12 to 110 in. Less
than 5 percent of this moisture falls from June to September. California
black oak grows best in a zone where 10 to 50 percent of the precipita-
tion occurs as snow. Mean daily temperatures in this climate range from
a minimum of 31 to 46 ∞F in January to a maximum of 66 to 82 ∞F in
July.
California black oak has a high tolerance to drought. Deep roots
help it avoid drought; it also withstands high levels of moisture stress.
Black oak leaves are injured by extreme heat after cool, wet weather.
Planted seedlings are susceptible to dieback from late spring frosts. Branches
and boles may break from heavy snow or ice, particularly at forks. Sound,
healthy trees are windfirm.
Elevation
In Oregon, California black oak is found at elevations of 450 to 3000
ft. In California, it is most abundant at elevations of 1000 to 6000 ft in
the north and 4000 to 7800 ft in the south. It is often restricted to
north aspects at low elevations, and south to west aspects at high eleva-
tions. At middle elevations, it is found on all aspects.
Soils
California black oak grows best on deep, well-drained soils of me-
dium to coarse texture. It is found on soils from a wide variety of parent
materials across a wide range of textures. California black oak is often
found on shallow, rocky soils, although its growth and form are poor. It
seldom grows on clay soils, particularly clay topsoils. It does not tolerate
poor drainage or flooding. California black oak may prefer relatively
high levels of soil nutrients; fertilization greatly stimulates growth of
seedlings in the wild.

California black oak starts to produce seed as early as 30 years of
age, but does not usually produce heavily before age 80. It flowers in
spring, from mid-March to mid-May, depending on the environment.
Separate male and female flowers are borne on the same plant. The
greenish-red male flowers arise from leaf axils of the previous year,
forming hairy aments 1.4 to 3.0 in. long. Female flowers emerge from
leaf axils of the current year.
35
Seed
California black oak acorns mature during the second summer after
pollination. The acorns are 0.7 to 1.7 in. long and 0.4 to 1.5 in. wide,
and number from 52 to 147/lb. Acorns should be collected from late
September to early November. The first acorns that fall (mid-August to
mid-September) are usually infested with insects. Acorns should be col-
lected soon after they fall to reduce losses in viability from extreme
temperatures and losses to animals.
California black oak acorns require after-ripening and should either
be planted immediately or stored under cool, moist conditions (33 to
34 ∞F) until spring planting. Germination capacity is quite variable, ranging
from 21 to 95 percent; average rates were 31 to 38 percent in one
large-scale test.

Natural regeneration from seed is uncertain and poorly distributed.
Establishment of seedlings is most frequent under parent trees. Animals
transport many seeds and facilitate the occasional establishment of seedlings
away from the parent tree. Seedlings establish best on undisturbed litter
or loose, well-aerated mineral soil. California black oak does not usually
colonize compacted or heavy clay surface soils. Seedlings rapidly de-
velop deep taproots (to 30 in.) the first year; shoot growth remains slow
for the first 6 or more years.

Most regeneration of California black oak results from basal sprouts,
which are profuse after cutting or burning. Larger parent trees produce
more abundant and more vigorous sprouts than do smaller trees. Sprout
development is best in completely open conditions; shelterwood cut-
tings are not recommended for regenerating black oak sprouts. Stumps
should be cut low to the ground in order to produce more vigorous,
well-formed sprouts.

The performance of California black oak planted on several sites in
California indicates fair potential for regenerating black oak in planta-
tions. Fertilization greatly enhances growth of planted trees. Drought
and pocket gophers are the most common causes of mortality. Late
spring frost causes top dieback; injured seedlings typically resprout.

Little site preparation is necessary for establishing stands from sprouts.
Regeneration and growth may be enhanced by burning or mechanically
removing slash that shades stumps.
On California black oak sites, shrubs are often sparse and not competi-
tive. If vegetative competition is significant on a site, however, vegetation
36
management will benefit California black oak sprouts or seedlings. Seed-
lings are most vulnerable to competition. Site preparation and later control
of competing vegetation may accelerate the typically long transition stage
from seedling to sapling.
Stand Management
California black oak sprouts initiate at high densities, after which
self-thinning and expression of dominance proceed rapidly. Thinning
young sprout clumps after 4 years is probably not beneficial. Thinning
in older stands can improve diameter growth (up to 2 times) while
favoring better quality trees. Results of one thinning study indicate that
optimum stand and tree growth may be maintained at stand densities
of 100 to 125 ft 2 per acre. Thinning can increase the size and quality of
epicormic branches. Density management strategies must be designed
to minimize the impact of epicormic branches on wood quality. Califor-
nia black oak is best managed in even-aged stands or patches.
Top light must be maintained for good growth of California black
oak in mixed stands. On good sites, associated tanoak and conifers may
need to be controlled to maintain California black oak in the long term.
Black oak may be used as a nurse tree to facilitate survival of conifer
regeneration on low-elevation sites. Black oak is resistant to the annosus
root rot (Heterobasidion annosum), which suggests a strategy of planting
black oak in root-rot areas within conifer stands.
Growth and Yield

Seedlings grow slowly, reaching heights of 4 to 6 in. the first year,
and often have slower shoot growth for 5 or more years while roots
establish. Sprout growth averages about 2 ft per year for the first 10
years. Site index (50-year base) ranges from 30 to 70 ft, with an aver-
age of 50 ft. In natural stands, average diameter increment is about 1.8
in. per decade during the first 60 years, slowing to about 1.5 in. per
decade by age 110.
California black oak stands in California average about 1213 ft 3 per
acre, with maximum volumes of 4000 ft 3 per acre over 5 acres. One
fully stocked stand of 70-year-old black oak had a volume of 5845 ft 3
per acre with an average diameter of 12 in. and height of 62 ft.

California black oak acorns are an important food source for birds,
rodents, deer, and bear. Fluctuations in deer populations are sometimes
correlated with the acorn crop. Large, hollow trees are common and
provide habitat for cavity-nesting animals. Foliage is browsed by deer
and elk.
37
Many species of insects live on California black oak, but they seldom
have significant impact. The carpenter worm (Prionoxystus robinae) damages
the wood of black oak; other insects causing damage include the pit
scales (Asterolecanium minus and A. quericola), the Pacific oak twig girdler
(Agrilus angelicus), the California oakworm (Phryganidia californica), and
the fruit-tree leafroller (Archips argyrospila).
Heart rots (Inonotus dryophilus and Laetiporus sulphureus) cause sig-
nificant damage in old trees and in trees injured by logging, fire, or
weather. The shoestring root rot, Armillaria ostoyae (A. mellea) com-
monly attacks older trees. Black oak is resistant to the annosus root rot
(Heterobasidion annosum), which kills many other species.
Genetics
California black oak hybridizes with Q. agrifolia (known as Quercus x
ganderi) and with Q. wislizenii (known as Quercus x moreha). The latter
hybrid is most common and has sparse evergreen foliage.

or equations to estimate total tree volume in cubic feet and sawlog
volume. California black oak trees have a tendency to split and “barber-
chair” during felling so care must be taken in making the undercut and
leaving holding wood.
Logs are generally weighed or sold by the truckload, and conversion
factors are used to convert back into board foot volumes. The logs
check easily during storage and should be end-coated to prevent splits
in the lumber. Log grades have also been used to effectively separate
logs of different product quality and are recommended for marketing.
Product Recovery
Sawlogs usually have a minimum small-end diameter of 10 in. Smaller
logs are chipped for pulp. Stain, discoloration, and end- and surface-
checking are common problems with California black oak lumber if it is
not processed within a short time after it is removed from the woods.
One study found that 72 percent of the board-foot-scale was recovered
as lumber; another study found that 60 percent of the weight is recov-
ered as lumber products, 20 percent as slabs and edgings, and 20 per-
cent as sawdust and shavings. The percentage of loss associated with
edgings may be reduced if care is taken to prevent “over-edging” in the
mill. Recovery of higher grade lumber from California black oak appears
to be somewhat low compared to other hardwood species (Appendix 1,
Table 2).
38
Wood Properties
Characteristics
The heartwood is light brown with pink to pale reddish-brown color;
the sapwood is a pale yellowish-white to brownish-white. California black
oak is a ring-porous wood, with earlywood pores that are large and
distinct and form a conspicuous band with each growth ring. The late-
wood pores are small and numerous, and require a hand lens to view.
The large earlywood vessels are almost always occluded by tyloses. Among
the red oaks, California black oak has one of the lowest percentages of
summer wood; for an oak, then, the wood is fairly fine-grained. The
rays are numerous, short in height, and wide. When the wood is dry, it
has no characteristic odor or taste. Distinctive burls are sometimes present.
California black oak is commercially classed as a red oak in USDA Forest
Service nomenclature.
Weight
California black oak weighs about 66 lb/ft 3 when green and 40 lb/ft 3
at 12 percent moisture content (MC). The average specific gravity is
0.51 (green) or 0.58 (ovendry).
Mechanical properties
Because of its lower specific gravity and lower percentages of sum-
mer wood, California black oak has lower clear-specimen strength val-
ues than many of the eastern red oaks. It still is an oak, however, and
possesses many desirable strength properties, including parallel and per-
pendicular compression resistance, and side hardness. It is suitable for
most furniture design applications, and the lower grades perform well
as pallet stock. It holds nails well, but will split unless it is prebored or
pneumatic nailers are used. See Appendix 1, Table 3 for average me-
chanical properties for small, clear specimens.

California black oak requires special care and attention to detail
during an extended kiln schedule to properly reduce MC to a level
suitable for interior products such as flooring, furniture, or millwork.
Drying defects can cause serious downgrade; end- and surface-checking
result from uncontrolled or overly rapid drying; honeycomb, collapse,
and ring failure occur from wetwood; iron stains form when tannins
contact certain metals; and grey sapwood staining will result if there is
poor air circulation. Additionally, the wood can be inhabited by bacte-
ria, which will complicate drying but will not affect the quality of the
final dry product. The green MC of California black oak averages 105
percent (ovendry basis). Shrinkage values for green to ovendry (based
on the original green size) are low, and average 3.6 percent radially and
39
6.6 percent tangentially. It is suggested that all the upper grades be air-
dried to 20 percent MC and then kiln-dried according to a time sched-
ule (See Table 3 for the appropriate kiln schedule).
Table 3. Kiln schedule—California black oak 4/4, 5/4, 6/4.

Equilibrium
1 Above 35 110 107 19.1 90 43.5 41.5

2 35 to 30 110 106 17.6 87 43.5 41.0
3 30 to 25 120 114 15.5 83 49.0 45.0
4 25 to 20 130 120 12.2 74 54.5 49.0
5 20 to 15 140 115 6.8 46 60.0 46.0
6 15 to final 160 110 3.4 21 71.0 43.5
Machining
The machining characteristics of California black oak are excellent.
Because of its moderate specific gravity and tight grain, the feed speeds
of machines can be greater for California black oak than for most of the
other oaks, but still produce quality surfaces when planing, shaping,
turning, boring, and sanding. The wood can be successfully bent when
it is properly steamed and bending forms are utilized.
Adhesives
California black oak bonds satisfactorily and there are no unusual
problems when gluing conditions are well controlled. Careful curing/
drying of glue joints is required to prevent sunken gluelines from subse-
quent machining.
Finishing
California black oak finishes well, although it may be necessary to
fill the grain to obtain a smooth finish. The heartwood/sapwood color
variation can present difficulties if uniform color is desired. Dyes and
transparent stains are better than heavily pigmented stains, which re-
quire the removal of any excess pigment from the wood.
Durability
The heartwood of California black oak is basically nondurable when
exposed to conditions that are favorable to wood decay organisms. Iron
staining will occur if ferrous products contact wet wood. Oxidative staining
can additionally degrade improperly handled logs and lumber.
40
Uses
Black oak has been successfully peeled into veneer and used in cabi-
net fronts. It is used for moulding, millwork, paneling, furniture, floor-
ing, veneer, edge-glued panels, pallets, chips for landscaping, and fire-
wood.
Related Literature
HALL, G., and R. ALLEN. 1981. Wood products from California oaks, Cal
Oak Lumber Company style. P. 362-368 in: Proceedings of the Sym-
posium on the Ecology, Management, and Utilization of California
Oaks. T.R. Plumb, tech. coord. USDA Forest Service, Pacific South-
west Forest and Range Experiment Station, Berkeley, California. Gen-
eral Technical Report PSW-44.
KAUFFMAN, J.B., and R.E. MARTIN. 1987. Effects of fire and fire suppres-
sion on mortality and mode of reproduction of California black oak
(Quercus kelloggii Newb.). P. 122-126 in Proceedings of the Sympo-
sium on Multiple-Use Management of California’s Hardwood Resources.
T.R. Plumb and N.H. Pillsbury, tech. coords. USDA Forest Service,
Pacific Southwest Forest and Range Experiment Station, Berkeley,
California. General Technical Report PSW-100.
KOENIG, W.D., R.L. MUMME, W.J. CARMEN, and M.T. STANBACK. 1994.
Acorn production by oaks in central coastal California: variation within
and among years. Ecology 75:99-109.
MALCOLM, F.B. 1962. California black oak - a utilization study. USDA
Forest Service, Forest Products Laboratory, Madison, Wisconsin. Re-
port No. 2237. 10 p.
McDONALD, P.M. 1969. Silvical characteristics of California black oak
(Quercus kelloggii Newb.) USDA Forest Service, Pacific Southwest Forest
and Range Experiment Station, Berkeley, California. PSW-53.
and California black oak in north-central California. USDA Forest Ser-
vice, Pacific Southwest Forest and Range Experiment Station, Berke-
ley, California. Research Note PSW-362. 6 p.
McDONALD, P.M. 1990. California black oak. P. 661-671 in Silvics of
North America. Volume 2, Hardwoods. R.M. Burns and B.H. Honkala,
coords. USDA Forest Service, Washington D.C. Agriculture Handbook
654.
tems for the Major Forest Types of the United States. R. Burns, tech.
compil. USDA Forest Service, Washington, D.C. Agriculture Hand-
book 445.
PLUMB, T.R., and P.M. McDonald. 1981. Oak management in California.
USDA Forest Service, Pacific Southwest Forest and Range Experiment
Station, Berkeley, California. General Technical Report PSW-54. 11 p.
41
POWELL, C. 1978. The California black oak: from saddle-soaped leather
to pumpkin orange. Environment Southwest 481:3-50.
POWERS, R.F. 1972. Site index curves for unmanaged stands of Califor-
nia black oak. USDA Forest Service, Pacific Southwest Forest and
Range Experiment Station, Berkeley, California. Research Note PSW-
262. 5 p.
QUARLES, S.L. 1992. Acoustic emission associated with oak during dry-
ing. Wood and Fiber Science 24:2-12.
42
CALIFORNIA-LAUREL
UMBELLULARIA CALIFORNICA
PACIFIC-MYRTLE
OREGON-MYRTLE
CALIFORNIA-BAY
PEPPERWOOD
SPICE-TREE
California-laurel is the only tree of the family Lauraceae found in the

western United States. It is a broadleaved evergreen tree with distinctly
aromatic “bay” leaves. Often referred to as myrtlewood, California-laurel
is one of the best known and most valuable western hardwoods.

California-laurel Typical California-laurels are 40 to 80 ft tall and 18 to 30 in. in
Range
diameter. On good bottomland sites, mature trees can attain diam-
eters of 36 to 72 in. (159 in. maximum) and heights of over 100 ft
BRITISH (175 ft maximum). California-laurels may live at least 200 years. The
COLUMBIA
trees often have forked or multiple stems with ascending branches, which
form dense, round-topped crowns. In forest stands, the stems or limbs
are relatively straight, vertical, and clear of smaller branches. Open-
WA grown trees have broad spreading crowns (often wider than tree height)
supported by several main stems or branches. The root system of Cali-
fornia-laurel is wide and spreading, although it varies from shallow to
OR deep, depending on the soil and drainage.
ID
Geographic Range
The range of California-laurel extends from Reedsport, Oregon (lat
44∞N) to San Diego, California (lat 33∞N). It is not found more than 160
CA miles from the Pacific Coast. California-laurel is found in the Coast Ranges,
the southwestern Cascade Range, and all along the western Sierra Ne-
vada.
43
Timber Inventory
The total inventory of California-laurel is about 520 MMCF of grow-
ing stock, of which 93 MMCF occurs in southwestern Oregon (Appen-
dix 1, Table 1, excluding federal lands in southwestern Oregon, for
which there are no recent estimates). Many of the best trees are found
in parks and riparian areas. According to some representatives of the
myrtlewood industry, available supplies of the high-value, figured wood
are getting scarce.

California-laurel is intermediate in tolerance of shade. Seedlings estab-
lish and grow at low light levels (1 to 18 percent of full sun), and saplings
and intermediate trees are common with coniferous overstories.
Ecological role
California-laurel can be a climax species, as it is long-lived and re-
produces in the understory. A component of California-laurel is typically
maintained by resprouting after fires in subclimax forests, which are
most common outside of riparian areas. The leaf litter from California-
laurel may have toxic effects on other vegetation; these toxic effects are
a suspected cause for the typically sparse cover under California-laurel
trees.
On good sites, pure stands of California-laurel are generally restricted
to small patches, while short or shrubby California-laurel often domi-
nate more extensive areas. California-laurel is most commonly found in
mixed stands; a great number of tree species are associated with laurel
across its range. Common tree associates in the Pacific Northwest in-
clude red alder, cottonwood, willows, tanoak, Pacific madrone, wax
myrtle, Oregon ash, Sitka spruce, Douglas-fir, Port-Orford-cedar, and
redwood. Common shrub associates include salmonberry, evergreen
huckleberry, red huckleberry, dewberry, snowberry, poison-oak, Pacific
rhododendron, Oregon-grape, serviceberry, and honeysuckle.

The best conditions for growth of California-laurel occur on moist,
protected bottomlands or lower slopes with deep soils. Large trees can
develop under these conditions, even in otherwise hot and dry locali-
ties. California-laurel will establish and grow on a much wider range of
44
sites, but it is limited to a shrubby or prostrate form on dry, rocky, or
exposed sites.
The capability of a site for growing California-laurel should be evalu-
ated by examining growth and form of older trees. Good growth poten-
tial is indicated by the following characteristics:
• Top height of at least 80 ft on mature trees, up to 150 ft on the
best California-laurel sites
• Sustained height growth of 1 to 2 ft per year between ages 5 to
20 years
Climate
California-laurel grows under conditions that range from the cool,
humid climate of coastal forests to the hot, dry climates of the chaparral
or interior woodlands. Within its range, annual precipitation varies from
13 to 83 in., with 0.7 to 17.0 in. falling from April to September.
Temperature extremes are -13 to 118 ∞F, although a milder climate is
more typical, with average minimums of 31 to 50 ∞F (January) and
average maximums of 56 to 84 ∞F (July).
Moisture is the major factor limiting growth of California-laurel. Lau-
rel will establish and grow in hot, dry regions but its distribution is
more limited to moist microsites. Considerable damage and breakage of
stems and branches is caused by snow and wind. Windthrow of laurel is
common on wet soils. California-laurel crowns are commonly deformed
in areas of frequent wind on coastal or ridge sites.
Elevation
Over most of its range, California-laurel grows at elevations from sea
level to 4000 ft. At the southern limit of its range, it grows from 2000
to 5000 ft.
Soils
California-laurel grows on a wide variety of soils derived from allu-
vial, sedimentary, volcanic, or metamorphic parent materials. The best
growth occurs on deep, well-watered, and well-drained soils, typically
on alluvial benches, valley bottoms, and coastal slopes.

Flowers may occur on stems as young as one year of age; abundant
fruiting usually occurs after 30 to 40 years. Flower buds form during the
fall before flowering, and flowers emerge before new leaves, beginning
as early as November (far south) and continuing through late spring.
Pale yellow flowers (perfect), about 0.6 in. in diameter grow in clusters
(umbels) originating from leaf axils. The fruits are round drupes, which
ripen in the first autumn after flowering, changing in color from green
to yellow, brown, or purple when ripe.
45
Seed
Each fruit contains one nutlike seed (0.6 in. in diameter); there are
about 300 seeds/lb of fruit. California-laurel seed may be collected from
the ground during late fall and winter, although it should be collected
soon after it falls to reduce losses in viability from exposure. Germina-
tion and seedling emergence occur in autumn, soon after seedfall or in
late winter and spring.
Seed should be planted immediately or stored under cool, moist
conditions (37 ∞F for up to 6 months) until spring planting. Germina-
tion of fresh seed may take up to 3 months; scarification or stratifica-
tion can shorten this time to 2 months. Rates of germination are not
affected by light; the highest rates occur with high humidity in moist,
but not wet soil (moisture tension of -4 to -10 bar).

Abundant crops of seed are usually borne every year. Establishment
from seed is best where seed is covered by soil, either from soil distur-
bance or from soil deposition after high water. Seedlings commonly
establish and grow at low light levels (1 to 8 percent of full sun) under
forest or brush canopies, although growth is increased with light levels
that exceed 18 percent of full sun.

California-laurel sprouts vigorously from the root collar or along the
trunk with increased exposure to light. To promote development of
better quality sprouts, stumps should be cut close to the ground.

Regeneration of California-laurel with planted seedlings has been rare.
Seedlings typically develop a taproot and may be difficult to transplant
unless they are grown in containers. Seedlings grow slowly for the first few
years after transplanting. Under culture, seedlings appear to grow well
across a range of temperature, moisture, and nutrient conditions.

Little site preparation is necessary for establishing stands of sprout
origin after forest clearing. Regeneration and growth are enhanced by
burning or mechanically removing slash that shades stumps. Pre-estab-
lished roots of the parent tree and rapid growth of sprout clumps make
California-laurel a superior competitor in new stands. Rapid develop-
ment of California-laurel cover at high densities of parent stumps will
inhibit establishment and growth of competing species. With seed-
lings, or at lower densities of California-laurel sprouts, control of com-
peting herbs and shrubs should improve the growth of young Califor-
nia-laurel stems.
46
Stand Management
There has been little management of California-laurel, and there is
no available data from experimental treatments. As demonstrated for
many other sprouting species, thinning dense clumps of California-lau-
rel sprouts may be a viable method for selecting better stems and im-
proving diameter growth. Care must be taken to avoid damage to re-
sidual trees during management activities, since California-laurel is eas-
ily invaded by wood decays. Given its strong tendency to form curved,
multiple stems, maintenance of moderate and uniform stand density
and evenly spaced stems may be needed to encourage straight, well-
pruned stems.
There are many opportunities for growing California-laurel in mixed
stands, as it readily establishes in the understory and may be grown in
any crown position in even-aged or uneven-aged stands.
Growth and Yield

Initial growth of sprouts is rapid; seedlings develop more slowly in
typical understory environments. Long-term height growth of Califor-
nia-laurel is slow (<1 ft per year) on many sites, but rates of 1 to 2 ft
per year are possible on good sites. Diameters of 15 to 16 in. may be
achieved in 50 years.
California-laurel (along with tanoak) has the highest net annual vol-
ume growth rates of any hardwood in California (3.5 percent per year).
Average stand volumes for California-laurel forest types in California
were 1677 ft 3 per acre with maximum volumes of 3125 ft 3 per acre.
Almost half of the California-laurel stands in California exceed 100 ft 2
per acre of basal area.

California-laurel provides food and cover for various animals. Seeds
are an important food source for squirrels, woodrats, mice, and birds.
Deer browse young shoots during the summer. Heart rot in larger trees
often provides cavity nesting habitat.

Old or scarred trees are commonly invaded by decay fungi, particu-
larly the heart rot, Ganoderma applanatum. Decay is common around
branch wounds and root collars. Occasional dieback of branches is caused
by Botryosphaeria spp. A stem canker (Nectria galligena) is common on
stems stressed by wind or snow. Although a variety of pathogens attack
foliage, serious damage is rare.
Insects usually do not cause serious problems for California-laurel.
The cottony cushion scale (Icerya purchasi) has been quite damaging in
47
the past. Some oak bark beetles (Pseudopityophthorus spp.) will attack
injured trees. Powderpost beetles (Ptilinus basalis) can cause serious damage
to stored logs or lumber. A variety of insects consume foliage, although
high concentrations of volatile compounds in California-laurel reduce
the incidence of damage.
Genetics
There are some racial varieties of California-laurel, one of which is
known for its distinctly pendulous branchelets (Umbellularia californica
forma pendula).

volume. Log grades have not been developed. Premium prices may be
paid for burls, butts, and logs 12 in. and larger in diameter.
Product Recovery
Sawlogs usually have a minimum small-end diameter of 6 in. The
relatively small volume of California-laurel that is harvested is fully uti-
lized, and demand for California-laurel wood is high. Lumber grade
recovery has not been studied.
Wood Properties
Characteristics
California-laurel is a moderately heavy, moderately hard wood with
an even texture and a fine grain. The sapwood is whitish to light brown
and typically thick. The heartwood is light brown or greyish-brown,
frequently with darker streaks of pigment figure. The growth rings are
distinct and can be delineated by a dark band of denser latewood. The
wood is diffuse-porous, with evenly distributed, distant small pores that
are barely visible to the naked eye. These pores are either solitary or in
groups of two or three, and are encircled by a whitish sheath. The fine
rays require a hand lens to see. When freshly cut, the wood has a very
characteristic spicy odor, but its volatile oils impart no taste to the
wood. Burls are sometimes produced and some of the wood has inter-
locked grain. When soaked in water, the wood darkens appreciably.
48
Weight
California-laurel weighs about 54 lb/ft 3 when green and 39 lb/ft 3 at
12 percent moisture content (MC). The average specific gravity is 0.51
The strength properties of California-laurel are satisfactory for the
typical uses and product applications. The wood has good resistance to
indentation and splitting, but the wood has moderately low bending-
strength properties. The wood holds nails and fasteners well, but holes
should be prebored to prevent splitting. See Appendix 1, Table 3 for
average mechanical properties of small, clear specimens.

California-laurel is one of the western hardwoods that can be suc-
cessfully kiln-dried when green, without first being airdried. A mild schedule
is recommended to prevent surface checking and honeycomb. A suit-
able general schedule is shown in Table 4. End-checking from refractory
wood can be a problem, so an appropriate end-coating should be ap-
plied. The green MC of the wood averages 70 percent. The shrinkage
values indicate that California-laurel is a relatively stable wood; it has a
radial shrinkage value of 2.8 percent (green to ovendry) and a tangen-
tial shrinkage value of 8.1 percent. For comparison, black cherry has
values of 3.7 percent and 7.1 percent, respectively.
Table 4. Kiln schedule—California-laurel 4/4, 5/4, 6/4.

Equilibrium
1 Above 30 120 113 14.4 80 49.0 45.0

2 30 to 25 130 120 12.2 74 54.5 49.0
3 25 to 20 140 125 9.6 64 60.0 51.5
4 20 to 15 150 125 6.8 49 65.5 51.5
5 15 to 10 180 140 4.5 36 82.0 60.0
6 10 to final 180 130 3.5 26 82.0 54.5
Machining
California-laurel ranks very high in machinability for turning, boring,
and mortising. In planing and shaping, however, chip-out difficulties
arise because of the interlocked grain and the small burls sometimes
present. It is advisable to consider saw-sized lumber and abrasive plan-
ing as alternatives to planer degrade. The fewest defects were obtained
with hook angles of 20∞. Finish sanding requires a generally fine grade
of paper to avoid surface scratching.
49
Adhesives
California-laurel produces satisfactory glue bonds of good strength if
conditions are well controlled.
Finishing
California-laurel is a very beautiful and distinctive wood when fin-
ished. It takes finishes well, without the need to fill the grain. Figured
wood and burls are exceptionally attractive when clear-coated. The wood
is generally not stained, but when color changes are desired, dyes and
transparent stains are usually preferable to pigmented types. Thick sur-
face coatings should be avoided.
Durability
California-laurel is a nondurable species that is susceptible to wood
decay; it is not suitable where conditions are favorable to rot. It is also
susceptible to powder-post beetle infestation and sap staining.
Uses
California-laurel is used for novelties and craft items, wooden ware,
turnery, furniture, paneling, flooring, veneer, and gun stocks.
Related Literature
STEIN, W.L. 1990. California-laurel. P. 826-834 in Silvics of North America.

Volume 2, Hardwoods. R.M. Burns and B.H. Honkala, coords. USDA
Forest Service, Washington, D.C. Agriculture Handbook 654.
TINNIN, R.O., and L.A. KIRKPATRICK. 1985. The allelopathic influence
of broadleaf trees and shrubs on seedlings of Douglas-fir. Forest
Science 31:945-952.
50
GIANT CHINKAPIN
CASTANOPSIS CHRYSOPHYLLA
GOLDEN CHINKAPIN
GIANT EVERGREEN CHINKAPIN
CHINKAPIN
CHINQUAPIN
GOLDENLEAF CHESTNUT
Giant chinkapin, an evergreen member of the beech family (Fagaceae),

is the only tree-sized species of its genus found in the United States.
Most of about 110 different species of Castanopsis occur in China, India,
and Malaysia. With its dense, evergreen foliage (bright golden-yellow
on the underside of the leaf) giant chinkapin is a distinctive, although
minor, component of mixed-evergreen forests in western Oregon.
Giant Chinkapin
Range Size, Longevity, and Form
BRITISH Mature giant chinkapin are typically 60 to 80 ft tall (150 ft maxi-
COLUMBIA
mum) and 12 to 30 in. in DBH (96 in. maximum). Chinkapin may live
400 to 500 years. In forest stands, giant chinkapin develops a dense,
ovoid to conical crown on a straight, clear bole (50-70 percent of bole
length). Chinkapin displays apical dominance even under open condi-
WA
tions. Open-grown trees develop a more spreading crown on a highly
tapered bole. A shrub form of giant chinkapin is also common across
OR the range of the species. Young giant chinkapins often develop a tap-
ID root, while older trees have a well-developed lateral root system.
CA
Geographic Range
Giant chinkapin is native to the West Coast, from west-central Wash-
ington (lat 47∞N) to central California (lat 35∞N). The tree form of chinkapin
is most common in the Coast and western Cascade ranges, from Lane
County, Oregon to northern California.
51
Timber Inventory
The total volume of giant chinkapin in Oregon is about 86 MMCF
(Appendix 1, Table 1) and is about equally divided between the West-
Central and Southwest subregions. Another 50 MMCF occurs in the
Northwest subregion of California. Most giant chinkapin in Washington
are shrubs or small trees; their volume is negligible.

The tree form of giant chinkapin is intermediate in tolerance. Chinkapins
often occur in intermediate canopy positions, although they decline in
vigor and may die after prolonged overtopping by associated conifers
in older stands. Shrub forms of giant chinkapin are quite tolerant of
shade and will persist in the understory.
Ecological Role
Giant chinkapin appears to be most competitive and persistent on
droughty, infertile sites; this is where the oldest trees are usually found.
Young chinkapins are also relatively aggressive and competitive during
early succession on poor sites. Periodic disturbance (fire, logging, wind)
may be required to maintain a component of giant chinkapin on better
sites.
Giant chinkapin rarely occurs in pure stands, but it is a minor com-
ponent in many different forest types. Common associate trees are Douglas-
fir, incense-cedar, sugar pine, Pacific madrone, tanoak, western white
pine, western hemlock, white fir, ponderosa pine, California black oak,
Port-Orford-cedar, canyon live oak, and knobcone pine. Common shrubs
include Pacific rhododendron, salal, Oregon-grape, baldhip rose, dew-
berry, snowberry, oceanspray, hazel, poison-oak, manzanitas, modest
whipplea, and prince’s-pine.

Giant chinkapin can grow relatively well on harsh, droughty, or
infertile sites. Good growth and development of chinkapin can be ex-
pected on better sites, but management may be required to maintain
chinkapin among taller, more competitive associates. The capability of
a site for growing chinkapin should be evaluated by examining growth
and form of older trees. Good growth potential is indicated by the
following characteristics:
52
• Top height of at least 60 ft on mature trees
• Sustained height growth of 1 to 2 ft per year for ages 10 to 30 years
Climate
Giant chinkapin grows in a mild climate characterized by winter rain
and summer drought, wherein annual precipitation ranges from 20 to
130 in., with very little falling from June to September. Snow is com-
mon at higher elevations over chinkapin’s range, particularly in the
Siskiyou Mountains and Oregon Cascade Range.
Judging from its superior competitiveness (relative to associated tree
species) on harsh sites, giant chinkapin appears to have a high toler-
ance to extremes of heat, drought, and cold. The shrub form becomes
more common under extreme climates.
Elevation
Chinkapin grows at elevations from sea level to 6000 ft.
Soils
The tree and shrub growth forms of giant chinkapin are found over
a wide variety of soils derived from parent materials that include basalt,
diorite, sediments, metasediments, and serpentine. The best develop-
ment of giant chinkapin occurs on relatively deep soils that often have
some nutrient deficiencies. The shrub form of chinkapin is predominant
on shallow, rocky, droughty soils; the tree form is typically found on
deeper soils and under more moderate moisture stresses.

Trees of seedling origin produce sound seed at 40 to 50 years of
age. Some fruiting occurs on younger stems; sprouts as young as 6
years old have produced seed.
Giant chinkapin is monoecious, and flowering usually begins in June
or July. Male flowers form in dense catkins 1 to 3 in. long. From one to
three female flowers develop within involucres either at the base of
male flowers or separately on the stem. Flowers are pollinated by wind
and perhaps by insects; chinkapin flowers are thought to impart a bad
taste to honey.
The fruit is composed of one to three nuts within a spiny, golden-
brown bur 0.6 to 1.0 in. across. Fruit ripens in the second autumn after
pollination.
Seed
The seeds (nuts) are about 0.5 in. across and average about 830 to
1100/lb. Nuts are dispersed by gravity or by animals from September to
December. Limited information indicates that germination rates are low
53
(14 to 53 percent) compared to other hardwoods. Germination is not
increased by cold stratification; however, seed germinates in spring un-
der natural conditions.

Little is known of requirements for natural regeneration of chinkapin.
Partial shade and a light litter layer appear to favor seedling establish-
ment. In northern California, establishment of seedlings was best on
moist sites with sparse understory vegetation. Initial growth of seedlings
is slow (6 to 18 in. after 4 to 12 years).

Giant chinkapin produces vigorous basal sprouts after cutting, fire,
or other injury. To promote development of better quality sprouts, stumps
should be cut low to the ground.

There are no documented cases of regeneration of chinkapin from
planted seedlings.

origin. Sprout regeneration and growth may be enhanced by burning or
by mechanically removing slash that shades giant chinkapin stumps.
Site-preparation treatments that leave some leaf mulch and partial
protection (debris, vegetation) may be best for promoting establishment
and growth of chinkapin from seed.
The response of giant chinkapin to control of competing vegetation
has not been studied. Vegetation management may be required to maintain
chinkapin on more fertile or moist sites where other tree species will
eventually suppress it in the absence of disturbance (historically, fire).
Stand Management
Giant chinkapin sprouts initiate at high densities, after which self-
thinning and expression of dominance proceed rapidly. As with tanoak,
thinning young sprout clumps (at ages 3 to 10 years) is probably not
effective because of the abundant resprouts.
There is no information specific to managing giant chinkapin. As for
other sprouting hardwoods, it is probable that growth and quality of
stems may be improved by thinning in older stands of chinkapin.
Giant chinkapin is most commonly found as a minor associate in
mixture with conifers. Particularly on better sites, periodic disturbance
54
(or management) is required to maintain a component of chinkapin.
Under these conditions, maintenance of more open stand conditions
can allow chinkapin to grow tall, relatively well-formed stems.
Growth and Yield

There is no information on volume growth within pure stands or
patches of chinkapin. A chinkapin component of 1100 to 1500 ft 3 per
acre comprised 11 to 21 percent of the total stand volume in 80-to-
100-year-old stands of Douglas-fir (site class III to IV) in three sample
stands in the Coast and Cascade ranges. Average DBH of chinkapin in
these stands ranged from 7.8 to 13.4 in.

There have been few studies of the specific importance of giant
chinkapin to wildlife. Chinkapin nuts probably provide a nutritious food
for various birds and mammals. Typical chinkapin in the understory of
conifer forests provide structural canopy diversity, which can improve
habitat for various animal species.

Insects and pathogens seldom cause serious problems for chinkapin.
Heart rot fungi, including Phellinus igniarius, commonly cause defect in
old or injured stems.
Insects such as the filbertworm (Melissopus latiferreanus) can have
significant impacts on chinkapin seed crops. One study found that nearly
100 percent of the seed crop had been attacked by insects on one of
three study sites. The California oak-worm (Phryganidia californica), which
causes serious defoliation of oak species, also attacks chinkapin and
reduces its growth.
Genetics
The shrub and tree growth forms of giant chinkapin may be geneti-
cally distinct in some cases. There appear to be three ecotypes of chinkapin:
the tree form common at lower elevations, a high-elevation shrub type
adapted to cold temperatures and heavy snow, and a chapparal shrub
adapted to dry sites. Some hybridization may also occur between giant
chinkapin and evergreen chinkapin (Castanopsis sempervirens).
Harvest and Utilization

Diameter at breast height and total height can be used with equa-
tions to estimate total tree volumes in cubic feet and sawlog volumes.
55
Log grades have been applied to giant chinkapin; there were large
differences by log grade in the value of lumber recovered. Purchasers
generally buy chinkapin as sawlogs (>10 in.) or pulp logs, without ap-
plying more detailed log grades.
Product Recovery
Sawlogs usually have a minimum small-end diameter of 10 in.; smaller
logs are chipped for pulp. One study indicates that recovery of No. 1
Common and Better grade green lumber from giant chinkapin com-
pares favorably to other hardwoods; however, Select and Better lumber
has much lower recovery than other hardwoods (Appendix 1, Table 2).
The relatively small volume of chinkapin that is harvested is fully uti-
lized, and demand for lumber is high.
Wood Properties
Characteristics
The wood of giant chinkapin is of moderately fine texture and is
moderately hard and heavy. The thin sapwood is the same color or
slightly lighter than the light brown, pinkish-tinged heartwood. It is a
ring porous wood with large earlywood pores that are generally singu-
lar or, occasionally, in pairs. Emanating radially from the large early-
wood pores are flame-shaped clusters of smaller latewood pores not
readily visible to the naked eye. Still finer are the pores across the
growth ring and between the flame-shaped patterns. The rays are very
fine and are barely visible with a hand lens. When the wood is dry,
there is no characteristic odor or taste.
Weight
Chinkapin weighs about 61 lb/ft 3 when green and 32 lb/ft 3 at 12
percent moisture content (MC). The average specific gravity is 0.42
Because of its limited availability and its uniqueness, giant chinkapin
is rarely used for building or structural applications. Instead, it is most
often used for fine furniture or exceptional paneling. Giant chinkapin
performs well in these applications, if the furniture is adequately de-
signed. See Appendix 1, Table 3 for average mechanical properties of
small, clear specimens.
56
There are some distinct difficulties in kiln-drying green chinkapin.
These, coupled with the uncommonness of this minor species, presently
limit commercial availability. Much of the work on drying giant chinkapin
was done before the mid-1950s, and indicated that chinkapin is subject
to considerable collapse (excessive shrinkage in thickness) unless the
wood is air-dried to below 20 percent MC before insertion into a kiln.
Besides collapse, the defects of honeycomb and checking can be present.
Table 5a shows a schedule for drying green, 4/4 giant chinkapin. This
schedule should be considered experimental; the final quality of lumber
produced by this schedule is not well demonstrated. In Table 5b, there
is a schedule that could be used if the material is first air-dried to the
recommended 20 percent. For chinkapin, average initial MC is reported
to be 134 percent (ovendry basis). The radial shrinkage (green to ovendry)
averages 4.6 percent; tangentially, shrinkage averages 7.4 percent.
Table 5a. Kiln schedule—Giant chinkapin 4/4 (green).

Equilibrium
1 Above 70 105 102 19.0 90 40.5 39.0

2 70 to 60 105 101 17.5 87 40.5 38.0
3 60 to 50 110 104 15.1 81 43.0 40.0
4 50 to 40 110 99 11.4 67 43.0 37.0
5 40 to 30 120 98 7.4 45 49.0 37.0
6 30 to 20 130 96 4.9 30 54.0 35.5
7 20 to final 180 135 3.8 30 82.0 57.0
Table 5b. Kiln schedule—Giant chinkapin 4/4 (air-dried to <20%).

Equilibrium
Temperature ∞F moisture Relative Temperature ∞C
Dry- Wet- content humidity Dry- Wet-
Step Time (h) bulb bulb (%) (%) bulb bulb
1 12 110 104 15.1 81 43.0 40.0

2 12 110 99 11.4 67 43.0 37.0
3 12 120 98 7.4 45 49.0 37.0
4 12 130 96 4.9 30 54.0 35.5
5 As needed 180 135 3.8 30 82.0 57.0
to final MC
Specific procedure for partially air-dried stock (less than 20 % MC)—Set the dry-bulb temperature at the value pre-
scribed in Step 1. With the vents closed, add steam spray only as needed to keep the wet-bulb depression from
exceeding 10 ∞F, but do not allow the depression to become less than 5 ∞F, or moisture will condense on the wood.
After the prescribed dry-bulb temperature has been reached, run a minimum of 12 hours on each of the first four wet-
bulb depression steps of the schedule (still allow for the minimum 5 ∞F wet-bulb depression limit during changes). Then
adjust to the conditions prescribed for the moisture content of the controlling samples at the last step.
57
Machining
Giant chinkapin is a good wood for machining. It does not plane as
well as the oaks or Pacific madrone, but there are fewer defects than
with similar runs of walnut, red alder, and maple. Shaping and turning
operations must be carefully controlled. Chinkapin sands well, with a
minimum of both scratching and fuzzing.
Adhesives
Giant chinkapin bonds well with a wide variety of adhesives if condi-
tions are moderately well controlled. The joint strength of chinkapin is
considered to be excellent.
Finishing
From the limited literature on this subject, it appears that there are
no apparent difficulties in staining or coating this wood. Its light brown
color is very pleasant when clear-coated. Although filling is recommended
for high-gloss finishes, the open grain characteristics of this wood allow
for dramatic highlighting with glazes or lightly pigmented stains.
Durability
Information on the natural resistance of chinkapin to wood-decay
organisms is very limited and conflicting. The heartwood is classified as
nondurable in one source and “somewhat more than moderately du-
rable” in another. Infestation with powder-post beetle (Ptilinus basalis)
often occurs if the wood is stored improperly, and sap staining can
affect its appearance.
Uses
Chinkapin is used for furniture, veneer, paneling, doors, and fire-
wood.
Related Literature
HARRINGTON, T.B., J.C. TAPPEINER, II, and R. WARBINGTON. 1992.

Predicting crown sizes and diameter distributions of tanoak, Pacific
madrone, and giant chinkapin sprout clumps. Western Journal of
Applied Forestry 7:103-108.
tems for the Major Forest Types of the United States. R.M. Burns,
58
tech compil., USDA Forest Service, Washington, DC. Agriculture Handbook
445.
McKEE, A. 1990. Giant chinkapin. P. 234-239 in Silvics of North America.
Forest Service, Washington D.C. Agriculture Handbook 654.
PRESTEMON, D.R., F.E. DICKINSON, and W.A. DOST. 1965. Chinkapin
log grades and lumber yield. California Agriculture Experiment Sta-
tion, Berkeley, California. California Forestry and Forest Products No.
42.
ROY, D.F. 1955. Hardwood sprout measurements in northwestern Cali-
fornia. USDA Forest Service, California Forest and Range Experiment
Station, Berkeley, California. Forest Research Note 95. 6 p.
59
OREGON ASH
FRAXINUS LATIFOLIA
Oregon ash, a member of the olive family (Oleaceae), is one of 16

species of ash in the United States. It is the only ash native to the Pacific
Northwest.

Mature Oregon ash attain heights of 60 to 80 ft and DBH of 16 to
30 in. Ash may live at least 250 years. In forest stands, Oregon ash
develop short, narrow crowns with small branches on long, straight
Oregon Ash boles. Open-grown trees have broad, rounded crowns with large
Range limbs on short boles. The root system of ash is moderately shallow,
wide-spreading, and densely fibrous.
BRITISH
COLUMBIA
Geographic Range
Oregon ash is native from northern Washington (lat 48∞N) to south-
WA ern California (lat 35∞N) on the west side of the Cascade Range and the
Sierra Nevada. It is most common in valleys and along rivers.
OR
ID
Timber Inventory
The total inventory of Oregon ash (about 184 MMCF) is scattered
across all subregions of Oregon and Washington (Appendix 1, Table 1).
It is most abundant in the Columbia Basin and tributary streams and
CA
valleys in northwestern Oregon and southwestern Washington. The availability
of Oregon ash for timber harvest is significantly restricted by regula-
tions governing forestry practices in riparian areas and wetlands.
60

Oregon ash is intermediate in tolerance, and commonly occurs in
intermediate or codominant crown positions. Young seedlings will grow
in fairly dense shade. Overtopped trees of any age can respond to
release with openings in the overstory.
Ecological Role
Oregon ash is a long-term dominant in riparian areas along slow
streams and in other poorly drained areas subject to seasonal flooding.
It may replace cottonwood on heavy soils and on swampy ground with
poor drainage during the growing season. Oregon ash seedlings will
also colonize wet areas in grasslands and abandoned fields.
Tree species commonly associated with Oregon ash are black cot-
tonwood, red alder, white alder, bigleaf maple, Oregon white oak, Cali-
fornia-laurel, California sycamore, and various willows. Douglas-fir, grand
fir, and ponderosa pine are associated with Oregon ash on the drier
margin of typical ash sites. Understory vegetation is sparse under dense
stands; sedges often dominate under ash. Common shrub associates
include snowberry, hawthorn, serviceberry, mockorange, crabapple, and
himalaya berry.

Oregon ash is particularly suited to heavy soils and poorly drained
areas, which are often too wet for any other tree, including cotton-
wood. Ash may grow very well outside of such areas, although manage-
ment may be required to maintain it. Growth and stem form are often
quite poor on poorer sites; the suitability of sites should be evaluated
before effort is invested in management of Oregon ash.
The capability of a site for growing Oregon ash should be evaluated
by examining growth and form of older trees. Good growth potential is
indicated by the following:
• Top height on mature trees of at least 60 ft
• Sustained height growth of 1 to 2 ft per year at ages 5 to 20 years
Climate
Oregon ash thrives in a mild, humid climate characterized by rela-
tively cool, humid summers and wet, mild winters. In this climate, annual
61
precipitation ranges from 20 to 118 in., with very little rain in July and
August; mean annual temperatures range from 46 to 54 ∞F.
The bottomlands where Oregon ash is most common provide a
relatively cool, moist environment, even in hot interior valleys. Oregon
ash has a fairly high tolerance to summer drought; however, it gener-
ally ceases growth and drops its leaves when conditions become hot
and dry. Oregon ash that establish on drier sites often persist in a
stunted, crooked form.
Elevation
Oregon ash usually grows at elevations from sea level to 3000 ft,
although it may be found as high as 5000 ft.
Soils
Oregon ash typically grows on deep, poorly drained clays or silty
clay loams that are rich in humus. It will also grow on sandy, rocky, and
gravelly soils in riparian areas or areas with seasonal flooding. Oregon
ash also grows on upland forest soils adjacent to more typical ash habi-
tat.

Oregon ash begins producing seed at about 30 years of age. Male
and female flowers grow on separate plants. The greenish-white flowers
(male and female) are borne in dense panicles, which appear at the
base of new foliage in April or May. The fruits are oblong to elliptical
samaras 1 to 2 in. long. They ripen in August or September, turning
from green to light brown.
Seed
The winged seeds (10,000 to 14,000/lb) are dispersed by wind dur-
ing September and October. Ash seeds can remain viable for more than
one year; they usually require moist, cold stratification in order to ger-
minate. Germination rates are medium to high.

Open-grown trees produce good seed crops almost every year. In
forest stands, heavy crops of seed are produced every 3 to 5 years. Wet
or moist soils high in organic matter provide the best conditions for
germination and establishment of Oregon ash. Although many seeds
are transported by flood waters, few seedlings establish from seeds
deposited on sandy or gravelly stream beds. Seedlings commonly estab-
lish in the understory of existing riparian forests after floods that de-
posit silt. Ash seedlings also establish in wet grasslands or fields, par-
ticularly after disturbance such as plowing (in the absence of fire or
grazing).
62
Oregon ash sprouts vigorously from the root collar after cutting.
Stumps should be cut low to the ground in order to produce well-
formed sprouts of good quality.

The performance of Oregon ash has been poor to fair in limited
outplanting trials. There are no known examples of operational forest
plantations. Oregon ash is widely planted and does very well in orna-
mental applications, which indicates its potential for good performance
in managed plantations.

origin. Both mechanical and chemical treatment options may be limited
on typical, wet Oregon ash sites. Although Oregon ash would probably
benefit from control of competing vegetation on a site, established
seedlings seem to grow well amidst substantial competition in old fields.
Stand Management
Oregon ash is not generally managed for timber production. Young
ash trees grow fairly rapidly, but are commonly crooked or forked. Young
stands are typically dense; early thinning to allocate stand growth to
well-formed trees can improve stand quality. Older ash trees also re-
spond to thinning; the diameter growth of crop trees doubled after
crown-thinning in a dense, 45-year-old stand.
The shade tolerance of young Oregon ash trees and the ability of
overtopped trees to respond to openings provide flexibility for manage-
ment of ash in mixed species or age classes. Oregon ash are often
mixed with cottonwood and bigleaf maple in bottomland forests, where
flood waters maintain a variable disturbance regime. Management of
these areas must integrate the range of growth rates, shade tolerance,
and regeneration needs among associated species.
Growth and Yield

Growth of Oregon ash is moderately rapid for the first 60 years (1
to 2 ft per year). Thereafter, height growth is negligible and top height
is relatively short (60 to 80 ft), although good diameter growth may
continue. Early growth of stump sprouts is quite rapid (3 to 4 ft per
year). There are few estimates of volume growth or yield for Oregon
ash. Measurements in a pure stand of 45-year-old Oregon ash showed
63
an average height of 70 ft and basal area of 140 to 160 ft 2 per acre,
which yield approximate volumes of 3200 to 3800 ft 3 per acre. Annual
growth was about 70 ft 3 per acre in both lightly thinned and unthinned
stands.

Ribbonlike forests of Oregon ash along streams and sloughs provide
an important forest habitat in valleys that are otherwise cultivated farm
or pasture. Oregon ash provide food and habitat for beaver and nutria,
which also significantly damage the trees. Deer and elk also browse
Oregon ash seedlings and stump sprouts.

Weevils (Thysanocnemis spp.) destroy significant amounts of seed
(up to 60 percent). Various other insects feed on twigs and foliage and
may be pests in ornamental plantings. A variety of fungi cause leaf spot
and powdery mildew. True mistletoe grows on Oregon ash. A heart rot
causes extensive defect in older trees, and hollow “stovepipe” ash are
prevalent in some stands.
Genetics
Some of the stand-to-stand variation in the form of Oregon ash may
have a genetic basis. There are no recognized varieties of Oregon ash,
although it may hybridize with velvet ash (F. velutina) in the southern
Sierra Nevada.

There are no equations or tables for calculating tree volume. Log
grades have not been developed. Premium prices may be paid for logs
12 in. and larger. Harvesting of Oregon ash may often be subject to
restrictions for protection of riparian zones or wetlands.
Product Recovery
Sawlogs usually have a minimum small-end diameter of 6 in. The
relatively small volume of sawlogs that are harvested are fully utilized,
demand for lumber is relatively high. Lumber grade recovery has not
been studied.
64
Wood Properties
Characteristics
The wood of Oregon ash is moderately hard and heavy, with distinct
growth rings. The sapwood is nearly white and is wide. The heartwood
is a yellowish-brown, with a slight greyish cast. The wood is somewhat
lustrous. It has no characteristic odor or taste. Oregon ash wood is ring
porous; the earlywood vessels are large, forming a band that is 2 to 4
pores in width and distinctly visible to the naked eye. The transition to
latewood is abrupt. Latewood pores are small, barely visible to the na-
ked eye. The variability between earlywood and latewood is especially
apparent on flat-sawn surfaces, where the denser latewood markedly
contrasts with the porous, grainy earlywood. Rays are not easy to distin-
guish with the naked eye.
Weight
Oregon ash weighs about 48 lb/ft 3 when green and 38 lb/ft 3 at 12
percent moisture content (MC). The average specific gravity is 0.50
Because Oregon ash is moderately hard and heavy, it is rated inter-
mediate in bending strength and stiffness. Its impact resistance, how-
ever, is rated as high. Its hardness and compressive strength is good;
thus it is well suited for most furniture, paneling, or flooring applica-
tions. Nail and screw splitting are less problematic than with other,
more dense woods such as oak. The nail-holding ability of Oregon ash is
good. Appendix 1, Table 3 provides information on clear strength val-
ues for Oregon ash, as well as comparative values of other species.

Under moderately controlled conditions, Oregon ash dries rapidly
and with minimal degrade. Drying defects are most often associated
with wetwood, or are caused by uncontrolled air-drying, which results
in surface- and end-checking. Sapstains and sticker stains can also de-
grade the wood. Green MC of Oregon ash wood averages 49 percent.
Shrinkage values for Oregon ash dried from green to ovendry are 4.1
percent radially and 8.1 percent tangentially. These values are similar to
other ash species. Table 6 provides an appropriate dry-kiln schedule. For
thicker stock, contact the Forest Product Department at OSU for addi-
tional schedules.
65
Table 6. Kiln schedule—Oregon ash 4/4, 5/4, 6/4.
Equilibrium
1 Above 35 130 123 14.3 81 54.5 50.5

2 35 to 30 130 120 12.2 74 54.5 49.0
3 30 to 25 140 125 9.6 64 60.0 51.5
4 25 to 20 150 125 6.8 49 65.5 51.5
5 20 to 15 160 120 4.5 31 71.0 49.0
6 15 to final 180 130 3.5 26 82.0 54.5
Machining
Oregon ash machines without much difficulty. It planes and shapes
favorably; ash is comparable to bigleaf maple in shaping qualities, and
slightly below the oaks in planing qualities. It turns well and, when
holes are bored in ash, the holes are smooth with minimal size varia-
tion. Fuzzing from sanding is rare, although surface scratching can be
apparent, especially on the latewood parts of flat-sawn boards. The best
results in planing are obtained with tooling that has a 25∞ hook angle.
Tool wear is moderate.
Adhesives
Oregon ash bonds satisfactorily and there are no unusual problems
when conditions are well controlled. Because of its light color, the glue
line can be visible on sapwood if darker resins are used.
Finishing
The natural luster of Oregon ash is enhanced with clear finishes or
transparent dyes. The earlywood vessels may require filling to minimize
surface texture. Heavily pigmented stains tend to darken the more po-
rous earlywood without penetrating the denser latewood. Color varia-
tion between sapwood and heartwood can present problems if a uni-
form color is desired. Surface scratching in the dense latewood can
sometimes be a problem.
Durability
Oregon ash is a nondurable species that is susceptible to wood de-
cay. Untreated posts of this wood average only 6 years of service before
failing. Heartwood of ash species is considered easily penetrated with
66
preservative solutions. The wood is subject to attack by powder post
beetle (Ptilinus basalis) and is moderately susceptible to sapstain, mold,
and iron stain.
Uses
Oregon ash is used for furniture, cabinets, paneling, veneer, tool
handles, flooring, millwork, pallets, crates, boxes, and firewood.
Related Literature
FRENKEL, R.E., and E.F. HEINITZ. 1987. Composition and structure of

Oregon ash (Fraxinus latifolia) forest in William L. Finley National
Wildlife Refuge, Oregon. Northwest Science 61:203-212.
HIBBS, D.E., and G.R. AHRENS. Unpublished data. Forest Science De-
partment, Oregon State University, Corvallis.
OWSTON, P.W. 1990. Oregon ash. P. 339-343 in Silvics of North America.
67
OREGON WHITE OAK
QUERCUS GARRYANA
GARRY OAK
OREGON OAK
BREWER OAK
Oregon white oak, a member of the beech family (Fagaceae), is one

of only four deciduous oaks native to the West Coast. The massive,
branching trunks and broad crowns of old white oaks are characteristic
features of valley woodlands in the Pacific Northwest.

Oregon White Mature Oregon white oaks are 50 to 90 ft tall (120 ft maximum)
Oak Range and 24 to 40 in. in DBH (97 in. maximum). Oregon white oaks may
live 500 years. In forest stands on good sites, Oregon white oaks
BRITISH develop narrow crowns with small branches on straight, clear stems.
COLUMBIA
In more open stands and on poor sites, boles are typically short and
crooked; shrubby stands of stunted trees are common. Open-grown
trees develop very broad, rounded crowns (crown width may equal
total height) with massive, crooked branches on short, massive boles.
WA
The root system of this species is composed of a deep taproot and well-
developed laterals.
OR
Geographic Range
Oregon white oak has a wide latitudinal range from Vancouver Is-
CA land (lat 49∞N) to southern California (lat 34∞N), although it takes on a
shrub form toward the southern end of this range. It is well distributed
throughout the valleys west of the Cascade Range and Sierra Nevada on
inland slopes of the Coast ranges.
68
Timber Inventory
A substantial inventory of Oregon white oak (450 MMCF) is distrib-
uted throughout western Oregon and northwestern California. A much
smaller volume (13 MMCF) occurs in western Washington (Appendix 1,
Table 1).

Oregon white oak is generally intolerant, although this depends on
the environment and associated vegetation. Sparse development of branches
in closed stands indicates intolerance to shade. Although it can repro-
duce in its own shade, Oregon white oak will die after overtopping by
Douglas-fir.
Ecological Role
Oregon white oak is a persistent climax or sub-climax species on dry
sites or under regimes of periodic fire. Large oaks have thick bark and
are resistant to fire. Smaller trees are generally killed or badly injured by
fire. Oregon white oak is an early successional species on better sites,
where it is replaced by Douglas-fir and bigleaf maple in the absence of
fire. Historically, periodic fires were a major factor maintaining Oregon
white oak woodlands. After a century of fire exclusion, many acres have
progressed from open Oregon white oak, to closed white oak, to Dou-
glas-fir. Fire prevention is probably causing continued decline in the
extent of Oregon white oak type forests. The white oak type will con-
tinue to diminish without periodic fire.
Many distinct Oregon white oak associations are recognized. Com-
mon associate trees of Oregon white oak are Douglas-fir, grand fir,
ponderosa pine, bigleaf maple, Pacific madrone, California black oak,
Oregon ash, and cherry. Common shrubs include hazel, hawthorn, snowberry,
serviceberry, poison-oak, wild rose, and oceanspray. Herbaceous associ-
ates include many different grasses, western swordfern, western bracken,
wild strawberry, bedstraw, and sweetroot. A great variety of other plant
species grow with Oregon white oak in other forest types.

Oregon white oak is particularly suited to exposed, droughty sites at
the margins of more productive forest land. It is also well suited for
69
areas near rivers that are very wet in winter but droughty in the sum-
mer. Oregon white oak will grow well on better sites, but requires man-
agement to persist among more competitive Douglas-fir and maple.
The capability of a site for growing Oregon white oak should be
evaluated by examining growth and form of older trees. Good potential
for growth of this species is indicated by the following site characteris-
tics:
• Top height on mature trees of at least 60 ft
• Sustained height growth of 1 to 2 ft per year for trees 10 to 30
years old
Climate
Oregon white oak grows across a diverse range of climates, most of
which have moderate to extreme summer drought and annual precipita-
tion of 10 to 100 in. Within its range, mean daily temperatures may
vary from lows of 13 to 50 ∞F in January to highs of 60 to 84 ∞F in July.
Oregon white oaks are well adapted to hot, dry conditions. With
adequate moisture early in the season, relatively large trees may de-
velop on sites where severe summer drought limits other species. Exten-
sive stands of small, shrubby white oak, often mixed with Pacific mad-
rone, grow on sites that are often too dry to support any other tree
species.
Of the western hardwoods, Oregon white oak is one of the most
resistant to damage from ice and snow. Twigs and buds have moderate
resistance to cold injury. Healthy Oregon white oak are not prone to
windthrow or breakage.
Elevation
Oregon white oak is usually found at lower elevations in the interior
valleys. It grows from sea level to 3800 ft in the north and at elevations
of up to 7500 ft at the southern end of its range.
Soils
While common on droughty soils, Oregon white oak is also competi-
tive on soils that are poorly drained during the wet season and droughty
during the summer. This commonly occurs on heavy clays, coarse-tex-
tured flood plains, and river terraces in the interior valleys. Oregon ash
is also common under these conditions.

The age at which Oregon white oak first produces seed has not been
determined. Oregon white oak flowers appear with new foliage in spring.
Oregon white oak usually flowers later than common associates, in March
in the south, and June in the north. Separate male and female flowers
70
grow on the same tree. Male flowers (greenish-yellow catkins) emerge
from existing buds and at the base of new shoots, expanding to a size
of 1.2 to 3.9 in. when fully developed. Small, red female flowers appear
in the axils of new leaves. The acorns ripen from August to November
during the first season after flowering.
Seed
Oregon white oak has large acorns, which are about 1.2 in. long
and half as wide, and average 85 seeds/lb. Acorns should be collected
from September to November. They must be kept cool and moist until
germination. Although germination usually occurs in spring, seeds will
germinate soon after dispersal under warm, moist conditions; they also
germinate prematurely in cool, moist storage. As with other Oregon
white oaks, sowing seeds in the fall may be best. Limited tests indicate
rates of viability that are greater than 75 percent. Seeds remain viable
for only one season.

Natural regeneration from seed is often quite good where there has
been soil disturbance, particularly in the absence of fire or grazing.
Seedlings rapidly develop a deep taproot, which may account for their
ability to establish in grass and in droughty soils. The shoot of natural
seedlings often remains small and shrubby for many years, perhaps to
accommodate development of deep roots. This is followed by a sapling
stage with relatively rapid growth. Regeneration from seed is greatly
improved when seeds are protected from rodents and other predators.

Oregon white oak sprouts vigorously after cutting or fire. Sprouts
arise from dormant buds at the root collar and along the trunk. Both
the vigor and the abundance of sprouts increase as the size of the
parent tree increases. Stumps should be cut low to the ground in order
to produce well-formed sprouts of good quality.

Nursery culture of Oregon white oak is relatively easy. Experience
with other white oaks indicates that with vigorous nursery stock, Or-
egon white oak has good potential for management in plantations.
Performance of Oregon white oak has been poor to fair in limited outplanting
trials, however. There are no known examples of operational forest plan-
tations.

Cultivated seedlings of Oregon white oak grow rapidly and do not
display the prolonged shrub stage observed for many wild seedlings.
71
This suggests that site preparation and control of competition from
grasses and shrubs could greatly improve growth of Oregon white oak
seedlings in the field.
Stand Management
Oregon white oak has not been extensively managed for timber
production. It shows good potential for management in closed, even-
aged stands, as indicated by the relatively good stand growth and high
stem quality in closed stands established after the exclusion of fire.
Thinning in dense, sapling-sized stands can increase diameter growth.
Competitive associates such as Douglas-fir and bigleaf maple must
be controlled to maintain Oregon white oak on better sites.
Growth and Yield

Growth of Oregon white oak is generally slow. Height growth is
usually less than 1 ft per year and diameter growth is often 15 to 20
rings/in. Faster growth, particularly in diameter, is possible (3 to 10
rings/in.). Stump sprouts may grow as much as 3 ft per year during the
first 3 years. Oregon white oak stands may achieve basal area of up to
265 ft 2 per acre and volume as high as 4500 ft 3 per acre.

Oregon white oak trees and stands are very important for wildlife.
Oak woodlands and forests provide food and habitat for many species.
Two of note are the acorn woodpecker and Merriam’s wild turkey.
Diversity of bird species is often higher in oak forests than in adjacent
conifer forests. Both acorns and foliage provide high-protein food for
many animals.

Filbertworm (Melissopus latiferreanus) and filbert weevils (Curculio
occidentalis) attack acorns of Oregon white oak. Hundreds of other
insect species live on Oregon white oaks, although few cause signifi-
cant damage. The most damaging insect is the western oak looper
(Lambdina fiscellaria somniaria), which can defoliate trees over large
areas; tent caterpillars also have a preference for oak. Gall wasps are
common on Oregon white oak. Noticeable damage is often inflicted on
oak by Bassettia ligni, which girdles and kills branches.
Numerous pathogens attack Oregon white oak. The hairy mistletoe
is widespread. Shoestring root rot, Armillaria ostoyae (A. mellea), and
white pocket root and butt rot (Polyporus dryophilus) cause significant
damage. One episode of anthracnose disease (Gnomonia quericina) ap-
peared to cause significant damage in Washington in 1968.
72
Genetics
Oregon white oak hybridizes with four other oaks in California. There
is relatively little genetic variation within the species, despite its wide
latitudinal range.

volume. Log or tree grades are not used for Oregon white oak; how-
ever, recent studies have shown that there are differences in the value
of lumber that can be recovered from each of the log grades. Log
grades developed for eastern hardwoods may be a useful marketing tool
in the future. Oregon white oak logs check easily during storage and
should be end-coated to prevent splits.
Product Recovery
Sawlogs usually have a minimum small-end diameter of 8 in., smaller
logs are chipped for pulp. Rot, checks, and cross-grain have been prob-
lems in the lumber.
There is increasing interest in using Oregon white oak for cooperage
for wine barrels, flooring, and chairs. It has been used for pulp and is
frequently used for firewood.
Wood Properties
Characteristics
Oregon white oak is a hard, heavy wood that has distinct growth
rings and very prominent rays. The sapwood is whitish to light brown;
the heartwood is a pale, yellowish, grey-brown, often with a slight greenish
cast. The dry wood is has no characteristic odor or taste. Oregon white
oak is ring porous; the earlywood pores are large and distinct, forming a
conspicuous band with each growth ring. The latewood pores are small
and numerous, and require a hand lens to view. Rays are of two types,
broad and narrow. The broad rays are readily visible to the naked eye
and are separated by several to many narrow rays. When oaks are quartersawn,
these rays appear as a pronounced fleck. The earlywood pores are plugged
with a membranous growth known as tyloses, which makes the wood
73
impenetrable to fluids. For commercial purposes, Oregon white oak is
classed with the other white oaks in USDA Forest Service nomenclature.
Weight
Oregon white oak weighs about 69 lb/ft 3 when green and 50 lb/ft 3
at 12 percent moisture content (MC). The average specific gravity is
0.72 (green) or 0.75 (ovendry).
The wood of Oregon white oak has exceptional strength properties
and is noted for its hardness, toughness, resiliency, and resistance to
abrasion. It holds nails well, but, because of its density and hardness,
will split without preboring. See Appendix 1, Table 3 for average me-
chanical properties for small clear specimens.

Oregon white oak requires special care and attention to detail dur-
ing an extended kiln schedule to properly reduce MC to a level suitable
for interior products such as flooring, furniture, or millwork. Drying
defects can cause serious downgrade. End- and surface-checking result
from uncontrolled or overly rapid drying; honeycomb, collapse, and
ring failure occur because of wetwood. Iron stains form when tannins
contact certain metals, and grey sapwood staining will result if there is
poor air circulation. Green MC of the wood is generally 67 to 72 per-
cent (ovendry). Shrinkage values for green to ovendry (based on the
original green size) average 4.2 percent radially and 9.0 percent tan-
gentially. It is suggested that all the upper grades be air-dried to 20
percent MC and then kiln-dried according to a time schedule (See Table
7 for the appropriate kiln schedule).
Table 7. Kiln schedule—Oregon white oak 4/4, 5/4, 6/4, 8/4.

Equilibrium
1 Above 35 110 107 19.1 90 43.5 41.5

2 35 to 30 110 106 17.6 87 43.5 41.0
3 30 to 25 120 114 15.5 83 49.0 45.0
4 25 to 20 130 120 12.2 74 54.5 49.0
5 20 to 15 140 115 6.8 46 60.0 46.0
6 15 to final 160 110 3.4 21 71.0 43.5
74
Machining
Species in the white oak group, including Oregon white oak, gener-
ally machine well. They plane well (87 percent defect-free pieces), turn
well (85 percent defect-free) and yield accurately sized, smooth-sided
holes when bored and mortised. White oaks also bend exceptionally
well after steaming. On white oak, sanding produces a smooth, rela-
tively scratch-free surface with little or no fuzzing. The hardness, tannin
content, and density of these woods cause considerable tool dulling and
sandpaper wear. It is recommended that saws and other tools have
hook angles of 15 to 20∞ and sharpness angles of 55∞ for optimum
performance on white oaks. Care should be taken not to overfeed this
wood or attempt to remove too much stock at once because machine
burn or surface roughness may result.
Adhesives
Oregon white oak bonds satisfactorily, and there are no unusual
problems when gluing conditions are well controlled. Careful curing/
drying of glue joints is required to prevent sunken gluelines from subse-
quent machining.
Finishing
All white oaks finish well, although it may be necessary to fill the
grain. White oaks color best with dyes or transparent stains, especially if
dramatic color changes are sought. Heavily pigmented stains can also
be used if care is taken to remove excess pigment from the wood.
Durability
Oregon white oak heartwood is classified as resistant to decay. In
tests conducted by staff of the OSU Forest Products Department, un-
treated fence posts lasted an average of 18 years before failure. The
sapwood has no decay resistance and will deteriorate rapidly. Iron stain
and oxidative stain sometimes occur on Oregon white oak.
Uses
Oregon white oak is used for furniture, flooring, railroad ties, tight
cooperage, turnings, veneer (sliced), millwork, fence posts, mine tim-
bers, handles, boxes, crates, pallets, caskets, pulp chips, and fuelwood.
75
Related Literature
COBLENTZ, B.E. 1980. Production of Oregon white oak Quercus garryana

acorns in the Willamette Valley, Oregon utilization by deer. Wildlife
Society, Bulletin 8:348-350.
EVANS, D. 1972. Alternate generations of gall cynipids (Hymenop-
tera:Cynipidae) on Garry oak. Canadian Entomologist 104:1805-1818.
GUMTOW-FARRIOR, D.L., and C.M. GUMTOW-FARRIOR. 1992. Manag-
ing Oregon white oak communities for wildlife in Oregon’s Willamette
Valley: a problem analysis. Unpublished report prepared for the Or-
egon Department of Fish and Wildlife Nongame Program.
HIBBS, D.E., and B.J. YODER. 1993. Development of Oregon white oak
seedlings. Northwest Science 67:30-36.
tems for the Major Forest Types of the United States. R. Burns, tech
compil. USDA Forest Service, Washington, DC. Agriculture Hand-
book 445.
PLUMB, T.R., and P.M. McDONALD. 1981. Oak management in Califor-
nia. USDA Forest Service, Pacific Southwest Forest and Range Experi-
ment Station, Berkeley, California. General Technical Report PSW-
54. 12 p.
RIEGEL, G.M., B.G. SMITH, and J.F. FRANKLIN. 1992. Foothill oak wood-
lands of the interior valleys of southwestern Oregon. Northwest Sci-
ence 66:66-76.
STEIN, W.I. 1990. Oregon white oak. P. 650-660 in Silvics of North
America. Volume 2, Hardwoods. R.M. Burns and B.H. Honkala, USDA
76
PACIFIC MADRONE
ARBUTUS MENZIESII
MADRONE
MADRO~NA
MADRO~NO
Pacific madrone is one of the largest of about 14 species of Arbutus

in the world, and one of the two Arbutus species in North America.
Pacific madrone is a broadleaved evergreen tree and a member of the
heath family (Ericaceae). It is distinguished by its smooth trunk, or-
ange-red deciduous bark, white flowers, and red berries.

Pacific madrones attain heights of 80 to 125 ft and diameters of
24 to 48 in. The largest trees may be as much as 400 years old; ages
Pacific Madrone of 200 to 250 years have been counted. Pacific madrone can develop
Range a clear, straight bole under good conditions in forest stands, particu-
larly in canyons and dense stands. Open-grown individuals and trees
BRITISH
COLUMBIA
growing on lower quality sites often have multiple stems, which origi-
nate from sprouts or root burls that often are J-shaped and forked. The
tree may become shrubby on poor sites. Pacific madrone generally de-
velops a deep and spreading system of lateral roots, often in association
WA with large root burls. Seedlings have a tap root.
OR
ID
Geographic Range
CA Pacific madrone is found from San Diego (lat 33∞N) to eastern Vancouver
Island (lat 51∞N). In Oregon and Washington, it is restricted to the
Coast Range and the west slopes of the Cascade ranges. In California, it
is also found in the Coast Range, throughout much of the Klamath
Mountains, and in some areas west of the Sierra Nevada.
77
Timber Inventory
Pacific madrone is the most abundant hardwood in the Siskiyou
Mountains and interior coast ranges of the Southwest subregion of
Oregon. This is the only subregion of Oregon that has a substantial
inventory of Pacific madrone timber (Appendix 1, Table 1). Much of
the Pacific madrone in Oregon is on federal lands, although volume
estimates are not readily available. Pacific madrone is the second-most
abundant hardwood in northern California. In Washington, it is com-
mon in the Puget Sound and Olympic subregions.

Pacific madrone most commonly occurs as a codominant or inter-
mediate tree in a canopy of mixed-hardwood species that often have
some overstory of conifers. Pacific madrone is intermediate in toler-
ance. Tolerance appears to be lower for older trees and for trees at the
northern end of the range. Seedlings establish best in partial shade,
and young trees can survive in fairly dense shade. Top light is required
for good growth; older trees may require top light to survive. Pacific
madrone will grow toward openings, leaning as much as 15 to 20
degrees.
Ecological Role
Pacific madrone can be subclimax or climax in successional status; a
substantial component of madrone is often maintained by periodic fires
in the southern and central parts of its range. Although the thin-barked
stems are easily killed by fire, Pacific madrone often dominates post-fire
vegetation via vigorous regeneration of sprouts. It can also persist as a
component of the mixed Douglas-fir/tanoak/Pacific madrone forest type.
In the heart of its range, Pacific madrone is a major component of a
widespread mixed-evergreen forest, which is characterized by an over-
story of Douglas-fir and a secondary canopy of mixed hardwoods. Un-
derstory vegetation is often sparse under mature stands containing Pa-
cific madrone. Pacific madrone is a common associate in a variety of
other major cover types in the region.
Common tree species associated with Pacific madrone include Dou-
glas-fir, ponderosa pine, sugar pine, white fir, western hemlock, tanoak,
Oregon white oak, California black oak, giant chinkapin, bigleaf maple,
bitter cherry, and California-laurel. Small trees commonly associated
with Pacific madrone include vine maple, black hawthorn, red osier
dogwood, willow, hazel, and red elderberry. Numerous shrub associ-
78
ates include manzanitas, Oregon-grape, ceanothus, salal, oceanspray,
poison-oak, gooseberry, wood rose, snowberry, huckleberry, and thimbleberry.

Pacific madrone is particularly suited for warm, dry sites in the Northwest,
especially on south and west aspects. Many of these sites may be mar-
ginal for production of other tree species, particularly in the absence of
intensive vegetation management. On such sites, Pacific madrone’s abil-
ity to maintain forest cover and produce usable wood becomes an im-
portant asset, one that may be improved with management. Relatively
good growth and stem quality can be produced on better sites, al-
though species such as Douglas-fir and tanoak are also more competi-
tive on these sites. There are no established guides or site-index curves
for estimating the productivity of a site for Pacific madrone. A site with
good potential for growth of Pacific madrone is indicated by site trees
with the following characteristics:
• Top height on mature trees of 80 to 100 ft
• Rapid juvenile height growth of 1 to 3 ft per year
• Sustained height growth from age 15 to 30 of 1 to 2 ft per year
Climate
Pacific madrone prefers a climate characterized by mild, wet winters
and dry, cool summers. Within its range, annual precipitation varies
from 25 to 118 in. and average temperatures range from 36 ∞F in Janu-
ary to 77 ∞F in July.
Pacific madrone tolerates warm, dry conditions better than most
tree species in the Northwest. It is one of the most drought-tolerant
trees in the region and it has superior ability to extract water from soil
or rock. Its roots can penetrate up to 12 ft in fractured bedrock, giving
it access to substantial moisture unavailable to shallow-rooted species.
Established and resprouting Pacific madrone are thus able to maintain
relatively good growth on shallow, rocky soils where it may be difficult
for seedlings of any species to establish and grow.
Pacific madrone is relatively sensitive to cold and snow. Its broad,
evergreen leaves and brittle branches are vulnerable to breakage from
heavy wet snow. Foliar damage and die-back are commonly observed
after severe freezing or unseasonable frost. At the northern end of its
range, Pacific madrone is one of the least frost-resistant tree species.
Pacific madrones are relatively windfirm because of their deep, spreading
root systems.
Elevation
At the southern end of its range, Pacific madrone is found from
2000 to 4260 ft in elevation. In the north, it ranges from sea level to
3000 ft.
79
Soils
Towards the southern and middle part of its range, Pacific madrone
grows on soils derived from a wide variety of parent materials. In the
north, it is usually found on soils derived from glacial sands and gravels or
hard glacial till. It is often found on rocky soils and on soils with low
moisture retention. Pacific madrone is generally restricted to soils with
good internal drainage; it will not tolerate poor soil drainage or flooding.

Pacific madrone produces seed as early as 3 to 5 years of age. Trees
begin flowering in early spring, from mid-March to May, depending on
the elevation. The blossoms are dense, drooping clusters (terminal panicles)
of small, white, urn-shaped flowers. The fruit is a berry (0.3 to 0.5 in.),
which ripens in the fall, turning from yellow-green to bright red or
reddish-orange.
Seed
Berries number from 630 to 1130/lb and contain an average of
about 20 seeds per berry. Seeds are small, numbering from 197,000 to
320,000/lb. The berries are fleshy and relatively heavy; the seed are
thus dispersed by gravity or by animals. The berries are eaten by many
birds and mammals.
To obtain seeds, berries should be collected soon after they ripen in
the fall. The following methods have been suggested for treatment of
berries and seeds (Jane Smith, USDA Forest Service, PNW Station, Corvallis,
Oregon). Berries can be dried at room temperature and stored at 34 ∞F
(4 ∞C) for at least 2 years. Seeds should be separated from the pulp of
fresh or dried berries. To extract seeds from dried berries, berries can be
soaked in water (overnight) and blended in cold water in a blender at
low speed for 3 to 10 minutes. Moist stratification for at least 4 to 6
weeks at 1 to 2 ∞C may improve germination.

In the northern parts of its range, Pacific madrone usually produces
seed every year. Very good crops may occur as frequently as every 2
years, while very light seed crops may occur only once in 10 years. At
the southern end of its range, good seed crops may occur as infre-
quently as once every 10 years. Seeds usually germinate in the first year
after ripening. Natural rates of survival are often very low (0 to 10
percent) after seedling emergence because of drought, fungi, or preda-
tion.
Seedlings of Pacific madrone establish naturally in disturbed soils
along roads, near uprooted trees, or in partially open forests. Bare min-
eral soil provides the best seedbed; very few seedlings establish in un-
disturbed litter. Seedlings also need partial shade to establish. Early
growth of seedlings under natural conditions is slow (2 to 4 in. per
year).
80
Most reproduction of Pacific madrone arises from sprouts after fire
or cutting. Death of the main stem stimulates profuse sprouting (up to
300 sprouts per parent), which originate from dormant buds near the
root collar. These sprouts provide reliable regeneration and have rapid
growth potential, which is due to carbohydrate reserves and soil access
provided by pre-existing roots. Sprouts may grow as much as 5 ft in
height the first year and attain an average height of 10 ft after 3 years.
To produce vigorous, high-quality sprouts, stumps should be cut low to
the ground (<8 in.), with a slight angle to the stump surface. Pacific
madrone sprouts in partial or shelterwood cuttings have relatively poor
growth and quality. Moderate to large clearings with little competitive
vegetation produce the best growth of sprouts.
Plantation Regeneration
Little effort has been made to regenerate Pacific madrone from planted
seedlings. Commercial seedling production methods have not been de-
veloped, although good quality seedlings have been produced for some
research applications. Mortality rates have been high in field transplantings
to date.

origin. Regeneration and growth may be enhanced by burning or me-
chanically removing slash that shades Pacific madrone stumps. Rapid
growth of sprout clumps makes Pacific madrone a superior competitor
in the new stand. Control of competing herbs and shrubs can greatly
improve the growth of young sprouts.
Site-preparation treatments that produce bare mineral soil while leaving
some partial shade (debris, vegetation) may be best for promoting es-
tablishment and growth of Pacific madrone seedlings.
Stand Management
The growth and quality of Pacific madrone stands may be greatly
improved through management. Diameter growth of madrone is re-
sponsive to increased growing space within or between sprout clumps.
Sprouts should be thinned after dominant stems have emerged, at 5
to 10 years. Thinning should select well-formed, dominant stems that
originate near the ground and are evenly distributed around the stump.
One early thinning is probably adequate for production of firewood,
which may be done in 15- to 20-year rotations. A second thinning
(yielding firewood) may be beneficial if sawtimber production is de-
sired.
Thinning in older existing stands can increase diameter growth on
residual trees by 2 to 5 times. Pacific madrone stands (pure or mixed)
are often quite dense, and sometimes stagnant, with little or no diam-
81
eter growth. Periodic thinning may be necessary to avoid stagnation and
maintain stand growth.
Selective harvesting or dense shelterwoods are not recommended for
management of Pacific madrone sprouts. Uneven-aged management may
be feasible over a large area, with clearing in patches larger than 0.2
acres. Thinning will be necessary within patches or sprout clumps.
Pacific madrone typically occurs as a component or patch within
mixed stands. Management of mixed stands is complex, and may re-
quire periodic treatments to maintain growth of diverse components.
Pacific madrone stump sprouts may need to be controlled or thinned to
avoid early suppression of associated conifer seedlings. Later treatments
may be needed to maintain growth of Pacific madrone, particularly on
better sites where conifer species are superior competitors.
Growth and Yield

Most natural Pacific madrone stands originate from sprouts. Dense
sprout regeneration grows rapidly under open conditions. By age 10,
the average height of sprouts may reach 15 to 22 ft and stand basal area
may reach 100 ft 2 per acre on a good site. Typical mature trees (50 to
70 years old) are 50 to 80 ft tall and 10 to 20 in. in diameter. Diameter
growth in natural stands is relatively slow, averaging 12 to 15 rings per
in.
Mature stands or patches may attain basal areas of 140 to 200 ft 2
per acre. The best stands of Pacific madrone may exceed 4000 ft 3 per
acre over several acres. Average stand volume of Pacific madrone forest
types in California is 1705 ft 3 per acre.
There are few examples of growth and yield from managed stands.
One test with 45-year-old Pacific madrone on a poor site suggested that
thinning in dense, stagnant stands can greatly increase diameter growth
(as much as 5 times) while maintaining or even increasing total annual
volume growth per acre (33 to 37 ft) after removal of up to 65 percent
of the stand basal area. Another study of Pacific madrone in mixed
hardwood stands in northern California showed annual growth rates of
85 ft 3 per acre among all species combined, after removal of 40 to 50
percent of the original stand basal area.

Pacific madrone berries are an important food for many birds and mammals.
The berries are a particularly significant component in the diet of doves and
pigeons during the fall. Deer eat the berries and also browse young shoots.
Damage caused by animals is relatively minor on Pacific madrone. Live
trees with rotten heartwood provide excellent habitat for cavity-nesting
birds. Pacific madrones in mixed-conifer forests provide a middle canopy
story, an important element in forest structural diversity.
82
Significant mortality and damage is caused by a fungus commonly
known as “madrone canker” (asexual stage, Fusicoccum aesculi; sexual
stage Botryosphaeria dothidea). The canker causes a dieback of branches
from the tip, and cankers may spread to the bole and kill the tree. The
bark of dead branches becomes blackened, somewhat resembling fire
damage. The disease reproduces from spores in the outer bark, which
are spread by insects and, possibly, rain and wind.
A basal canker, Phytophthora cactorum, also has significant impact.
The annosus root rot, Heterobasidium annosum, has potential to cause
serious damage.
Insects such as defoliators, wood borers, and bark beetles are com-
mon but cause only minor damage.
Genetics
No natural varieties or hybrids of Pacific madrone are recognized,
although there may be some horticultural cultivars.

Cruising and harvesting
Diameter at breast height and total height of Pacific madrone can be
used in tables or equations to estimate total tree volume in cubic feet and
sawlog volume. Tests of the eastern hardwood grades have found no dif-
ference in value between log grades for this species, but have found a
significant relation between log diameter and value. Stump burls offer an
additional harvesting and management option for Pacific madrone.
Product Recovery
logs are chipped for pulp. The percentage of No. 1 Common and Better
green lumber recovered from Pacific madrone logs compares favorably
with the grade recovery from eastern oaks (Appendix 1, Table 2). Pa-
cific madrone burls are highly prized and valued for their appearance,
and are used in novelty items such as tables and clocks.
Wood Properties
Characteristics
Pacific madrone is a hard, heavy wood with a fine grain and little
texture. The sapwood is white or cream-colored with a pinkish tinge;
83
the heartwood is a light reddish-brown. The wood is without any char-
acteristic odor or taste. Pacific madrone wood is diffuse porous; the
pores are nearly uniform, numerous, and minute. With a hand lens, the
growth rings are barely visible. The rays range from barely visible to
readily visible.
Weight
Pacific madrone weighs about 60 lb/ft 3 when green and 45 lb/ft 3 at
12 percent MC. The average specific gravity is 0.58 for green volume
and 0.69 for ovendry.
Pacific madrone wood has good strength properties. For most of its
common applications (e.g., flooring or furniture), its resistance to inden-
tation and abrasion is a plus. Pacific madrone has exceptional resistance
to breakage, making it suitable for joinery. Because of its hardness, nail-
ing is difficult and splitting is likely unless the wood is prebored. See
Appendix 1, Table 3 for average mechanical properties for small, clear
specimens.

Pacific madrone requires special care during drying because of its
wetwood, which can contribute to collapse. Green MC for this wood
ranges from 68 to 93 percent. Its shrinkage values are considerably
higher than for most other woods, which may result in increased drying
degrade from warp. The radial shrinkage (green to ovendry) is 5.4 per-
cent and the tangential shrinkage is 11.9 percent. For comparison, the
respective values for alder are 4.4 percent and 7.3 percent, and for
white oak are 4.2 percent and 9.0 percent. Lumber cut in a quarter-
sawn pattern will minimize some of the high shrink/warp potential; oth-
erwise, careful design consideration is a must. Because the tree does not
always grow straight, tension wood sometimes forms, which will con-
tribute to nonuniform shrinkage. Presteaming the kiln charge and stickering
at a closer interval has been used successfully to control warp. (See
Tables 8a and 8b for the appropriate kiln schedules).
Prior to kiln drying, Pacific madrone can develop a chemical oxida-
tive stain that appears as blue or purple streaking in the wood. It does
not show on rough-sawn surfaces of the wood and is apparent only after
planing. To minimize staining, madrone should be dried as soon as
possible after sawmilling, and tight stacking of wet lumber should be
avoided.
Machining
Of all the hardwoods of the Pacific Northwest, Pacific madrone ranks
highest (fewest machining defects) for planing, shaping, boring, and
turning. Because of its high density, it should not be processed too fast
84
Table 8a. Kiln schedule—Pacific madrone 4/4, 5/4, 6/4.
Equilibrium
1 Above 35 110 106 17.6 87 43.5 41.0

2 35 to 30 110 105 16.3 84 43.5 40.5
3 30 to 25 120 112 13.5 77 49.0 44.5
4 25 to 20 130 116 10.1 65 54.5 46.5
5 20 to 15 140 110 5.8 38 60.0 43.5
6 15 to final 180 130 3.5 26 82.0 54.5
Table 8b. Kiln schedule—Pacific madrone 8/4.

Equilibrium
1 Above 35 110 107 19.1 90 43.5 41.5

2 35 to 30 110 106 17.6 87 43.5 41.0
3 30 to 25 120 114 15.5 83 49.0 45.0
4 25 to 20 130 120 12.2 74 54.5 49.0
5 20 to 15 140 115 6.8 46 60.0 46.0
6 15 to final 160 110 3.4 21 71.0 43.5
(overfeed). It is recommended that saws and other tooling have a hook

angle of 20∞ and a sharpness angle of 55∞ for optimum performance. As
with other fine-grain, hard woods such as birch or maple, surface scratching
(cross-grain or swirls) during sanding can be a problem with Pacific
madrone.
Adhesives
Pacific madrone bonds well; there are no unusual problems with
this wood when gluing conditions are moderately well controlled. Care-
ful curing/drying of glue joints is required to prevent sunken gluelines
from subsequent machining.
Finishing
Pacific madrone finishes well, without the need to fill the grain; it
colors best with dyes or transparent stains. Heavily pigmented stains
tend to be muddy in appearance. Pacific madrone can be successfully
ebonized.
85
Durability
Pacific madrone is a nondurable species that is susceptible to wood
decay. Untreated wood posts in ground contact have an average service
life of 6 years. Mold and oxidative staining are moderate problems.
Uses
Pacific madrone is used for furniture, flooring, turnings, paneling,
veneer for hardwood plywood faces and core stock, pulpwood, and
firewood.
Related Literature
DICKINSON, F.E., D.R. PRESTEMON, and W.A. DOST. 1965. Pacific madrone
log grades and lumber yield. California Agriculture Experiment Sta-
tion, Berkeley, California. California Forestry and Forest Products No.
43.
McDONALD, P.M. 1992. Estimating seed crops of conifer and hardwood
species. Canadian Journal of Forest Research 22:832-838.
compil. USDA Forest Service, Washington, DC. Agriculture Hand-
book 445.
McDONALD, P.M., and J.C. TAPPEINER II. 1990. Pacific madrone. P.
124-132 in Silvics of North America. Volume 2, Hardwoods. R.M.
Burns and B.H. Honkala, coords. USDA Forest Service, Washington
D.C. Agriculture Handbook 654.
RAPHAEL, M.G. 1987. Use of Pacific madrone by cavity-nesting birds. P.
198-202 in Proceedings of the Symposium on Multiple-Use Manage-
ment of California’s Hardwood Resources. T.R. Plumb and N.H. Pillsbury,
tech. coords. USDA Forest Service, Pacific Southwest Forest and Range
Experiment Station, Berkeley, California. General Technical Report
PSW-100.
TAPPEINER, J.C., II, P.M. McDONALD, and T.F. HUGHES. 1986. Survival
of tanoak (Lithocarpus densiflorus) and Pacific madrone (Arbutus menziesii)
seedlings in forests of southwestern Oregon. New Forests 1:43-55.
TINNIN, R.O., and L.A. KIRKPATRICK. 1985. The allelopathic influence
of broadleaf trees and shrubs on seedlings of Douglas-fir. Forest
Science 31:945-952.
ZUMRAWI, A.A., and D.W. HANN. 1989. Equations for predicting the
height to crown base of six tree species in the central western Willamette
Valley of Oregon. Forest Research Laboratory, Oregon State Univer-
sity, Corvallis. Research Paper 52. 12 p.
86
RED ALDER
ALNUS RUBRA
OREGON ALDER
WESTERN ALDER
PACIFIC COAST ALDER
Alders are members of the birch family (Betulaceae). Of the ten

species of Alnus native to the United States, red alder is the only one
Red Alder Range that reaches commercial size and abundance. It is also the most com-
YUKON
mon and important of the hardwoods in the Pacific Northwest.
BRITISH
COLUMBIA
Mature red alder trees are typically 70 to 120 ft in height (130 ft
maximum) and 10 to 34 in. in diameter (70 in. maximum). Red alder
are mature at 60 to 70 years; they seldom survive beyond 100 years. In
forest stands, red alder develops a clear (60 to 70 percent of total
height), slightly tapered bole with a narrow, domelike crown. Open-
grown trees form broadly conical crowns and highly tapered boles, of-
ten with large forks and branches. The root system of red alder is shal-
low and spreading where limited by poor drainage; a deep-root system
develops on soils with better drainage.
WA
OR Geographic Range
ID
The range of red alder extends from southeastern Alaska (lat 60∞N)
to southern California (lat 34∞N), generally within 125 miles of the
ocean. Red alder is common at low elevations throughout the Coast and
CA north Cascade ranges but is restricted to riparian areas or moist microsites
farther south.
87
Timber Inventory
Historical inventories indicate that the abundance of red alder has in-
creased about 20-fold since the 1920s, though this trend may be reversed
by modern forest practices, which favor conifers. The current inventory of
about 7.4 billion cubic feet of red alder comprises 60 percent of the total
hardwood volume in the Northwest (Appendix 1, Table 1). The greatest
volume occurs in the Puget Sound and Northwest Oregon subregions. A
significant portion of the red alder resource is not available for harvest;
forest practices rules constrain timber management in riparian areas where
red alder is most abundant. Also, very little red alder is sold from public
lands, although substantial inventory occurs there.

Red alder is intolerant of shade, and it must maintain a dominant or
codominant canopy position. Trees of intermediate or suppressed-crown
classes do not survive long. Both pure and mixed-species stands are
predominantly even-aged. In mixed stands, red alder are usually grouped.
Ecological Role
Red alder is a pioneer species that establishes rapidly in openings
created by forest disturbance; it commonly invades newly bared soils
after landslides, logging, or fire. Red alder can maintain or improve
soils via rapid input of organic matter and nitrogen. Its roots fix atmo-
spheric nitrogen via symbiosis with the actinomycete, Frankia. Red alder
does not reproduce in the absence of soil disturbance.
Red alder often occurs in mixture with other tree species. Common
associates include Douglas-fir, western redcedar, western hemlock, grand
fir, Sitka spruce, bigleaf maple, vine maple, black cottonwood, Pacific
willow, and bitter cherry. Common shrubs and herbs associated with
red alder are salmonberry, thimbleberry, red elderberry, devil’s-club,
whortleberry, osoberry, evergreen blackberry, western swordfern, and
hedge nettle.

The suitability of specific sites should be carefully assessed before
red alder management is planned. Although red alder colonizes a wide
variety of sites, many of those sites present high risks of tree mortality,
persistent damage, or poor growth and are thus unsuitable for timber
88
management. Good sites for red alder are generally found along streams,
in moist bottomlands, and on lower slopes. Growth of red alder can also
be quite good on upland sites (below 2000 ft) with adequate soil mois-
ture and a favorable climate.
When representative red alder trees are present, site index should be
estimated with either the 20-year base age (Harrington and Curtis 1985)
or the 50-year base age (Worthington et al. 1960). Harrington’s 1986
study, “A method of site quality evaluation for red alder,” should be
used for evaluating a site when there are no representative red alder
present.
Climate
The typical climate in the range of red alder is mild and humid.
Most precipitation occurs as rain in the winter; summers are generally
cool and dry. Better red alder sites receive occasional rain and frequent
morning fog during the summer. Annual precipitation ranges from 16
to 220 in. (405 to 5600 mm) and temperatures range from -22 to 115
∞F (-30 to 46.1 ∞C).
For red alder, risks of excessive mortality and damage from sunscald,
heat, or drought are high on southerly aspects, particularly inland on
steep slopes. Planted red alder seedlings are particularly susceptible.
Near the coast, higher humidity and soil moisture provide more favor-
able conditions on any aspect.
Good development of trees occurs where annual precipitation ex-
ceeds 40 in. or where roots have access to ground water. Red alder do
poorly under droughty conditions, which may result from inadequate
annual or seasonal precipitation, low moisture-holding capacity of the
soil, or high evapotranspiration, together or singly.
Severe freezing or unseasonable frost hazards can greatly limit man-
agement of red alder. Local frost pockets and flat areas that accumulate
cold air from large, cold-air drainages are poor sites for red alder. Both
late spring and early fall frost can be disastrous to young plantations.
Cumulative effects of periodic frosts produce poor quality stands.
Periodic exposure to high winds can greatly reduce stem quality and
height growth of red alder. Areas exposed to periodic high winds (>50
mph) and coastal sites that are not protected from prevailing winds
should be avoided.
Elevation
Management of red alder should generally be restricted to eleva-
tions below 3000 ft at the southern end and 1000 ft at the northern
end of red alder’s range.
Soils
Although red alder is found on a wide range of soils, the most
productive stands occur on deep, well-drained loams and sandy loams
derived from marine sediments or alluvium. There are also good red
89
alder sites on soils of volcanic origin. Plentiful soil moisture during the
growing season is necessary for good development of red alder. Exces-
sive drought is produced by soils with low water-holding capacity in-
cluding coarse-textured soils (sandy loams or sands) or soils with high
rock fragment contents (>40 percent by volume). Coarse soils with con-
sistent subsurface moisture (flood plains, riparian areas) are acceptable,
although drought hazards are still high during stand establishment, par-
ticularly if competing vegetation is present.
Red alder tolerates poor drainage and occasional flooding during
the growing season. Sites with very poor drainage or sites subject to
prolonged flooding during any season are not suitable for management
of red alder plantations.
Soils low in available phosphorus (P) greatly limit establishment and
growth of red alder, although specific criteria for determining deficiency
of P in soils have not been developed for red alder. Deficiency of P in
red alder is indicated by foliar concentrations of less than 0.16 percent.
Deficiency of soil nitrogen (N) is of lesser concern for red alder. Nitro-
gen fixation via red alder’s symbiotic association with Frankia can com-
pensate for deficiencies in soil N.

Trees reach sexual maturity as early as 3 to 4 years of age. Domi-
nant trees in a stand usually begin to produce seed at 6 to 8 years of
age. Red alder is monoecious, having separate male and female flowers
on the same individual. Male catkins develop in clumps that hang down.
In late winter, they elongate from 1 to 3 in. and turn from green to
reddish-brown, releasing their pollen in late winter and early spring.
Female flowers are borne in clumps of upright catkins, which later de-
velop into cone-like strobiles that bear the seed. The “cones” begin to
ripen in September or October, changing from green to yellow-green or
brownish-green to brown.
Seed
The seeds are small, winged nutlets borne in pairs at the base of
bracts within the strobiles. Seeds are very light (350,000 to 1,400,000
seeds/lb) and they can be carried long distances by the wind. Seed
dispersal may begin in late September; most seeds are released from
late fall through winter. Seed should be collected from a local source to
ensure that seedlings are adapted to conditions on the outplanting site.
Cones should be collected from numerous trees of good growth and
form that are well distributed within a stand. The quality and quantity
of the cone crop should be assessed in July or August. Collection of
cones may begin when the color of a cone has changed to about 50
percent yellow. Another test for crop maturity is to twist cones along
the long axis. Seeds are ripe if the cone twists easily and the bracts
separate.
After collection, cones should be airdried in paper or cloth bags.
Care must be taken to provide adequate ventilation and prevent mold-
ing. When cones have dried, seed should be extracted via thrashing in a
tumbler or by hand (for small lots). Yield may be increased by repeated
90
wetting, redrying, and extracting. Extracted seeds are screened to re-
move large debris. Air column machines can be used to remove small
trash and empty seed. For short-term storage, dry seed can be stored in
sealed containers in the refrigerator with no loss in viability. Red alder
seed may be stored for 5 to 10 years with little loss in viability when
dried to less than 10 percent moisture content (MC) and stored in
sealed containers in the freezer.

Dissemination of light red alder seed by the wind commonly pro-
duces widespread colonization on disturbed soils under a variety of
conditions. Very little work has been done to develop methods of inten-
tional regeneration of red alder from seed, however. Establishment from
seed generally requires open conditions and bare mineral soil; red alder
seedlings become established on organic substrates only under very
moist conditions. Excessive heating or drying of the soil surface at any
time greatly limits establishment of red alder from seed.
High humidity and soil moisture near the coast or at the north end
of red alder’s range provide favorable conditions on almost any aspect.
In the interior Coast Range or Cascade foothills, establishment from
seed is practically zero on southern aspects, and it may be limited to
wet microsites and lower slopes on northern aspects.
Adequate distribution of seeds can be provided by well-distributed
seed trees or a seed “wall” adjacent to the selected unit. Smaller clear-
ings (<20 acres) with a seed source on at least two sides can regenerate
well. Isolated seed trees left after harvest may not stand very long. Seed
trees on the north side of a unit are preferable, since dispersal is ac-
complished primarily by drying north winds in the late fall and winter.
Conditions favorable for natural regeneration of red alder often pro-
duce an overabundance of seedlings (exceeding 100,000 stems per acre),
and early precommercial thinning may be necessary to prevent stagna-
tion or poor growth.

Young red alder will sprout vigorously after cutting (coppicing).
Coppices with rotations of 4 to 6 years have been managed successfully
for a few rotations. Red alders more than 10 years old do not sprout
well after cutting; regenerating red alder by coppicing older stands is
not feasible.
Red alder are not easily established from unrooted cuttings. Cut-
tings of greenwood from young trees can be rooted by dipping in
indole-3-butyric acid and culturing in a warm, well-aerated medium.
Tests of operational regeneration from rooted cuttings have been mini-
mal.

Planting of seedlings allows greater flexibility in site selection and
provides greater control over spacing and seed source compared to
91
regeneration from seed. Vigorous, planted red alder seedlings will have
an advantage over competing vegetation. Seedlings of good quality,
planted on well-prepared sites can reach heights of 4 to 7 ft after the
first growing season.
Plantations of red alder can be successfully established with a variety
of seedling stocktypes, but many efforts have failed because of poor
quality seedlings, extreme weather, and other hazards. Consistent suc-
cess requires a careful evaluation of regeneration hazards, along with
adequate seedling quality, and good site-preparation and planting prac-
tices. Red alder seedlings that will have the best survival rate, growth
rate, and resistance to damage over a range of conditions are character-
ized as follows:
• Height of 12 to 36 in. and basal diameter (caliper) of at least 0.16
in. (4 mm)
• Stocky, rather than tall and thin
• Healthy buds or branches along the entire length of the stem,
particularly the basal portion
• Full, undamaged fibrous root systems
• Free of disease.

Vigorous red alder seedlings can compete successfully with little or
no site preparation when levels of competing vegetation are low to
moderate. Moderate amounts of slash, debris, and vegetation shelter
new seedlings and may also improve establishment. With high levels of
competing vegetation, site preparation is required to achieve adequate
stocking and good performance. Growth of red alder seedlings may be
lower if the cover of competing vegetation exceeds 90 percent during
the first year. Survival may be reduced by competition from 125 to 150
percent cover with overtopping in the first year.
Broadcast burning often provides adequate site preparation where
levels of slash and/or shrub cover are high. Chemical site preparation
may be most cost-effective for controlling both shrubby and herbaceous
competitors. When a site has been heavily invaded by herbs, herbicide
treatments just before planting can make the difference between suc-
cess and failure of hardwoods.
When regeneration is directly from seed, site preparation should
produce an even distribution of bare mineral soil. Mechanical scarifica-
tion, broadcast burning, or piling and burning will do this in most
situations. To prevent overabundant regeneration, one method is to
minimize soil disturbance during harvest and then mechanically scalp
evenly spaced spots throughout the unit. Closely spaced red alder
seedlings (<9 ft) can effectively dominate a site within 2 to 4 years,
thereafter, site-preparation treatments are unnecessary. Red alder at wider
spacings (10 to 20 ft) are vulnerable to the prolonged effects of vegeta-
tive competition. At these wider spacings, maintenance of weed-free
conditions after establishment can double to quadruple seedlings’ growth
in comparison to unweeded trees.
92
Stand Management
Natural stands of red alder generally establish at high densities (10,000
to 100,000 stems per acre); intense competition causes rapid self-thin-
ning and slow diameter growth. Management of lower initial densities
(300 to 600 stems per acre) can increase diameter growth rates on crop
trees 15 to 20 percent compared to unmanaged stands during the first
15 years. Continued thinning (pulpwood, fuelwood, precommercial thinning)
can maintain diameter growth rates up to 30 percent higher than those
in unmanaged stands, at least until age 25. Managed stands are ex-
pected to attain an average diameter of 12 in. by age 30 or before; the
average natural stand would take 45 years (SI 50 = 100 ft).
Guidelines for management of stand density are provided by the
density management diagram (Puettmann et al. 1993). Thinning must
favor trees with good growth potential (dominant or codominant trees
less than 15 to 20 years old). It is not worthwhile to thin older stands or
to leave suppressed trees because the remaining trees will not have
adequate capacity for growth response.
Some crowding is necessary to maintain dominance of red alder and
to reduce branching, forking, and stem taper. The goal is to manage
spacings that optimize growth while maintaining the benefits of crowd-
ing. Moderate crowding will induce lower branch mortality with mini-
mal reductions in diameter growth. Relatively uniform spacing in man-
aged stands will also improve stem form by producing straighter stems.
Red alder grow towards the light; clumpy spacing and large holes in the
stand increase lean and sweep.
Initial spacings of 9 to 10 ft between trees should shade out lower
branches 30 to 40 ft up the bole by ages 8 to 15 years. A subsequent
thinning, combined with pruning of dead branches (many are broken
off during thinning) will maintain diameter growth on a high-quality
bole. Pruning of live branches may also increase wood quality, although
little work has been done on this.
Because of red alder’s ability to improve soils via N-fixation and
addition of organic matter, there is particular interest in managing red
alder in mixture with conifers in order to maintain or improve site pro-
ductivity. Management of mixtures can be difficult because of red alder’s
rapid height growth and great sensitivity to competition. Under favor-
able moisture conditions, red alder will overtop and suppress conifers
established at the same time. Low proportions of red alder may be
difficult to maintain over the long term, because red alder must main-
tain codominance in order to thrive.
Strategies for managing mixtures include (1) delaying the establish-
ment of red alder for at least 3 to 6 years, (2) maintaining a low propor-
tion of red alder in the stand (10 to 20 percent by stem count) and, (3)
managing mixtures in small patches of single species, similar to most
natural mixtures.
93
Growth and Yield
On good sites, height growth may exceed 6 ft/year for the first five
years, and trees may attain heights of 60 to 80 ft in 20 years. Mean
annual production rates in young stands have been estimated at 6.8 dry
tons per acre. Growth slows substantially after the juvenile stage, par-
ticularly on poor sites. Site index ranges from 33 to 82 ft for base age
20 years and 60 to 120 ft for base age 50.
Yield tables based on site index and stand basal area (Chambers
1983) are available for estimating volumes of red alder in natural stands.
Maximum volume per acre for red alder typically occurs at age 50 to
70, ranging from 5000 to 7000 ft 3 per acre. On very good sites, annual
volume growth rates may average 300 ft 3 per acre for the first 10 years
and 200 ft 3 per acre over 30 years.
Relatively little information is available on growth and yield in man-
aged stands of red alder. Major gains in average stem diameter and stand
basal area appear to be possible with management of spacing in young
stands. Optimistic projections anticipate sawlog rotations of 30 to 35 years
for managed stands compared to 45 to 50 years for natural stands.

For wildlife, red alder provides an important deciduous component
in the predominantly coniferous forests of the Northwest. Typically,
shrub and herb vegetation under red alder is quite different from that
of conifer-dominated areas. A variety of animals seem to prefer or de-
pend on red alder for food or habitat. Maintenance of a red alder
component can provide greater habitat diversity within or between co-
nifer stands.
Browsing, antler rubbing, and trampling by deer and elk can cause
serious problems in young plantations. Red alder are very sensitive to
this damage; effects on young trees include decreased growth, multiple
stems, and poor stem form. Rapid growth and close spacings generally
ensure that an adequate number of crop trees will escape serious dam-
age. Risks of permanent damage are highest with plantations estab-
lished at wide spacings (>12 ft). Areas of concentrated use by elk or
deer should not be managed for red alder.
Both mountain beaver and fur beaver can cause substantial damage
to seedlings. Planted seedlings may be the major food source for moun-
tain beaver during the first years after burning or chemical site prepara-
tion. Preventative measures such as trapping should be considered if
there is evidence of a significant mountain beaver population. Fur bea-
ver can cause extensive mortality of saplings and trees up to 150 ft
from streams.
Voles, mice, and other rodents often severely damage seedlings,
particularly in grassy or marshy areas. Basal netting or tubing can pro-
tect seedlings from rodents.

Young, undamaged red alder stands are fairly free of problems from
insects and disease. Stem cankers are common in some young stands,
94
although they seldom have significant impact except under stressful
environments. Although red alder has long been perceived as highly
susceptible to decay, some recent work shows that healthy, living trees
are exceptionally resistant to decay after typical stem injuries.
Occasionally, serious outbreaks of defoliating insects can cause growth
reductions in healthy stands and mortality in stressed stands. Tent cat-
erpillars (Malacosoma disstria, M. californicum), red alder flea beetle (Altica
ambiens), red alder woolly sawfly (Eriocampa ovata), striped red alder
sawfly (Hemichroa crocea), and a leaf beetle (Pyrrhalta punctipennis) have
all caused damage.
Genetics
Major gains in growth and quality may be possible with selective
breeding of red alder. This is because red alder has a large amount of
genetic variation, early sexual maturity, frequent seed production, rapid
growth, and the capability of vegetative propagation. Little effort has
been made to establish breeding programs.

Both cubic-foot and board-foot volume tables have been developed
to estimate volume in standing trees from DBH and total height. Stan-
dard log grades, adapted from eastern hardwood log grades, have been
developed for red alder. Most pricing decisions, however, are based on
log diameter, length, and grade specifications developed by the specific
log buyer.
Harvesting and transport costs for red alder are often higher than
those for softwoods, although no special logging equipment is required.
Red alder typically has lower volumes per acre and smaller, shorter
trees. Red alder has a high green-weight-to-volume ratio, and natural
stands produce a high percentage of logs with sweep and crook, which
reduces the amount of logs that can be loaded on a truck. Most logging
takes place in the dryer months; harvest volume declines in the rainy
winter months because of road and site conditions.
Logs are generally scaled with Scribner log scale rules. Logs are also
sold by weight or by the truckload. To prevent staining, red alder logs
must be removed from the woods and processed within 6 to 8 weeks in
the summer and 8 to 12 weeks in the winter.
Product Recovery
logs are chipped for pulp. Lumber is graded under special National
Hardwood Lumber Association (NHLA) rules for red alder; grades in-
clude Selects and Better, No. 1 Shop, No. 2 Shop, No. 3 Shop, and
Frame. Unlike the standard NHLA grading rules, these grades are gener-
ally based on the best face of the piece, whereas the other NHLA rules
95
are based on the poorer face. Grades can be applied to rough, surfaced,
green, or dry lumber; in practice, lumber is usually dried and surfaced
before grading. A considerable volume of the low-grade material is sawn
into 1 X 4, 1 X 6, and 2 X 4 for making pallets.
Recent studies show that the cubic volume of red alder that is recov-
erable as lumber ranges from 30 percent in small diameter logs to 50
percent in larger logs. Grade recovery also varies by log size or log
grade; e.g., 85 percent of the surface-dried lumber produced from 7-in.
logs was pallet material, but 75 percent of the surface-dried lumber
from 20-in. logs was No. 1 Shop and Select. An earlier study conducted
with NHLA standard grades (rather than the modified red alder and
maple grades) showed that the average green lumber grade recovery
from alder logs was lower than that of other eastern and western hard-
woods for a given log grade (Appendix 1, Table 2). For a given log
diameter, grade recovery from butt logs is much higher than that for
logs higher in the tree.
Most of the high-grade lumber is used for furniture, cabinets, and
turned products. Lumber prices have remained high and are competi-
tive with prices for eastern hardwoods. Red alder lumber is marketed
internationally, with strong markets in the Pacific Rim countries and in
Europe, especially Italy and Germany.
Red alder is peeled into veneer for both low-grade core stock and
high-grade face material. Veneer logs are an increasingly important market
that is competitive with sawlogs. Red alder is also widely used for pulp,
both domestically and overseas, but staining and fiber deterioration are
a problem in storing pulp chips for more than a few months. An evalua-
tion of red alder as a raw material for structural panels, such as oriented
strand board, found no problems in producing flakes, bonding with
resins, or meeting structural design values.
Wood Properties
Characteristics
The wood of red alder is evenly textured with a subdued grain pat-
tern, and is of moderate weight and hardness. Red alder is a light-
colored or white wood when it is freshly sawn, but with exposure to air,
the wood darkens and changes to a light brown hue with a reddish tint.
There is no color distinction between heartwood and sapwood.
The growth rings are distinct, delineated by either a whitish or brownish
line at the outer margin. The pores are uniformly distributed within a
growth ring (diffuse porous). Rays are present and of two types, narrow
(simple) and broad (aggregate). Both the pores and the rays are indis-
tinct to the naked eye. The wood is without any characteristic taste or
odor.
96
Weight
Red alder weighs about 46 lb/ft 3 when green and 28 lb/ft 3 when
dried to 12 percent MC. The average specific gravity is 0.37 for green
and 0.43 for ovendry.
Because of its moderate specific gravity, red alder is not an excep-
tionally strong wood. In many applications this will be apparent as
indentations on the surface of the wood. In furniture applications, it
may be necessary to redesign joints and the sizes of structural parts to
compensate for the often slightly lower strength values of red alder.
Red alder holds nails well and does not readily split when nails are
driven into it. Lower grades of red alder perform adequately as pallet
material. See Appendix 1, Table 3 for average mechanical properties for
small, clear specimens.

Red alder lumber 5/4 and thinner is one of the easiest North Ameri-
can wood species to dry. Establishing and maintaining uniform color
requires special handling and storage of logs and freshly cut lumber,
and specially developed dry-kiln schedules. Variable coloration is due to
the oxidation of extractives present in the wood. Colors may range
from yellow to deep red and may be mottled.
Kiln-drying the lumber as soon as possible after sawing prevents
mottling. Steaming the kiln charge at different temperatures for differ-
ent lengths of time will result in different colored wood (from white to
dark red); this technique allows the kiln operator to select the desired
final color. See Table 9 for a standard kiln schedule. Other schedules
are available for either lighter or darker final coloring of the wood.
Shrinkage values for green to ovendry wood based on original green
sizes are low and average 4.4 percent in the radial direction and 7.3
percent tangentially. The green MC of the wood averaged 98 percent
(ovendry basis).
Table 9. Kiln schedule—Red alder (standard color) 4/4, 5/4, 6/4.

Equilibrium
1 Above 50 140 133 14.2 82 60.0 56.0

2 50 to 40 140 130 12.0 75 60.0 54.5
3 40 to 35 140 125 9.6 64 60.0 51.5
4 35 to 30 140 115 6.8 46 60.0 46.0
5 30 to 25 150 110 4.4 28 65.5 43.5
6 25 to 20 160 110 3.4 21 71.0 43.5
7 20 to 15 170 120 3.5 24 76.5 49.0
8 15 to final 180 130 3.5 26 82.0 54.5
97
Machining
Red alder has an excellent reputation for machining. Due to the
moderate specific gravity and the even texture of the wood, high throughput
of material is possible. Quality surfaces can be obtained if sharp cutting
edges are used. Some tear-out is possible during planing and shaping if
tooling becomes dull or if feed rates are excessive. Red alder sands well
without scratching and with a minimum of fuzzing. Its turning charac-
teristics are similar to those of black cherry.
Adhesives
The ease of gluing red alder is well known in the industry. It bonds
well and there are no unusual problems when conditions are moder-
ately well controlled.
Finishing
Because of its uniform, small pore structure and the consistency of
color, red alder is a preferred wood for finishing. It accepts a variety of
stain types and has been successfully substituted for other woods when
properly colored stains are applied.
Durability
Red alder is a non-durable wood when subjected to conditions that
are favorable to decay. We recommend that it be rapidly processed into
lumber after harvest to prevent staining and decay. A reddish-purple
stain develops in solid-piled lumber that has not been dried or treated
with anti-stain chemicals. In-ground tests indicate that untreated, peeled
round posts will decay and fail in 3 years on average, while split posts
will last only 5 years.
Uses
Uses for red alder include face veneer, furniture, cabinets, paneling,
edge-glued panels, core-stock and cross-bands in plywood, millwork,
doors, pallets, woodenware and novelties, chips for waferboard, pulp-
wood, and firewood.
Related Literature
AGER, A.A., P.E. HEILMAN, and R.F. STETTLER. 1993. Genetic variation
in red alder (Alnus rubra) in relation to native climate and geogra-
phy. Canadian Journal of Forest Research 23:1930-1939.
AHRENS, G.R., A. DOBKOWSKI, and D.E. HIBBS. 1992. Red alder: guide-
lines for successful regeneration. Forest Research Laboratory, Or-
egon State University, Corvallis. Special Publication 24. 11 p.
98
ATTERBURY, T. 1978. Alder characteristics as they affect utilization. P.
71-81 in Utilization and Management of Alder. D.G. Briggs, D.S.
DeBell, and W.A. Atkinson, compils. USDA Forest Service, Pacific
Northwest Forest and Range Experiment Station, Portland, Oregon.
General Technical Report PNW-70.
BRIGGS, D.G., D.S. DeBELL, and W.A. ATKINSON, compilers. 1978. Uti-
lization and management of alder. USDA Forest Service, Pacific Northwest
Research Station, Portland, Oregon. General Technical Report PNW-
70. 379 p.
CHAMBERS, C.J. 1983. Empirical yield tables for predominantly alder
stands in western Washington. Washington Department of Natural
Resources, Olympia, Washington. DNR Report N. 31. 70 p.
CLEAVES, D.A. 1992. Marketing alder and other hardwoods. Oregon
State University Extension Service, Corvallis, Oregon. Extension Cir-
cular 1377. 8 p.
CURTIS, R.O, D. BRUCE, and C. VanCOEVERING. 1968. Volume and
taper tables for red alder. USDA Forest Service, Pacific Northwest
Forest and Range Experiment Station, Portland, Oregon. Research
Paper PNW-56. 35 p.
DeBELL, D.E. Unpublished data. USDA Forest Service, Pacific Northwest
Research Station, Olympia, Washington.
FEDDERN, E.T. 1978. Harvesting of red alder. P. 61-70 in Utilization and
Management of Alder. D.G. Briggs, D.S. DeBell, and W.A. Atkinson,
compils. USDA Forest Service, Pacific Northwest Research Station,
Portland, Oregon. General Technical Report PNW-70.
GEDNEY, D.R. 1990. Red alder harvesting opportunities in western Or-
egon. USDA Forest Service, Pacific Northwest Research Station, Portland,
Oregon. Resource Bulletin PNW-RB-173. 22 p.
HAEUSSLER, S., and J.C. TAPPEINER II. 1993. Effect of the light environ-
ment on seed germination of red alder (Alnus rubra). Canadian Jour-
nal of Forest Research 23:1487-1491.
HARRINGTON, C.A. 1986. A method of site quality evaluation for red
alder. USDA Forest Service, Pacific Northwest Research Station, Portland,
Oregon. General Technical Report PNW-192. 22 p.
HARRINGTON, C.A. 1990. Red alder. P. 116-123 in Silvics of North
America. Volume 2, Hardwoods. R.M. Burns and B.H. Honkala, coords.
USDA Forest Service, Washington, D.C. Agriculture Handbook 654.
HARRINGTON, C.A., and R.O. CURTIS. 1985. Height growth and site
index curves for red alder. USDA Forest Service, Pacific Northwest
Research Station, Portland, Oregon. Research Paper PNW-358. 12 p.
HIBBS, D.E., and A.A. AGER. 1989. Red alder: guidelines for seed collec-
tion, storage, and handling. Forest Research Laboratory, Oregon
State University, Corvallis, Oregon. Special Publication 18. 6 p.
HIBBS, D.E., D.S. DeBELL, and R. TARRANT, editors. 1994. The Biology
and Management of Red Alder. Oregon State University Press, Corvallis.
256 p.
HIBBS, D.E., W.H. EMMINGHAM, and M.C. BONDI. 1989. Thinning red
alder: effects of method and spacing. Forest Science 35:16-35.
99
JOHNSON, H.M., E.J. HANZLIK, and W.H GIBBONS. 1926. Red alder of
the Pacific Northwest: its utilization, with notes on growth and man-
agement. USDA, Washington, D.C. Department Bulletin 1437.
KOZLIK, C.J. 1987. Presteaming to minimize mottling in partially air-
dried red alder lumber. Forest Research Laboratory, Oregon State
University, Corvallis. Research Note 80. 6 p.
LENEY, L., A. JACKSON, and H.D. ERICKSON. 1978. Properties of red
alder (Alnus rubra Bong.) and its comparison to other hardwoods. P.
25-33 in Utilization and Management of Alder. D.G. Briggs, D.S.
DeBell, and W.A. Atkinson, compils. USDA Forest Service, Pacific
General Technical Report PNW-70.
LOWELL, E.C., and R.L. KRAHMER. 1993. Effects of lean in red alder trees
on wood shrinkage and density. Wood and Fiber Science 25:2-7.
MACKIE, D.M, and G.J. Williams. 1984. Growth and utilization of alder
within the Pacific coastal region. Pulp & Paper Canada 85(8):71-76.
PLANK, M.E., T.A. SNELGROVE, and S. WILLITS. 1990. Product values
dispel “weed species” myth of red alder. Forest Products Journal
40(2):23-28.
PUETTMANN, K.J., D.E. DeBELL, and D.E. HIBBS. 1993. Density manage-
ment guide for red alder. Forest Research Laboratory, Oregon State
University, Corvallis. Research Contribution 2. 6 p.
RAETTIG, T., G.R. AHRENS, and K. CONNAUGHTON. Hardwood supply
in the Pacific Northwest: a policy perspective. USDA Forest Service,
Pacific Northwest Research Station, Portland. In preparation.
RESCH, H. 1980. Utilization of red alder in the Pacific Northwest. Forest
Products Journal 30(4):21-26.
RESCH, H. 1988. Red alder: opportunities for better utilization of a re-
source. Forest Research Laboratory, Oregon State University, Corvallis.
Special Publication 16. 13 p.
RHOADES, C.C., and D. Binkley. 1992. Spatial extent of impact of red
alder on soil chemistry of adjacent conifer stands. Canadian Journal
of Forest Research 22:1434-1437.
WORTHINGTON, N.P, F.A. JOHNSON, G.R. STAEBLER, and W.J. LLOYD.
1960. Normal yield tables for red alder. USDA Forest Service, Pacific
Research Paper 36.
100
TANOAK
LITHOCARPUS DENSIFLORUS
TANBARK-OAK
Tanoak is a unique evergreen hardwood that belongs to the beech

family (Fagaceae). It has characteristics similar to both oak and chest-
nut. All other 100 to 200 species of Lithocarpus are native to southeast
Asia or Malaysia. In southwestern Oregon, tanoak is second to Pacific
madrone as the most abundant hardwood.

Mature tanoak trees typically attain heights of 50 to 150 ft (208 ft
Tanoak maximum) and diameters of 6 to 48 in. (109 in. maximum). Tanoaks
Range
may live at least 250 years, although age can be difficult to deter-
mine. In dense forest stands, tanoaks develop narrow crowns with
BRITISH
COLUMBIA ascending branches on single, straight stems that are clear for 30 to 80
ft. In open stands, tanoaks often have broad crowns with large horizon-
tal branches on short, forked boles. The root system of tanoak is com-
posed of both deep and extensive lateral roots.
WA
OR
Geographic Range
Tanoak is native to a relatively narrow range on the West Coast from
Santa Barbara, California, to Reedsport, Oregon (lat 34 to 43∞N). It is
most common in the Coast ranges and Siskiyou Mountains in California
and southwestern Oregon, and can also be found in the Sierra Nevada.
CA
Timber Inventory
Growing stock volume in southwestern Oregon is about 350 MMCF
(Appendix 1, Table 1), much of it on federal lands (269 MMCF). The
101
inventory in California is 1938 MMCF; this considerably larger volume has
supported better market development in California than in Oregon.

Tanoak is very tolerant, and it commonly occurs in any crown posi-
tion. Seedlings and saplings are quite common in the understory of
hardwood or conifer stands. Understory tanoak can grow rapidly in
response to death or removal of overstory trees. Tanoak of any age
seem to prefer some shading from codominant stems within clumps.
Sudden, excessive exposure of stems or crowns is detrimental. Tanoak
typically occurs as a codominant tree with other hardwoods, often with
an overstory of conifers. Pure stands of tanoak are also common during
early stages of succession after fire or logging, however.
Ecological Role
Tanoak is a persistent, versatile species capable of maintaining a
substantial component at any successional stage in many forest types.
While tanoak is considered to be a climax species capable of reproduc-
ing in undisturbed forests, it is also well-adapted to disturbance. It
sprouts vigorously after fire or cutting and often dominates large areas.
Fire is the most important factor determining the fate of individual
tanoaks and the abundance of the species in general. Fire often kills
tanoak stems, although non-fatal injury may be more common. Stands
with a substantial component of tanoak may be less flammable than
stands of pure conifers.
Tanoak is the most important hardwood species in the Mixed-Ever-
green Zone of vegetation. The most common associate trees are Dou-
glas-fir, Pacific madrone, and redwood. Other common associates are
giant chinkapin, canyon live oak, California black oak, California-laurel,
white fir, ponderosa pine, western hemlock, and Sitka spruce.
Shrubs commonly associated with tanoak include blueblossom, ha-
zel, evergreen huckleberry, salal, Pacific rhododendron, Oregon-grape,
western poison-oak, red-flowered currant, and thimbleberry. Important
herbaceous associates include western bracken, western swordfern, New
Zealand fireweed, bull thistle, western whipplea, California brome, Cali-
fornia fescue, and California sweetgrass.

A high proportion of tanoak in the canopy often indicates high site
productivity in general. The proportions of associated Pacific madrone,
California black oak, and canyon live oak tend to increase on warmer,
102
drier sites, which are less productive for tanoak. Rapid early growth of
tanoak stump sprouts occurs regardless of site quality. The capability of
a site for growing tanoak should be evaluated by examining growth and
form of older trees. Good growth potential for tanoak on a site is indi-
cated by the following:
• Top height on mature trees of at least 80 ft, up to 150 ft on the best
tanoak sites
• Sustained height growth of 1 to 2 ft per year from age 5 to 20 years
Climate
Tanoak thrives in a mild, humid climate characterized by dry sum-
mers and wet winters, where snow is infrequent and ephemeral. In this
climate, annual precipitation ranges from 40 to 140 in., typically with
less than 5 percent falling from June through September. Mean daily
temperatures range from a minimum of 36 to 42 ∞F in January to a
maximum of 60 to 74 ∞F in July.
Tanoak’s large, deep root system and sclerophylous leaves impart some
resistance to heat and drought. Compared to other sclerophylls (plants
with thick, shiny blue-green leaves), however, tanoak requires more mod-
erate temperatures and high levels of moisture; it shows little control over
stomatal water loss. Tanoak grows best close to the coast where rain and
fog, high humidity, and low clouds are plentiful. It prefers shade and dense
foliage, which indicates a relatively high sensitivity to heat or sunlight.
Mature tanoaks often decline in vigor when codominant neighbors are
removed, causing sudden increases in exposure.
Severe cold is rare in the natural range of tanoak. Significant die-
back of foliage has been observed after a hard freeze. Heavy snow or ice
may bend saplings to the ground and cause sprouts to break at their
connection to the stump. Sound, undamaged tanoaks are windfirm.
Elevation
At the southern limit of its range, tanoak grows at 2400 to 4700 ft
elevation on the coast and up to 6500 feet in the Sierra Nevada. It is
most abundant from 500 to 3000 ft in the central and northern part of
its range.
Soils
Tanoak grows best on deep, well-drained, sandy, or gravelly loams,
which may be derived from a variety of parent materials. The most
productive soils for Douglas-fir or redwood are also some of the best
soils for tanoak. It seldom thrives on heavy clay soils. Tanoak is also
common on shallow, stony soils, usually on north slopes. It will not
tolerate poor soil drainage or flooding.

Sprouts as young as 5 years old may produce seed. Abundant seed
production generally begins after 30 to 40 years. Most tanoaks bloom
103
during the summer months, although some flowering may occur in the
spring or fall. Separate male and female flowers are borne on the same
plant. Most flowers of both sexes grow from the axils of new leaves on the
current year’s shoot. The flowers are borne in dense, yellowish blossoms
composed of clusters of female flowers at the base of erect male catkins (2
to 4 in. long). Seed ripens in the second autumn after pollination.
Seed
A tanoak seed is similar to an oak acorn, about 0.75 in. long with a
shallow, hairy cup. There are about 110 acorns/lb. Tanoak seed may be
collected from late September to mid-November. Early acorns are often
infested with insects; later acorns are generally sound, especially in a mast
(high seed production) year. Acorns should be collected soon after they fall
to reduce losses in viability from exposure to heat or drying. Germination
and seedling emergence usually occur in spring. Acorns should be planted
immediately or stored under cool, moist conditions (33 to 34 ∞F) until
spring planting. Estimates of germination rate vary from 19 to 80 percent.
Acorns should be planted with the pointed end up.

Natural reproduction of tanoak from seed is often quite abundant.
The best conditions for seedling establishment occur in undisturbed
duff and litter under partial to full forest cover. There, regeneration
occurs as a steady accumulation over long periods of time. Although
acorns germinate under other conditions, their survival is poor in clearcuts
and heavily disturbed environments.
Even under good conditions, only a small percentage of seeds be-
come established seedlings. Regeneration is successful because tanoaks
produce large amounts of seed. Mature trees may produce as many as
110,000 seeds per year. Tanoaks generally produce some seeds every
year and usually bear heavy crops of acorns every two years.
The low percentage of acorns that become established seedlings is
primarily due to predation by insects, birds, rodents, deer, bears, and
raccoons. Predation is particularly high in clearcuts and exposed areas.

Tanoak produces vigorous basal sprouts under a variety of condi-
tions. Sprouting is most vigorous after fire or cutting. Sprouts usually
originate from belowground burls, which are characteristic of tanoak of
any size. To promote development of better quality sprouts, stumps
should be cut low to the ground.

Very little effort has been made to regenerate tanoak with planted
seedlings. Seedlings established from broadcast seeds have performed
poorly, particularly in well-prepared sites under open conditions that
104
are successful for plantations of other hardwoods. Better results may be
possible with the outplanting of vigorous nursery stock, as demonstrated
for some true oaks. In comparison to sprout regeneration, however,
seedlings have lower rates of survival and growth.

origin. Regeneration and growth are enhanced by burning or mechani-
cally removing slash that shades tanoak stumps. Pre-established roots
and rapid growth of sprout clumps make tanoak a superior competitor
in the new stand. With high densities of parent stumps, rapid develop-
ment of tanoak cover will inhibit establishment and growth of compet-
ing species. At lower densities of tanoak, control of competing herbs
and shrubs can improve the growth of young tanoak sprouts.
Site-preparation treatments that leave intact litter and partial pro-
tection (debris, vegetation) may be best for promoting establishment
and growth of tanoak seedlings.
Stand Management
Tanoak sprouts initiate at very high densities (up to 500 per par-
ent), after which self-thinning and expression of dominance proceed
rapidly. Thinning young sprout clumps at 3 to 10 years is not effective
because of the abundant resprouts.
Thinning may improve growth and quality in older stands of tanoak.
Density should not be reduced below about 100 ft 2 per acre of basal
area. To avoid excessive exposure of residuals, thinning should be done
in small steps of about 25 ft 2/acre at 3-to-5-year intervals, if necessary.
Thinning should leave well-formed, dominant stems that originate from
the ground. Companion sprouts with “V”-type connections should be
left intact or cut as a unit.
Tanoak can grow in any crown position in stands with mixed spe-
cies or age classes. Management of mixed stands is complex, however,
and will require aggressive control of tanoak to maintain diverse com-
ponents of hardwoods and conifers, which are generally less tolerant of
shade. Tanoaks grow very slowly in dense shade, but understory sap-
lings respond quickly to increased light. Larger, codominant trees may
decline if stands are opened too much.
Growth and Yield

Seedlings grow slowly, generally averaging 2 to 8 in. in annual
height growth during the first 5 years. Sprouts may grow up to 5 ft in
the first year, and average 2 ft per year for the first 15 to 20 years.
Fully stocked tanoak stands of sprout origin rapidly accumulate basal
area. On one good site, sprout basal area reached 100 ft 2 per acre after
105
9 years. After 50 years or more, typical basal areas can range from 160
to 260 ft 2 per acre.
Typical stand volumes range from 2000 to 4000 ft 3 per acre, with
volumes as high as 7000 ft 3 per acre occurring in small productive
patches of up to several acres. The net annual growth of tanoak in
California averages 3.5 percent of standing inventory, the highest rate
of any hardwood in California.
There has been very little management of tanoak stands. Average
annual growth rates of 85 ft 3 per acre were measured in 50-to-60-year-
old tanoak stands after they were thinned to a basal area of 102 to 125
ft 2 per acre on a high site in northern California. Unthinned stands on
the same site had growth rates of 198 ft 2 per acre.

Tanoak acorns are an important food source for many animals, in-
cluding birds, rodents, deer, bears, and raccoons. A component of tanoak
provides thermal cover, refuge, and nesting habitat for wildlife. North-
ern flying squirrels, Allens’ chipmunks, and dusky-footed woodrats are
closely associated with tanoak. Combined with an overstory of large
conifers, a component of tanoak may improve habitat for northern spot-
ted owls by providing canopy structure and habitat for prey such as
woodrats and northern flying squirrels.

Fungi commonly enter tanoak trunks that have been injured by fire.
The combination of fire injury followed by fungal decay is frequent,
producing a high incidence of defect in older trees. Common fungi that
decay living trees include a brown cubical rot (the beefsteak fungus,
Fistula hepatica), a white root rot, weeping conk (Inonotus dryadeus),
and a sap rot (Schizophyllum commune).
Genetics
There are no known hybrids of tanoak. A shrub form of tanoak (Lithocarpus
densiflorus var. echinoides) grows on moist sites at elevations higher
than the tree form.

volume. A recent study of tanoak lumber recovery found that, even
though eastern hardwood grades will separate logs into distinct value
106
classes, the impact of log diameter on lumber value is more important.
Shifts in lumber values and competition with eastern hardwood species
may make the use of log grades more important in the future.
Product Recovery
Sawlogs usually have a minimum small-end diameter of 6 to 10 in.,
smaller logs are generally chipped for pulp. If tanoak logs are not processed
shortly after their removal from the woods, the lumber commonly end-
splits. One study showed that the percentage recovery of No. 1 Common
or Better green lumber from grade 1 tanoak logs is slightly lower than the
recovery from eastern oaks. The lumber grade recovery from grade 2 and 3
logs compares favorably with the lumber grade recovery from eastern oaks
and is higher than from the other hardwoods (Appendix 1, Table 2). Even
high grade tanoak logs yielded a comparatively low percentage (29 per-
cent) of Select or Better lumber, however.
Tanoak has also been found to yield good quality veneer for furni-
ture production. Heat treatment of the blocks to 160 ∞F improved the
smoothness, tightness, and quality of the veneer without increasing the
amount of end splitting. Defects included surface checking and associ-
ated staining, and knots greater than 2 in. The interior cores produced
veneer that was rough and buckled, so the cores should be left larger
than normal. There were no problems drying the veneer.
Wood Properties
Characteristics
Tanoak is a hard, heavy wood that in many ways resembles the true
oaks; thus, tanoak is often included in discussions about lumber from
western oak species. The wood is a light, reddish-brown when freshly
cut, but it ages to a tannish, reddish-brown. The sapwood is very wide;
it is difficult to distinguish the heartwood from the sapwood. There is
some opinion that true heartwood does not exist in this species. The
growth rings are difficult to distinguish and are delineated only by a
faint narrow line of darker, denser tissue at the outer margin. The infre-
quent pores are barely visible to the naked eye, are unevenly distrib-
uted, and are inserted in light-colored tissue in streamlike clusters that
extend across several or many growth rings. As a result, the wood is
evenly textured, with little apparent grain. There are broad rays, how-
ever, which are conspicuous and especially prominent on quartersawn
surfaces. Narrow rays are also present, but require magnification to view.
When dry, the wood has no characteristic odor or taste.
Weight
Tanoak weighs about 62 lb/ft 3 when green and 41 lb/ft 3 at 11 per-
cent moisture content (MC). The average specific gravity is 0.54 (green)
or 0.66 (ovendry).
107
The wood of tanoak is highly rated for its strength properties. It is
noted for hardness, resistance to abrasion, stiffness, and bending strengths.
It holds fasteners well, but requires preboring before nailing to prevent
splitting. See Appendix 1, Table 3 for average mechanical properties for
small clear specimens.

Tanoak lumber requires special care and well-controlled conditions
during drying to properly lower MC without causing excessive degrade.
It is no more difficult to dry than some eastern or other West Coast oak
species, however, unless it has mineral streak. Poor or uncontrolled
drying will cause defects such as end and surface checking in the lum-
ber. In addition, tanoak lumber can be degraded with honeycomb and
collapse if the drying rate is too rapid. These problems increase when
lumber with mineral streak is dried; in many cases, it is not worth the
effort to dry mineral streak material.
Tanoak can stain if certain metals contact the wet wood; contact
with iron can produce a pronounced blue-black discoloration. Molds
and bacterial stains will develop if air circulation around the wet lumber
is inadequate. Proper air-drying of tanoak requires very mild conditions,
with moderate temperatures and high humidities, to avoid developing
molds and bacterial stains. Poor handling of green wood can also pro-
duce sap staining. Air-drying will minimize much of the drying degrade
and will reduce the total kiln times as well.
The green MC of tanoak is reported to be 115 percent (ovendry
basis). Shrinkage values for green to ovendry lumber (based on the
original green size) average 5.5 percent radially and 10 percent tangen-
tially. See Table 10 for the appropriate kiln schedule. For information
about schedules for thicker material or wood that is partially air-dried,
contact the Forest Products Department at OSU.
Table 10. Kiln schedule—Tanoak 4/4, 5/4, 6/4, 8/4.
Equilibrium
1 Above 35 110 107 19.1 90 43.5 41.5

2 35 to 30 110 106 17.6 87 43.5 41.0
3 30 to 25 120 114 15.5 83 49.0 45.0
4 25 to 20 130 120 12.2 74 54.5 49.0
5 20 to 15 140 115 6.8 46 60.0 46.0
8 15 to final 160 110 3.4 21 71.0 43.5
108
Machining
Tanoak’s machinability is comparable to or better than that of the
oaks with regard to planing, shaping, boring, and mortising. As with
the oaks, tanoak sands well with a minimum or scratching or fuzzing.
Both tooling and sandpaper dull moderately quickly because of the hardness
of the wood. For optimum planing results, the recommended hook angle
is 15∞. Machine burn is possible if feed speeds are too slow.
Adhesives
If conditions are well controlled, tanoak produces glue joints of good
strength.
Finishing
Tanoak finishes well because of its uniform color (between sapwood
and heartwood) and evenly textured, fine-grained appearance. Trans-
parent stains and dyes are better than pigmented stains in retaining
more readable finishes on tanoak. Clear-coated flooring products made
from tanoak have a warm, pleasant appearance.
Durability
Tanoak is a non-durable species when subjected to conditions favor-
able to wood decay organisms. On average, round, unpeeled posts de-
cay within 4 years of being placed in the ground. Tanoak can be effec-
tively treated with wood preservatives.
Uses
Tanoak is used for flooring, furniture, truck bedding, pallets, veneer,
paneling, ties and mine timbers, pulpwood, and firewood.
Related Literature
DICKINSON, F.E., and D.R. PRESTEMON. 1965. Tanoak log grades and
lumber yield. California Agriculture Experiment Station, Berkeley,
California. California Forestry and Forest Products No. 41.
ESPENAS, L.D. 1953. The seasoning of one-inch tanoak lumber. Oregon
Forest Products Laboratory, State Board of Forestry and School of
Forestry, Corvallis, Oregon. Bulletin 3. 46 p.
HARRINGTON, T.B., R.J. PABST, and J.C. TAPPEINER II. 1994. Seasonal
physiology of Douglas-fir saplings: response to microclimate in stands
of tanoak or Pacific madrone. Forest Science 40:59-82.
HARRINGTON, T.B., J.C. TAPPEINER II, and R. WARBINGTON. 1992. Pre-
dicting crown sizes and diameter distributions of tanoak, Pacific madrone,
and giant chinkapin sprout clumps. Western Journal of Applied For-
estry 7:103-108.
109
McDONALD, P.M. 1992. Estimating seed crops of conifer and hardwood
species. Canadian Journal of Forest Research 22:832-838.
compil. USDA Forest Service, Washington, D.C. Agriculture Hand-
book 445.
McDONALD, P.M., and J.C. TAPPEINER. 1987. Silviculture, ecology, and
management of tanoak in northern California. P. 62-70 in Proceed-
ings of the Symposium on Multiple-Use Management of California’s
Hardwood Resources. T.R. Plumb and N.H. Pillsbury, tech. coords.
USDA Forest Service, Pacific Southwest Forest and Range Experiment
Station, Berkeley, California. General Technical Report PSW-100.
RAPHAEL, M.G. 1987. Wildlife-tanoak associations in Douglas-fir forest
of northwestern California. P. 183-189 in Proceedings of the sympo-
sium on Multiple-Use Management of California’s Hardwood Resources.
T.R. Plumb and N.H. Pillsbury, tech. coords. USDA Forest Service,
Pacific Southwest Forest and Range Experiment Station, Berkeley,
California. General Technical Report PSW-100.
SCHOWALTER, W.E. 1949. The suitability of tanoak (Lithocarpus densiflora)
for the production of veneer. USDA Forest Service, Forest Products
Laboratory, Madison, Wisconsin. Administrative report. 7 p. On file
with: Timber Quality Research Team, Forestry Sciences Laboratory,
Pacific Northwest Research Station, P.O. Box 3890, Portland, Or-
egon 97208-3890.
TAPPEINER, J.C., II, P.M. McDONALD. 1984. Development of tanoak
understories in conifer stands. Canadian Journal of Forest Research
14:271-277.
TAPPEINER, J.C., II, P.M. McDONALD, and T.F. HUGHES. 1986. Survival
of tanoak (Lithocarpus densiflorus) and Pacific madrone (Arbutus menziesii)
seedlings in forests of southwestern Oregon. New Forests 1:43-55.
TAPPEINER, J.C., II, P.M. McDONALD, and D.F. ROY. 1990. Tanoak. P.
417-425 in Silvics of North America. Volume 2, Hardwoods. R.M.
Burns and B.H. Honkala, coords. USDA Forest Service, Washington
D.C. Agriculture Handbook 654.
110
APPENDIX 1
Comprehensive tables
Table 1. Volume of hardwood growing stock in subregions of northern California, Oregon, and Wash-
ington.
Volume of growing stock in million cubic feet (MMCF)

California Oregon Washington Region
Total
North- South- West- North- South- Puget Olympic
Species west west Central west Total west Sound Peninsula Total
Bigleaf maple* 72 118 345 446 909 321 587 281 1189 2170
Black cottonwood* 2 1 10 50 61 66 277 52 395 458
California black oak* 345 110 21 — 131 — — — — 476
California-laurel* 204 93 — — 93 — — — — 297
Giant chinkapin* 50 45 40 1 86 — — — — 136
Oregon ash* — 20 23 38 81 46 30 27 103 184
Oregon white oak* 148 60 128 114 302 4 1 8 13 463
Pacific madrone* 716 274 13 4 291 — 45 15 60 1067
Red alder 121 814 871 1404 3089 1248 1626 1352 4226 7436
Tanoak 1438 350 — — 350 — — — — 1788
*Inventory not available for these species on federal lands in SW Oregon.

Total combined volume of these species on federal land in SW Oregon was 449 MMCF in 1976.
Sources: NW California, all ownerships, 1981-1984 inventory, Resource Bulletin PNW-131, June 1986. Oregon
non-federal land, 1985-86 survey: Resource Bulletin PNW-RB-138, September 1986; Resource Bulletin PNW-RB-
140, September 1986; Resource Bulletin PNW-RB-143, March 1987. Oregon federal land, 1976 survey: Resource
Bulletin PNW-72, March 1979; Resource Bulletin PNW-76, 1978; Resource Bulletin PNW-82, March 1979.
Washington, all ownerships, 1988-89 survey: Resource Bulletin PNW-RB-191.
111
Table 2. Green lumber recovery percentages for some northwestern and eastern hardwood species.
Average Select No. 1 Common

Log diameter and Better or Better
Species grade (in.) (%) (%)
Northwestern
Bigleaf maple 1,3 1 23 41 80
2 19 58
3 18 31
California black oak 6 1 24 27 60
2 21 35
3 15 19
Giant chinkapin 7 1 18 63
2 45
3 30
Pacific madrone 5 1 21 33 69
2 18 52
3 14 36
Red alder 1,2 1 19 26 69
2 19 36
3 17 31
Tanoak 4 1 25 29 65
2 21 52
3 19 43
Eastern
Hard maple 1 19 46 73
2 16 48
3 14 17
Red oak 1 22 46 73
2 17 51
3 16 21
Yellow birch 1 18 48 73
2 17 49
3 13 13
White oak 1 19 41 67
2 19 45
3 17 23
1
Grade recovery is based on standard NHLA grades based on the poor face, not modified grades for red alder and bigleaf
maple.
2
Pfeiffer, J.R., and A.C. Wollin. 1954. Red alder log and lumber grading. Oregon Forest Products Laboratory, Corvallis, Oregon.
Report No. G-3. 21 p.
3
Wollin, A.C., and J.R. Pfeiffer. 1955. Oregon maple log and lumber grading. Oregon Forest Products Laboratory, Corvallis,
Oregon. Report No. G-4. 21 p.
4
Dickinson, F.E., and D.R. Prestemon. 1965. Tanoak log grades and lumber yield. California Agriculture Experiment Station,
Berkeley, California. California Forestry and Forest Products No. 41.
5
Dickinson, F.E., D.R. Prestemon, and W.A. Dost. 1965. Pacific madrone log grades and lumber yield. California Agriculture
Experiment Station, Berkeley, California. California Forestry and Forest Products No. 43.
6
Malcolm, F.B. 1962. California black oak—a utilization study. USDA Forest Service, Forest Products Laboratory, Madison,
Wisconsin. Report No. 2237. 10 p.
7
Prestemon, D.R., F.E. Dickinson, and W.A. Dost. 1965. Chinkapin log grades and lumber yield. California Agriculture Experi-
ment Station, Berkeley, California. California Forestry and Forest Products No. 42.
8
Vaughn, C.L., A.C. Wollin, K.A. McDonald, and E.H. Bulgrin. 1966. Hardwood log grades for standard lumber. USDA Forest
Service, Forest Products Laboratory, Madison, Wisconsin. Research Paper FPL-63. 52 p.
112
Table 3. Average mechanical properties for northwestern (bold-face) and some eastern hardwood
species.
Side
MOR MOE COMP hardness Impact Shear SP. GR.
Species (lb/in. 2 ) (lb/in. 2) (lb/in. 2 ) (lb) bending (lb/in. 2) 12% MC
Black cottonwood 8300 1.26 X 10 6 370 350 22 1020 0.35

Aspen 8400 1.18 X 10 6 370 350 21 850 0.38
Red alder 9800 1.38 X 10 6 540 590 20 1080 0.41
Yellow-poplar 10100 1.58 X 10 6 500 540 24 1190 0.42
American chestnut 8600 1.23 X 10 6 620 540 19 1080 0.43
Silver maple 8900 1.14 X 10 6 740 700 25 1480 0.47
Bigleaf maple 10700 1.45 X 10 6 930 850 28 1730 0.48
Giant chinkapin 10700 1.24 X 10 6 680 730 30 1260 0.48
Black ash 12600 1.60 X 10 6 760 850 35 1570 0.49
Cherry 12300 1.49 X 10 6 690 950 29 1700 0.50
Black walnut 14600 1.68 X 10 6 1010 1010 34 1370 0.55
California-laurel 8000 0.94 X 10 6 1400 1270 31 1860 0.55
Oregon ash 12700 1.36 X 10 6 1540 1160 33 1790 0.55
California black oak 8700 0.99 X 10 6 1440 1100 16 1470 0.57
Southern red oak 10900 1.49 X 10 6 870 1060 26 1390 0.59
Northern red oak 14300 1.82 X 10 6 1010 1290 43 1780 0.63
Sugar maple 15800 1.83 X 10 6 1470 1450 39 2330 0.63
Eastern white oak 1 10300 1.03 X 10 6 1200 1370 29 1810 0.65
Pacific madrone 10400 1.23 X 10 6 1620 1460 23 1810 0.65
Tanoak 16300 1.80 X 10 6 1080 1410 NA 2180 0.66
Oregon white oak 10300 1.10 X 10 6 2110 1660 29 2020 0.72
MOR—modulus of rupture: strength measurement of the load required to break a test sample.
MOE—modulus of elasticity: stiffness measurement.
COMP—compression perpendicular to grain.
Impact Bending—drop distance in in. for a 50-lb hammer to break sample.
SP. GR.—specific gravity.
1
Burr oak figures from Wood Handbook (USDA Ag. Handbook #72).
113
APPENDIX 2
Common and scientific names of tree species
Bigleaf maple Acer macrophyllum Pursh

Bitter cherry Prunus emarginata (Dougl. ex Eaton)
Black cottonwood Populus trichocarpa Torr. & Gray
Black hawthorn Crataegus douglasii Lindl.
California black oak Quercus kelloggii Newb.
California-laurel Umbellularia californica (Hook. & Arn.) Nutt.
California sycamore Platanus racemosa Nutt.
Canyon live oak Quercus chrysolepis Liebm.
Douglas-fir Pseudotsuga menziesii (Mirb.) Franco
Giant chinkapin Castanopsis chrysophylla (Dougl.) A. DC.
Grand fir Abies grandis (Dougl. ex D. Don) Lindl.
Knobcone pine Pinus attenuata Lemm.
Northwest willow Salix sessilifolia Nutt.
Oregon ash Fraxinus latifolia Benth.
Oregon white oak Quercus garryana Dougl. ex Hook.
Pacific dogwood Cornus nuttallii Aud.
Pacific madrone Arbutus menziesii Pursh
Pacific willow Salix lasiandra Benth.
Ponderosa pine Pinus ponderosa Dougl. ex Laws.
Port-Orford-cedar Chamaecyparis lawsoniana (A. Murr.) Parl.
Red alder Alnus rubra Bong.
Redwood Sequoia sempervirens (D. Don) Endl.
River willow Salix fluviatilis Nutt.
Scouler willow Salix scoulerana Barratt ex Hook.
Sitka spruce Picea sitchensis (Bong.) Carr.
Sugar pine Pinus lambertiana Dougl.
Tanoak Lithocarpus densiflorus (Hook. & Arn.) Rehd.
Western hemlock Tsuga heterophylla (Raf.) Sarg.
Western redcedar Thuja plicata Donn ex D. Don
Western white pine Pinus monticola Dougl. ex D. Don
White fir Abies concolor (Gord. & Glend.) Lindl. ex Hildebr.
114
APPENDIX 3
Common and scientific names of shrub species
Bearclover Chamaebatia foliolosa Benth.

Bearberry Arctostaphylos uva-ursi (L.) Spreng.
Blue elderberry Sambucus caerulea Raf.
Blueblossom Ceanothus thyrsiflorus Eschsch.
Buckbrush Ceanothus cuneatus (Hook.) Nutt. ex Torr. & A.
Gray
Ceanothus Ceanothus spp.
Common snowberry Symphoricarpos albus (L.) S.F. Blake
Creeping snowberry Gaultheria hispidula (L.) Muhl.
Deerbrush Ceanothus integerrimus Hook. & Arn.
Devil’s-club Oplopanax horridum (Sm.) Miq.
Evergreen blackberry Rubus laciniatus Willd.
Evergreen huckleberry Vaccinium ovatum Pursh.
Greenleaf manzanita Arctostaphylos patula Greene
Hairy manzanita Arctostaphylos columbiana Piper
Hazel Corylus cornuta Marsh.
Himalaya blackberry Rubus procerus P.J. Mull.
Mountain-lover Pachistima canbyi Gray
Oceanspray Holodiscus discolor (Pursh.) Maxim.
Oregon crab apple Malus fusca (Raf.) C.K. Schneid.
Oregon-grape Berberis aquifolium Pursh
Osoberry Oemleria cerasiformus (Hook. & Arn.) J.W. Landon
Pacific dewberry Rubus ursinus Cham. & Schlechtend.
Pacific rhododendron Rhododendron macropyllum D. Don ex G. Don
Red elderberry Sambucus callicarpa Greene
Red-flowered currant Ribes sanguineum Pursh.
Red huckleberry Vaccinium parvifolium Sm.
Red osier dogwood Cornus sericea L.
Salal Galtheria shallon Pursh.
Salmonberry Rubus spectabilis Pursh.
Snowbrush Ceanothus velutinus Dougl. ex Hook.
Sweet mock-orange Philadelphus coronarius L.
Thimbleberry Rubus parviflorus Nutt.
Vine maple Acer circinatum Pursh
Wax myrtle Myrica cerifera L.
Western poison-oak Rhus diversiloba Torr. & A. Gray
Western raspberry Rubus leucodermis Dougl. ex Torr. & A. Gray
Western serviceberry Amelanchier alnifolia (Nutt.) Nutt.
115
Western whipplea Whipplea modesta Torr.
Whiteleaf manzanita Arctostaphylos viscida Parry
Whortleberry Vaccinium corymbosum L.
Wood rose Rosa gymnocarpa Nutt.
116
APPENDIX 4
Common and scientific names of herb species
Angelica Angelica spp.

Bedstraw Galium spp.
Bittercress Cardamine spp.
Bull thistle Cirsium vulgare (Savi) Ten.
Buttercup Ranunculus spp.
California brome Bromus carinatus Hook. & Arn.
California fescue Festuca californica Vasey
California sweetgrass Hierochloe occidentalis Buckley
Canada violet Viola canadensis L.
Enchanter’s-nightshade Circaea alpina L.
False lily-of-the-valley Maianthemum canadense Desf.
False Solomon’s-seal Smilacina stellata (L.) Desf.
Golden-saxifrage Chrysosplenium L.
Hedge nettle Stachys L.
Honeysuckle Lonicera L.
Horsetail Equisetum spp.
Iris Iris spp.
Ladyfern Athyrium felix-femina (L.) Roth
Maidenhair fern Adiantum pedatum L.
New Zealand fireweed Erechtites arguta DC.
Prince’s-pine Chimaphila umbellata (L.) Bart.
Red woodsorrel Oxalis oregana Nutt.
Sedges Carex spp.
Skunkcabbage Lysichiton americanum Hult & St. John
Spiraea Spiraea spp.
Spreading sweetroot Osmorhiza chilensis (Molina) Hook. & Arn.
Stinging nettle Urtica dioica L.
Twinflower Linnaea borealis L.
Water celery Oenanthe sarmentosa K. Presl. ex DC.
Wild strawberry Fragaria virginiana Duchesne
Western swordfern Polystichum munitum (Kaulf.) K. Presl.
Western bracken Pteridium aquilinum L.
Western springbeauty Montia sibirica (L.) J.T. Howell
Youth-on-age Tolmiea menziesii (Pursh) Torr. & A. Gray
117
118
Niemiec, S.S., G.R. Ahrens, S. Willits, and D.E. Hibbs. 1995.
HARDWOODS OF THE PACIFIC NORTHWEST. Forest Research
Laboratory, Oregon State University, Corvallis. Research Con-
tribution 8. 115 p.
This publication brings together in one place information on the
general characteristics, biology and management, harvesting and
utilization, wood characteristics, and related literature of Pacific
Northwest hardwoods. Species included are bigleaf maple, black
cottonwood, California black oak, California-laurel, giant chinkapin,
Oregon ash, Oregon white oak, Pacific madrone, red alder, and
tanoak.
Niemiec, S.S., G.R. Ahrens, S. Willits, and D.E. Hibbs. 1995.

HARDWOODS OF THE PACIFIC NORTHWEST. Forest Research
Laboratory, Oregon State University, Corvallis. Research Con-
tribution 8. 115 p.
This publication brings together in one place information on the
general characteristics, biology and management, harvesting and
utilization, wood characteristics, and related literature of Pacific
Northwest hardwoods. Species included are bigleaf maple, black
cottonwood, California black oak, California-laurel, giant chinkapin,
Oregon ash, Oregon white oak, Pacific madrone, red alder, and
tanoak.
119
As an affirmative action institution that complies with Section 504 of the
Rehabilitation Act of 1973, Oregon State University supports equal educa-
tional and employment opportunity without regard to age, sex, race,
creed, national origin, handicap, marital status, or religion.
Non-Profit Org.
U.S. Postage
Oregon State University PAID
Forest Research Laboratory 227 Corvallis, OR
Corvallis, OR 97331-7401 Permit No. 200
Address Correction Requested

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Research Contribution 8 March 1995

HARDWOODS OF THE PACIFIC NORTHWEST

College of Forest Research Laboratory

Address Correction Requested

The history of hardwoods in the Pacific Northwest is sometimes char-

Common Name Scientific Name

Bigleaf maple Acer macrophyllum Pursh

In each chapter, what is and is not known about a given spe-

This chapter provides information that is general to most species

Size, Longevity, and Form

Biology and Management

Suitability and Productivity of Sites

Flowering and Fruiting

Regeneration from Seed

Regeneration from Vegetative Sprouts

Regeneration from Planting

Growth and Yield

Interactions with Wildlife

Insects and Diseases

Harvesting and Utilization

Drying and Shrinkage

BOLSINGER, C.L. 1987. Major findings of a statewide resource assess-

Bigleaf maple is second to red alder among native hardwood species

Size, Longevity, and Form

Biology and Management

Tolerance, Crown Position

• Top height of existing mature trees is 80 to 100 ft

Flowering and Fruiting

Natural Regeneration from Seed

Regeneration from Vegetative Sprouts

Regeneration from Planting

Site Preparation and Vegetation Management

Growth and Yield

Interactions with Wildlife

Insects and Diseases

Harvesting and Utilization

Cruising and Harvesting

Table 1. Kiln schedule—Bigleaf maple 4/4, 5/4, 6/4.

1 Above 50 130 123 14.3 81 54.5 50.5

BASTENDORFF, K.M., and A. POLENSEK. 1984. Strength and stiffness of

Black cottonwood is a large deciduous tree belonging to the willow

Size, Longevity, and Form

Biology and Management

Tolerance, Crown Position

Suitability and Productivity of Sites

Flowering and Fruiting

Regeneration from Vegetative Sprouts

Regeneration from Planting

Site Preparation and Vegetation Management

Growth and Yield

Interactions with Wildlife

Insects and Diseases

Cruising and Harvesting

Drying and Shrinkage

Table 2. Kiln schedule—Black cottonwood* 4/4, 5/4, 6/4.

1 above 70 140 130 12.0 75 60.0 54.5

BASSMAN, J.H., and J.C. ZWIER. 1993. Effect of partial defoliation on

California black oak is a deciduous member of the beech family

Size, Longevity, and Form