A B C D E F

1 Population Viability Analysis

2
3 N(0) Mean l Std (l) l value p(extinction)
4 10 1 0.5 0.1
5
6 Trial l value
7 1 1.71781786 2.107835572 0.6458227109 1.3039043793
8 2 1.99081361 0.324063136 0.5539065924 1.3278894243
9 3 1.25595439 1.500209565 1.1035663813 0.531965074
10 4 1.65197438 0.819763457 0.6348604751 0.5808430978
11 5 1.06233036 0.855713374 0.8375107391 1.4423393836
12 6 1.15108725 1.859386806 1.8068115687 0.0257991639
13 7 0.34192528 0.570326534 0.8277694574 0.531760128
14 8 0.10138691 0.591479753 0.9096569375 1.462275664
15 9 0.77036687 1.156269798 0.1783187431 0.9712888188
16 10 0.71682946 0.664504792 1.1472221847 1.2483579265
17
18 Year
19 Trial 0 1 2 3 4
20 1 10 17 36 23 30
21 2 10 20 6 3 4
22 3 10 13 20 22 12
23 4 10 17 14 9 5
24 5 10 11 9 8 12
25 6 10 12 22 40 1
26 7 10 3 2 2 1
27 8 10 1 0 0 0
28 9 10 8 9 2 2
29 10 10 7 5 6 7
30
31
32
33
34
35 Projection of 10 Population Simulations with Mean Lambda=1 and S
36 45
37
40
38
35
39
30
40
25
Population Size

41
42 20
15
10
5
0
0 0.5 1 1.5 2 2.5 3 3.5 4
 40
35
30
25

Population Size
A B C D20 E F
43 15
44 10
45 5
46 0
47 0 0.5 1 1.5 2 2.5 3 3.5 4
48 Year
49
 G H I J
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16 100 years, 100 trials
17 s p(extinction)
18 Persist? 0
19 0 = Yes, 1 = No 0.1
20 1 0.2
21 1 0.3
22 1 0.4
23 1 0.5
24 1 0.6
25 1 0.7
26 1 0.8
27 0 0.9
28 1 1
29 1
30 0.9
31
32
33
34
ations with Mean Lambda=1 and Standard Deviation=0.5
35
36
37 Row 20
38 Row 21
39 Row 22
40 Row 23
41 Row 24
42 Row 25
Row 26
Row 27
Row 28
Row 29

.5 2 2.5 3 3.5 4 4.5

 Row 20
Row 21
Row 22
Row 23
Row 24
Row 25
G H I J
Row 26
43
Row 27
44
Row 28
45 Row 29
46
.5 2 2.5
47 3 3.5 4 4.5
Year 48
49
Population Viability Exercise
Spreadsheet template for 100 years and 100 trials

N (0) mean l std (l) l deviate p (extinction)

600 0.8 0.25 1.0528910864
l Deviates
Trial 1 2 3 4 5
1 0.82008883 0.709369 0.7055119366 0.9295442805 0.8767076469
2 0.33690044 1.0020956 1.1034836646 0.7196154513 0.8004296571
3 0.79527352 0.79917525 0.7103010683 0.6704462554 0.9437828891
4 0.49308942 0.50687689 0.4719687965 0.5916928246 0.386317179
5 1.01468503 0.7067314 0.5058499068 1.207214392 0.9414536261
6 0.72704288 1.30551604 0.6024029515 1.1501891967 0.9671189057
7 0.55923975 0.97266223 0.726395283 0.5277884088 0.8868115501
8 0.77985678 0.86602635 0.9529728232 1.3077849107 1.1723215735
9 0.76545465 1.33165011 0.6451738467 0.7731350017 0.4634119599
10 0.88784 0.83047888 0.7590360727 0.3243353568 0.9454040999
11 0.956591 0.80667719 0.480049328 0.6127027725 1.5089156955
12 0.77986787 0.89010545 0.8889682018 0.632984896 0.9034208775
13 0.7065181 0.72224949 0.7245808402 0.5087141531 0.9582715399
14 0.25664265 0.87503515 0.4890212173 0.6855959981 0.9052804986
15 0.50874735 1.28001504 0.7423996269 1.0968012509 0.9322407936
16 0.93601513 1.16508076 0.893405124 1.1196425057 0.9608710507
17 0.4178222 0.47512488 0.6196122242 0.1753391207 1.1555428539
18 1.34199025 0.81881112 0.3899564627 0.5364305804 0.8460027797
19 0.90830333 0.93753071 0.8254836668 0.9622796849 0.9561582196
20 0.67057511 0.87686659 0.4763622558 0.7015715701 1.0383498025
21 0.80019483 0.45969249 0.4025559432 0.5476638048 1.0664173317
22 0.97137532 0.93098366 0.8829735872 0.6794944195 0.8248762108
23 0.82995986 0.38881056 1.2467738416 0.492608179 0.7313088083
24 0.83594179 1.14043667 0.8241092025 0.4075706592 1.0776656318
25 0.36972286 0.74880072 0.4755572815 0.3443884794 0.7925774758
26 0.9991391 0.87393434 0.876920184 1.1347227868 1.1161537121
27 0.6772168 0.70463722 0.5230557708 0.8397707565 0.723833545
28 0.69166486 0.34874765 1.1107295286 1.3560130324 0.7155368755
29 0.73655327 0.73312057 1.1014253554 1.1713074333 1.0645117375
30 0.78493975 0.77689772 0.8544617943 1.0740029557 0.7434597669
31 0.67403981 0.97998656 1.0722331115 1.1541615389 0.6697280742
32 0.80820206 0.78037244 0.3582573735 0.8095640128 1.0367904976
33 1.18641696 0.90850748 0.346469159 0.8369431266 0.5190258704
34 1.04709048 0.90635108 1.0595413393 0.8190122969 0.8361884879
35 0.86211789 0.57230949 1.1318824533 0.4397339104 0.2958073696
36 1.02005699 0.94237781 0.8479789048 0.4826978981 0.298776964
37 0.5425835 0.94776268 1.048382094 1.0313717848 1.0551707052
38 0.92419241 0.9269937 0.7067350751 0.582594378 0.9135678403
39 1.0108557 0.48682792 0.5993915011 1.0343247854 0.8706200129
40 0.79143168 0.81949196 0.7353575019 0.9673018286 0.487993144
41 0.84376367 1.0266005 0.7937949836 0.825798474 0.9897694509
42 0.70256147 1.0027751 0.6204400781 1.1354337444 0.9871598647
43 0.4209137 1.01387646 0.926756218 0.5884655591 1.1353587803
44 0.79139417 1.1500216 0.7043079592 0.6416278474 0.6998980405
45 0.83784033 1.01645264 0.5758462256 0.5501840565 0.5856471926
46 1.11096207 0.77187103 0.8421086081 0.2659013443 0.6278968593
47 0.79059739 0.833621 0.8142403856 0.531505728 1.0202518826
48 1.0192127 0.83154541 1.0011896903 0.2354036072 1.0084621427
49 1.11228624 0.78062217 0.8514118634 0.8062228432 0.5994830286
50 0.42218751 0.79203153 0.8092863523 0.7768447794 0.8164753237
51 0.70158817 0.74354003 1.1015805633 1.1490720332 0.7677702138
52 0.54003868 0.61988593 0.7058389036 1.410100529 0.8797147863
53 0.52088879 0.82242089 1.0137062198 0.6596674111 0.9251774739
54 0.83111739 0.44246755 0.3884386267 1.0092192951 0.6910916021
55 0.71637878 0.59096233 0.5711862392 0.6537910519 0.8290854317
56 0.49766171 0.80100292 0.7103128352 1.3280229326 0.9311882219
57 1.09002164 0.99976171 0.5090600997 0.6153474808 0.7118763906
58 0.4678969 0.93855842 0.7263875289 0.7076720447 0.5690790542
59 0.96922334 0.80734419 0.7443000991 0.5390990684 0.8835344867
60 0.50598975 1.2165946 0.6740045304 0.7205611898 0.8614885186
61 0.4875623 0.86191763 0.5587303135 0.9900031841 1.3224883439
62 0.91669898 1.05831765 0.6518359626 0.813775196 0.8542795363
63 0.84707204 1.09339293 0.7084132664 0.8935753803 0.28560478
64 0.97660025 0.76585553 0.853482791 0.6340586634 0.9300024176
65 0.74021063 0.63187571 1.0921581281 0.4209619251 0.8065723052
 66 0.5345673 0.77394238 0.5499116689 0.617486696 0.7423177613
67 0.3598484 0.57298673 0.6611056352 1.0075226284 0.5245196561
68 0.66032094 0.64319276 1.1043238142 0.5894087194 0.8385558022
69 0.84978725 0.78776921 0.9391109936 1.3337104488 0.9966282589
70 1.07345562 0.7712029 0.6387614279 1.019697205 0.7590577654
71 0.776114 0.76149664 0.5549897155 0.4873035165 0.9710917862
72 0.73726211 0.95882058 1.1698350628 0.4705059763 1.1361130224
73 0.60913006 0.84464503 0.9388186788 0.5652687858 0.5582974979
74 0.86317347 1.03966412 0.6273223784 0.6071014545 0.7959732073
75 0.15781113 0.77114237 0.8640139011 0.4492395863 0.8577952233
76 0.68449267 0.86710425 0.4590581106 0.7948122428 0.5795037919
77 0.7838754 1.3479348 1.0561257509 1.0282354518 0.3591150957
78 0.70031421 1.01576068 1.1993397944 0.867297727 1.0188286182
79 0.7863827 1.16955701 1.0465805696 0.9283426241 0.6304395077
80 0.57806586 0.2158867 0.5926224811 1.0902256945 0.4986013829
81 0.21034797 0.8348195 0.4433452715 0.5712203992 0.8619884956
82 1.2527497 0.76457969 0.6803041592 1.0176691856 0.8410341311
83 0.73011236 0.64321972 0.6411654598 1.007559032 0.6020461688
84 0.68448832 0.72861877 1.4818607669 0.9428790301 0.8016075394
85 0.58936872 0.84964939 1.1347862105 0.7245508279 1.0441700982
86 0.8189554 0.98072562 0.8507171537 1.1413238349 1.1187989753
87 0.6865987 0.77992862 0.4278406001 1.1793821908 0.6441944015
88 0.75810058 0.54386337 0.7971088975 0.5050734142 0.7952420859
89 0.53930927 0.7877821 0.8063919835 1.0076173951 0.812760758
90 0.58197283 0.43596116 0.8529487471 0.7365117465 0.9219364631
91 1.00204031 0.90812104 0.7286907746 0.9351722992 1.2025954087
92 0.65440162 0.91526626 0.4970611932 0.7163708308 0.3388143245
93 0.66125944 0.71405119 0.6616770531 0.4476581837 0.7254177383
94 0.48803007 0.95844964 0.8174752144 0.4247530796 0.6218963437
95 0.9225691 0.9382513 0.7377555215 0.6247719312 0.9100302245
96 1.1015116 0.67298235 0.4987968165 0.9848360904 0.9320179709
97 0.83583022 0.91164519 0.7735460922 0.8710298397 0.1634885954
98 0.71885487 0.48485342 0.8116774301 0.9365413539 0.9185758949
99 1.04365191 1.02542397 1.0351552236 0.8071757933 0.4427280249
100 0.84355786 0.42859918 1.0204652936 0.7444735867 1.0007624202

Year
Trial 0 1 2 3 4 5
1 600 492 349 246 229 201
2 600 202 202 223 160 128
3 600 477 381 271 182 172
 4 600 296 150 71 42 16
5 600 609 430 218 263 248
6 600 436 569 343 395 382
7 600 336 327 238 126 112
8 600 468 405 386 505 592
9 600 459 611 394 305 141
10 600 533 443 336 109 103
11 600 574 463 222 136 205
12 600 468 417 371 235 212
13 600 424 306 222 113 108
14 600 154 135 66 45 41
15 600 305 390 290 318 296
16 600 562 655 585 655 629
17 600 251 119 74 13 15
18 600 805 659 257 138 117
19 600 545 511 422 406 388
20 600 402 353 168 118 123
21 600 480 221 89 49 52
22 600 583 543 479 325 268
23 600 498 194 242 119 87
24 600 502 572 471 192 207
25 600 222 166 79 27 21
26 600 599 523 459 521 582
27 600 406 286 150 126 91
28 600 415 145 161 218 156
29 600 442 324 357 418 445
30 600 471 366 313 336 250
31 600 404 396 425 491 329
32 600 485 378 135 109 113
33 600 712 647 224 187 97
34 600 628 569 603 494 413
35 600 517 296 335 147 43
36 600 612 577 489 236 71
37 600 326 309 324 334 352
38 600 555 514 363 211 193
39 600 607 296 177 183 159
40 600 475 389 286 277 135
41 600 506 519 412 340 337
42 600 422 423 262 297 293
43 600 253 257 238 140 159
44 600 475 546 385 247 173
45 600 503 511 294 162 95
46 600 667 515 434 115 72
47 600 474 395 322 171 174
48 600 612 509 510 120 121
49 600 667 521 444 358 215
50 600 253 200 162 126 103
51 600 421 313 345 396 304
52 600 324 201 142 200 176
53 600 313 257 261 172 159
54 600 499 221 86 87 60
55 600 430 254 145 95 79
56 600 299 239 170 226 210
57 600 654 654 333 205 146
58 600 281 264 192 136 77
59 600 582 470 350 189 167
60 600 304 370 249 179 154
61 600 293 253 141 140 185
62 600 550 582 379 308 263
63 600 508 555 393 351 100
64 600 586 449 383 243 226
65 600 444 281 307 129 104
66 600 321 248 136 84 62
67 600 216 124 82 83 44
68 600 396 255 282 166 139
69 600 510 402 378 504 502
70 600 644 497 317 323 245
71 600 466 355 197 96 93
72 600 442 424 496 233 265
73 600 365 308 289 163 91
74 600 518 539 338 205 163
75 600 95 73 63 28 24
76 600 411 356 163 130 75
77 600 470 634 670 689 247
78 600 420 427 512 444 452
79 600 472 552 578 537 339
80 600 347 75 44 48 24
81 600 126 105 47 27 23
82 600 752 575 391 398 335
83 600 438 282 181 182 110
84 600 411 299 443 418 335
85 600 354 301 342 248 259
 86 600 491 482 410 468 524
87 600 412 321 137 162 104
88 600 455 247 197 99 79
89 600 324 255 206 208 169
90 600 349 152 130 96 89
91 600 601 546 398 372 447
92 600 393 360 179 128 43
93 600 397 283 187 84 61
94 600 293 281 230 98 61
95 600 554 520 384 240 218
96 600 661 445 222 219 204
97 600 501 457 354 308 50
98 600 431 209 170 159 146
99 600 626 642 665 537 238
100 600 506 217 221 165 165
 100 years, 100 trials 100 years, 100 trials
s (lambda) p(extinction) Population Size
0 100
0.1 200
0.2 300
0.3 400
0.4 500
0.5 600
0.6 700
0.7 800
0.8 900
0.9 1000
1
# persistence #extinctions p(extinction)

6 7 8 9 10 11 12
0.5855369622 0.5495680416 0.8106874513 1.012209 1.0694928434 0.620965642 0.8244262892
0.8117753151 1.4099026375 0.5198544402 1.024064 0.9383142182 0.4905319404 0.7073851995
1.2563738711 0.6720130005 0.8958475188 1.017162 0.7619722767 0.8985493621 1.094714245
1.147826296 1.0220391959 0.6184251573 0.908629 0.8716212127 1.0473867278 0.763740358
0.9577010038 0.6367136195 1.1177676267 0.870368 0.7393285848 0.4583135909 1.1292639467
0.9778570806 0.3383575885 0.6231637935 1.190595 0.7254109839 1.1900927482 0.5156030194
1.0512976048 0.5308983811 0.4934325679 1.056955 0.6615027578 0.831727672 0.9972157569
0.6438420031 1.3669424145 0.9828480747 0.589798 0.7276984096 0.690294885 0.3699693038
0.7096969323 1.1300982388 0.6980551298 0.574194 0.5731256613 1.1322380441 0.6358020639
0.2503700814 0.690293711 0.8822948042 0.584452 0.422205978 0.5099381632 0.8006319749
0.8339017738 0.3660543704 0.5210683564 0.769491 1.1038655384 1.0014704346 1.1459437995
0.8398938521 1.0433842678 1.1403772524 0.804015 0.7160042884 0.7987422745 0.7306134064
0.8993297308 1.0076385192 0.591169442 0.796021 0.3132668498 0.5394181213 0.5476269843
0.518737504 0.6880807283 1.153903916 0.769028 0.7428848714 0.8695569077 0.6714529001
0.8390839995 1.2092018492 1.2644897847 0.252942 1.4138725496 0.5489861441 0.9657043784
0.7258969047 0.9932413971 1.0553360292 0.82241 0.9561083814 0.9534158058 1.1852638923
0.7292562912 0.6166807346 0.64682764 0.610833 0.8734116868 0.6046404533 0.46381962
0.7129644839 0.5853057662 1.1250529375 0.346056 0.3900794079 1.0301188601 0.9364967269
0.5969543693 1.0815531862 1.1020862093 0.851638 1.2864828462 1.0704061106 1.1346054949
1.0204222594 0.7183267885 0.7077235837 0.77891 1.1860975637 0.5829354789 0.7721951697
0.8659422559 1.0852321111 0.9767825881 1.183347 0.4573466441 0.7442927825 0.6890003387
0.9549382822 0.672307566 0.5125172426 0.514243 0.841430971 1.1360545309 0.4958438131
0.6356685362 0.8199656885 0.355225722 0.82298 0.8982515061 0.5197586051 1.3906007136
0.6705797898 0.8292714779 1.3766076446 0.467658 0.539930921 0.7623735315 0.5548151855
0.7163260139 1.1286524967 0.8604451629 0.771522 0.7862081742 0.6633013794 1.0322915404
0.6136673222 1.0114374046 1.2065222546 0.432508 0.5894710324 0.8808966032 0.7448525193
1.2178613748 0.9086370707 0.7089239276 0.827925 0.7833005126 1.1693167306 1.0509362983
0.7099766345 0.5816293843 0.9833915023 1.295847 0.7502636604 1.0477435897 0.9772463064
0.6902237741 0.7811705246 0.2897295443 0.949981 0.4984013715 0.7995366288 0.1157723297
0.6637471672 1.0005727444 1.0217889794 1.121892 0.6099120412 0.9584600012 0.7680916387
0.7490899693 0.4964651486 0.8654996397 0.771342 0.7897994486 0.5576048696 0.7164113527
1.6272822264 0.4457862355 0.9778465303 0.839474 0.9812424334 0.7883789636 0.9961662693
0.8537495764 0.6224775948 0.1064323166 0.782952 0.8241455122 0.8613209574 0.6766520407
0.6554262646 0.7122190637 0.4916061961 0.980219 0.6562907976 1.1530260775 0.8357138903
0.7004933898 1.1978748627 1.1342932059 0.435264 1.0140944052 0.650476118 0.9585622467
0.8858090267 0.7336764786 0.8801579571 0.94668 0.6007969222 0.7712413262 0.7720782646
0.7027172001 1.0743302523 0.4667786736 1.096321 1.3055943245 1.5254424021 0.8352686172
0.8331978252 0.5701125714 0.3736449851 0.915623 0.5925631673 0.4155990524 1.029509212
0.8256446471 0.8779722186 0.3386136328 1.178774 1.1171680896 0.8498797521 0.2973223546
0.9199527108 0.8444183938 0.8384844251 0.581218 1.021341181 0.6541377838 0.8160852802
1.092440161 1.0413840517 0.8882383288 0.645669 0.9962882375 0.7484984931 0.7629320248
0.9563076063 1.038801566 0.6382723995 0.842324 0.5945069819 0.7321243176 0.7055303852
0.5095923499 0.6490132281 0.4691580752 0.661556 0.8970313159 0.7300533064 0.9448396603
0.9761656744 0.1613183392 0.8374825977 0.853599 0.9546906217 0.7873677628 0.6839898331
0.6750865558 0.8121861751 1.1017203433 0.815679 0.5222650533 1.0317323668 0.962802692
0.8259353784 0.4746435332 0.8863736232 0.703033 0.5559689975 1.0202930116 1.3508693817
1.1123584189 0.8071450276 0.8740414722 1.18482 0.6751982531 0.7609206325 0.5248335145
0.9402670319 0.6637969111 1.3787507864 0.904191 1.3201585392 0.6646529215 0.6248451554
0.4517469821 1.0242134068 0.7735831167 0.601402 0.9390336521 0.4070113874 0.8516264138
0.7866764217 0.4284031192 0.6265097847 0.823822 0.5880826528 0.5242399458 1.068835634
0.7673019728 0.796795698 0.425805858 1.144039 0.8230838481 0.5050085764 0.9242626202
1.0458353695 0.6995244667 0.754253222 1.082725 1.1876444043 0.8478454155 0.5764456069
0.7466486193 0.5416015627 0.6494705467 0.458437 0.5439406528 1.1886390317 1.0366153049
0.3883572613 0.8757636973 0.7837548056 0.868221 0.6999055487 0.7961267087 0.7511892477
0.8721766987 0.6971466855 0.9074677189 0.820292 0.9865267871 0.6834017078 0.6924005652
1.1032080303 0.7913525865 0.9057906104 0.711344 1.0073416643 0.2606447958 0.9207036333
1.2641415915 0.868966854 0.9657795915 0.54644 0.6524487919 1.4197397715 0.8395698357
1.1840460334 0.9062880445 0.6759653698 0.51929 0.6185639751 0.7596277107 0.8933684538
0.5993238823 0.7703990488 0.518931034 0.905465 0.7026006952 1.1286227585 1.0075993398
0.9023003715 0.6307377972 0.8572993699 0.59246 0.6560639648 0.9556069455 0.7798211143
0.833118188 0.9361318177 0.8456264256 0.814318 0.7244601007 0.1939292368 1.17301191
0.7673458858 0.8723053945 0.6065559044 0.306048 0.8509753387 0.9931637119 1.2730636361
1.1084975441 0.8168779865 0.5130366866 0.909503 0.8441659416 0.8833976775 0.8357712105
0.9560816708 1.3895252777 0.4244863187 0.978221 0.9153468972 0.9864209204 0.8401840874
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82 83 84 85 86 87 88
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