Y-DNA Haplogroup M and its Subclades -

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Version History Last revision date for this specific

Because of continuing research, the structure of the Y-DNA Haplogroup Tree changes amd will be updated
If the differences need clarification or if you find any broken links on this page, then:

LINKS: Glossary Main Page Y-DNA Tree Trunk SNP Index

FONT COLORS: Newly confirmed in 2019 within subclade Confirmed within subclade

SYMBOLS: ^ Indicates a next-generation sequencing entry which does not yet meet quality guidelines fo
^^ Indicates an entry which does not meet quality guidelines but may be helpful.
~ Indicates only an approximate location on the tree.
The criteria for a representative SNP printed in bold for a subclade is: traditional usage, testing done in mu
found listed
SNPs in the below
area ofinthe chromosome
italics used in
(colored black or recent research
red) are qualitystudies.
variants from next-generation sequencing rep
representing
situation, that subgroup
evidence seems to indicate they also seem to belong to the same subgroup, but some confirmatory
available.

M SK1828, Page93/S322
Z31169, Z31170, Z31171, Z31172, Z31173, Z31174, Z31175, Z31181, Z31182, Z31183, Z31184, Z311
M1 Z31194, Z31195, Z31196,M186,
Z31197, Z31198, Z33312, Z33344
FGC13408.2,/Z16369.2, M5991, Z30911, Z30929, Z30934.1, Z30945, Z30949, Z30955, Z3096
M1~ Z31097, Z31100, Z31140, Z31148, Z31164
M1a Z30983, Z30997, Z31022, Z31030, Z31069
M1a1 SK1846/Z43400, Z30917
BY8413.1/Z30938.1, SK1869, Z30835, Z30
M1a1a Z31079, Z31151, Z31158
M1a1a1~ P94
M1a1a2 Z30928, Z31007, Z31146
M1a1a2a
 M1a1a3~ Z33330, Z33345, Z42288
M1a1a3a~
M1a1a3b
M1a1a3b~

M1a2~ Z43411, Z43412, Z43413, Z43414, Z43415

M1b~ FT254675
M2 M353, M387
M2~ PF4350.1
M2a M177^^/SRY9138^^
M3 P117

MUTATION INFORMATION AT VERY BOTTOM HERE

NOTES:

Identical SNPs that were discovered separately are listed in alphabetical order, not necessarily in the order of discovery,
and separated by "/". Examples: M5/P73, M104/P22, M177, SRY9138
Removed P35_1, P35_2, P35_3, not meeting quality guidelines on 11 August 2016.
The chromosome location for M-P256 shown in Karafet (2008) seems off by one chromosome position.
All M samples from next-generation sequencing show the same mutation at adjacent chromosome positio
So this is the position tentatively listed in the ISOGG index. Listed 31 August 2016.

Y-DNA haplogroup M reaches its known peak in Papua New Guinea, totaling one-third to two-thirds of population.
The haplogroup M Aborigines in Australia seem comparatively to be the most recent arrivals among Abori

Geographic distribution so far of ancestry of modern-day haplogroup M subgroup men (not complete):
M1a1a2a2a M16 Papua New Guinea
M1a1a2a2b M83 Papua New Guinea
M1a1a3b1b Z42297 N. Australia (Aborigines)
M1a1a3b1c B269 Papua New Guinea (Kosibes)
M1a1a3b1d~ B272 Papua New Guinea (Kosibes)
M1a1a3b2 Y26293/Z43837 Papua New Guinea
M1a2 Z43411 Papua New Guinea
M1b FT254675 Indonesia (Maluku Isl.)
M2 M177 Papua New Guinea
M3 P117 Melanesia

References:

Bergstrom et al, Deep Roots for Aboriginal Australian Y Chromosomes. (pdf) Current Biology, 26:809-813, (2016).
Bergstrom et al, A Neolithic Expansion, but Strong Genetic Structure, in the Independent History of New Guinea Sc
Cox M P & Lahr M M, Y-Chromosome Diversity Is Inversely Associated with Language Affiliation in Paired Austr
Hurles et al, European Y-Chromosomal Lineages in Polynesians: A Contrast to the Population Structure Revealed by
Hurles et al, Y Chromosomal Evidence for the Origins of Oceanic-Speaking Peoples. Genetics, 160:289-303, Januar
Karmin et al, A Recent Bottleneck of Y chromosome Diversity Coincides with a Global Change in Culture. Genome
Karafet et al, New Binary Polymorphisms Reshape and Increase Resolution of the Human Y-Chromosomal Haplogr
Kayser et al, The Impact of the Austronesian Expansion: Evidence from mtDNa and Y Chromosome Diversity in the
Kayser et al, Independent Histories of Human Y Chromosomes from Melanesia and Australia. American Journal of H
Kayser et al, Melanesian and Asian Origins of Polynesians: mtDNA and Y-Chromosome Gradients across the Pacifi
Kayser et al, Melanesian Origin of Polynesian Y Chromosomes. Current Biology, Vol. 10, No. 20, September 2000.
Kayser et al. Reduced Y-Chromosome, but Not Mitochondrial DNA, Diversity in Human Populations from West Ne
Mona et al, Patterns of Y-chromosome Diversity Intersect with the Trans-New Guinea Hypothesis. Mol Biol Evol. 2
Rozen et al, Remarkably Little Variation in Proteins Encoded by the Y Chromosome's Single-Copy Genes, Implying
Scheinfeldt et al, Unexpected NRY Chromosome Variation in Northern Island Melanesia. Society for Molecular Bio

Additional Resources:
ISOGG Wiki What you need to know about Genetic Genealogy.

Corrections/Additions made since 1 January 2019:

None, but many mutations in brown upgraded to confirmed status on 10 April 2020.

Back to
Main Page
Y-DNA Tree Trunk
SNP Index
 Papers/Prentations Cited
Glossary
Listing Criteria

Copyright 2020, International Society of Genetic Genealogy. All Rights Reserved.

MUTATION INFORMATION

If copying this section just below, choose Paste Special, then Values, otherwise the haplogroup contents will not co
Name Haplogroup Other Names rs # Build 37 # Build 38 #
SK1828 M 8685231 8817190
Page93 M S322 rs34486382 15481435 13369555
S322 M Page93 rs34486382 15481435 13369555
M4 M1 rs3895 2744628 2876587
CTS7357.2 M1 rs763267999 17437224 15325344
FGC29215 M1 10007406 10169797
M5 M1 P73 rs3896 21609946 19448060
P73 M1 M5 rs3896 21609946 19448060
M106 M1 rs2032611 21866424 19704538
M189 M1 rs2032606 14945705 12833779
M296 M1 rs9341319 22745121 20583235
Page1 M1 rs34820858 6736812 6868771
PH4279 M1 rs765015214 19152655 17040775
PH3025 M1 rs776850039 17392059 15280179
SK1828 M1 8685231 8817190
SK1845 M1 8630327 8762286
SK1847 M1 21685085 19523199
SK1848 M1 rs972502323 6940886 7072845
SK1849 M1 18713579 16601699
SK1850 M1 19165242 17053362
SK1851 M1 rs1466997937 16174669 14062789
SK1852 M1 14945506 12833580
SK1853 M1 17077435 14965555
SK1854 M1 17844264 15732384
SK1855 M1 17961809 15849929
SK1856 M1 18679291 16567411
SK1857 M1 14788335 12676406
Y1422.2 M1 Z8214.2 rs369289787 8432768 8564727
Z8214.2 M1 Y1422.2 rs369289787 8432768 8564727
YSC0000696 M1 14976854 12864929
Y26288 M1 Z43397 14010417 11889711
Z43397 M1 Y26288 14010417 11889711
Y26291 M1 Z43398 15501203 13389323
Z43398 M1 Y26291 15501203 13389323
Y26294 M1 Z43404 19563479 17451599
Z43404 M1 Y26294 19563479 17451599
Y26298 M1 Z43408 22678773 20516887
Z43408 M1 Y26298 22678773 20516887
Y26299 M1 Z43396 6886974 7018933
Z43396 M1 Y26299 6886974 7018933
Y26300 M1 Z43410 24493037 22346890
Z43410 M1 Y26300 24493037 22346890
Y27828 M1 Z43395 6595053 6727012
Z43395 M1 Y27828 6595053 6727012
Y27835 M1 Z43399 16333594 14221714
Z43399 M1 Y27835 16333594 14221714
Y27836 M1 Z43403 17596363 15484483
Z43403 M1 Y27836 17596363 15484483
Z30832 M1 14346900 12226195
Z30839 M1 2854695 2986654
Z30840 M1 2859636 2991595
Z30841 M1 2865710 2997669
Z30890 M1 6619307 6751266
Z30891 M1 6662594 6794553
Z30892 M1 6679553 6811512
Z30893 M1 6689680 6821639
Z30894 M1 6706463 6838422
Z30895 M1 6748654 6880613
Z30896 M1 6795966 6927925
Z30897 M1 6815542 6947501
Z30898 M1 6827825 6959784
Z30899 M1 6886974 7018933
Z30900 M1 6909278 7041237
Z30901 M1 6912648 7044607
Z30902 M1 7008618 7140577
Z30903 M1 7053576 7185535
Z30904 M1 7078921 7210880
Z30905 M1 7131506 7263465
Z30908 M1 7306041 7438000
Z30909 M1 7326477 7458436
Z30910 M1 7364734 7496693
Z30912 M1 7386101 7518060
Z30913 M1 7389976 7521935
Z30914 M1 7395770 7527729
Z30915 M1 7401903 7533862
Z30916 M1 7405762 7537721
Z30918 M1 7557145 7689104
Z30919 M1 7664098 7796057
Z30921 M1 7750041 7882000
Z30922 M1 7789873 7921832
Z30923 M1 7794047 7926006
Z30924 M1 7826071 7958030
Z30925 M1 7844929 7976888
Z30926 M1 7992575 8124534
Z30930 M1 8101772 8233731
Z30931 M1 8684095 8816054
Z30932 M1 8755344 8887303
Z30933 M1 8789706 8921665
Z30935 M1 8846070 8978029
Z30936 M1 8858186 8990145
Z30939 M1 8881443 9013402
Z30940 M1 8893392 9025351
Z30942 M1 9391070 9553461
Z30943 M1 9392733 9555124
Z30944 M1 9400716 9563107
Z30948 M1 9644123 9806514
Z30950 M1 9817653 9980044
Z30951 M1 9823756 9986147
Z30952 M1 9833039 9995430
Z30953 M1 9885801 10048192
Z30954 M1 9887780 10050171
Z30956 M1 9918361 10080752
Z30957 M1 9926992 10089383
Z30958 M1 rs1477971071 10037299 10199690
Z30959 M1 10080437 10242828
Z30960 M1 10082081 10244472
Z30961 M1 10082642 10245033
Z30962 M1 13607472 11451796
Z30963 M1 13651743 11496067
Z30964 M1 13653454 11497778
Z30965 M1 13654014 11498338
Z30967 M1 13667892 11512216
Z30968 M1 13830963 11710257
Z30972 M1 13904301 11783595
Z30975 M1 14039586 11918880
Z30976 M1 14063409 11942703
Z30977 M1 14103794 11983088
Z30979 M1 14124885 12004179
Z30980 M1 14149253 12028547
Z30981 M1 14177187 12056481
Z30982 M1 14189708 12069002
Z30984 M1 14194554 12073848
Z30985 M1 14202391 12081685
Z30986 M1 14241165 12120459
Z30987 M1 14264491 12143785
Z30988 M1 14297259 12176553
Z30989 M1 14337877 12217172
Z30991 M1 14408047 12287345
Z30992 M1 14476980 12356249
Z30993 M1 14521025 12409226
Z30994 M1 14556522 12444723
Z30995 M1 14634528 12522597
Z30996 M1 14751028 12639097
Z30998 M1 14868592 12756662
Z30999 M1 14931961 12820026
Z31001 M1 15080736 12968826
Z31002 M1 15132373 13020460
Z31004 M1 15429680 13317800
Z31006 M1 15449612 13337732
Z31008 M1 15580350 13468470
Z31009 M1 15585370 13473490
Z31011 M1 15650036 13538156
Z31012 M1 15755127 13643247
Z31013 M1 15967363 13855483
Z31014 M1 16095034 13983154
Z31015 M1 16132693 14020813
Z31016 M1 16132852 14020972
Z31017 M1 16278592 14166712
Z31018 M1 16328036 14216156
Z31019 M1 16382498 14270618
Z31020 M1 16399716 14287836
Z31021 M1 16457180 14345300
Z31023 M1 rs990233096 16546532 14434652
Z31024 M1 16575466 14463586
Z31025 M1 16628741 14516861
Z31028 M1 16725197 14613317
Z31029 M1 16725361 14613481
Z31031 M1 16874027 14762147
Z31032 M1 rs985809826 16906766 14794886
Z31033 M1 16909563 14797683
Z31034 M1 17032128 14920248
Z31035 M1 17032134 14920254
Z31036 M1 17098116 14986236
Z31037 M1 17150371 15038491
Z31038 M1 17241411 15129531
Z31039 M1 17280140 15168260
Z31040 M1 17280893 15169013
Z31041 M1 17289694 15177814
Z31042 M1 17302581 15190701
Z31043 M1 17314001 15202121
Z31044 M1 17314491 15202611
Z31045 M1 17400046 15288166
Z31048 M1 17545760 15433880
Z31049 M1 17575999 15464119
Z31050 M1 17590506 15478626
Z31051 M1 17596639 15484759
Z31052 M1 17623639 15511759
Z31054 M1 17893290 15781410
Z31055 M1 17922264 15810384
Z31056 M1 18034383 15922503
Z31058 M1 18052198 15940318
Z31059 M1 18096942 15985062
Z31061 M1 18113885 16002005
Z31063 M1 18413781 16301901
Z31064 M1 18628116 16516236
Z31065 M1 18634599 16522719
Z31066 M1 18740907 16629027
Z31067 M1 18782507 16670627
Z31068 M1 18849664 16737784
Z31072 M1 19005336 16893456
Z31073 M1 19042711 16930831
Z31074 M1 19090266 16978386
Z31075 M1 19107344 16995464
Z31076 M1 19172087 17060207
Z31077 M1 19241561 17129681
Z31078 M1 19334083 17222203
Z31080 M1 19436907 17325027
Z31081 M1 19443332 17331452
Z31082 M1 19520060 17408180
Z31083 M1 20816739 18654853
Z31085 M1 21069302 18907416
Z31086 M1 21142313 18980427
Z31087 M1 21179019 19017133
Z31088 M1 21202963 19041077
Z31089 M1 21209376 19047490
Z31090 M1 21235835 19073949
Z31094 M1 21314389 19152503
Z31095 M1 21317402 19155516
Z31096 M1 21323767 19161881
Z31099 M1 21506368 19344482
Z31101 M1 21569659 19407773
Z31102 M1 21588611 19426725
Z31103 M1 21607514 19445628
Z31104 M1 21806552 19644666
Z31105 M1 21876169 19714283
Z31106 M1 21885884 19723998
Z31107 M1 21919199 19757313
Z31108 M1 21929233 19767347
Z31110 M1 rs915283150 22001168 19839282
Z31111 M1 22003568 19841682
Z31112 M1 22006880 19844994
Z31113 M1 22006975 19845089
Z31114 M1 22028307 19866421
Z31115 M1 22140515 19978629
Z31116 M1 22147651 19985765
Z31117 M1 22158004 19996118
Z31133 M1 22538867 20376981
Z31134 M1 22541105 20379219
Z31136 M1 22580381 20418495
Z31137 M1 22585364 20423478
Z31138 M1 22591707 20429821
Z31141 M1 22647025 20485139
Z31143 M1 22733730 20571844
Z31144 M1 22773249 20611363
Z31145 M1 22794375 20632489
Z31147 M1 22833461 20671575
Z31149 M1 22864231 20702345
Z31150 M1 22885471 20723585
Z31152 M1 22998408 20836522
Z31153 M1 23032344 20870458
Z31154 M1 23071887 20910001
Z31155 M1 23123650 20961764
Z31156 M1 23125531 20963645
Z31157 M1 23130833 20968947
Z31159 M1 23159757 20997871
Z31160 M1 23163023 21001137
Z31161 M1 rs920979989 23338962 21177076
Z31162 M1 23372003 21210117
Z31163 M1 23388612 21226726
Z31165 M1 23443020 21281134
Z31166 M1 23445191 21283305
Z31167 M1 23461113 21299227
Z31168 M1 23563089 21401203
Z31169 M1 23586950 21425064
Z31170 M1 23596565 21434679
Z31171 M1 23622179 21460293
Z31172 M1 23635202 21473316
Z31173 M1 23749328 21587442
Z31174 M1 23781867 21619981
Z31175 M1 23786763 21624877
Z31181 M1 23980433 21834286
Z31182 M1 23996887 21850740
Z31183 M1 24401989 22255842
Z31184 M1 24419496 22273349
Z31185 M1 24433677 22287530
Z31187 M1 28778515 26632368
Z31188 M1 28787742 26641595
Z31189 M1 8187045 8319004
Z31190 M1 8225946 8357905
Z31191 M1 8309974 8441933
Z31192 M1 8320153 8452112
Z31193 M1 8365184 8497143
Z31194 M1 8369453 8501412
Z31195 M1 8369465 8501424
Z31196 M1 8378998 8510957
Z31197 M1 8480806 8612765
Z31198 M1 8645197 8777156
Z33312 M1 8340502 8472461
Z33344 M1 21072781 18910895
FGC13408.2 M1~ Z1639.2 rs1039187893 21834863 19672977
Z1639.2 M1~ FGC13408.2 rs1039187893 21834863 19672977
M186 M1~ 14923461 12811526
M5991 M1~ rs556563930 21229717 19067831
Z30911 M1~ 7378628 7510587
Z30929 M1~ 8064301 8196260
Z30934.1 M1~ rs775425105 8811347 8943306
Z30945 M1~ 9411414 9573805
Z30949 M1~ 9767079 9929470
Z30955 M1~ 9899837 10062228
Z30966 M1~ 13666831 11511155
Z30978 M1~ 14122830 12002124
Z31010 M1~ 15631065 13519185
Z31026 M1~ 16700935 14589055
Z31057 M1~ 18045037 15933157
Z31070 M1~ 18969468 16857588
Z31071 M1~ 18975759 16863879
Z31093 M1~ 21270308 19108422
Z31097 M1~ 21392977 19231091
Z31100 M1~ 21531897 19370011
Z31140 M1~ 22622775 20460889
Z31148 M1~ 22852111 20690225
Z31164 M1~ 23441702 21279816
Z30983 M1a 14193991 12073285
Z30997 M1a 14796147 12684218
Z31022 M1a 16527122 14415242
Z31030 M1a 16764117 14652237
Z31069 M1a 18854932 16743052
SK1846 M1a1 Z43000 16635832 14523952
Z43400 M1a1 SK1846 16635832 14523952
Z30917 M1a1 7416383 7548342
BY8413.1 M1a1a Z30938.1 8872065 9004024
Z30938.1 M1a1a BY8413.1 8872065 9004024
SK1869 M1a1a 8547532 8679491
Z30835 M1a1a 16324762 14212882
Z30907 M1a1a 7244645 7376604
Z30917 M1a1a 7416383 7548342
Z30920 M1a1a 7731507 7863466
Z30928 M1a1a 8022892 8154851
Z30937 M1a1a 8867037 8998996
Z31000 M1a1a 15006247 12894332
Z31005 M1a1a 15441425 13329545
Z31007 M1a1a 15576350 13464470
Z31046 M1a1a 17417323 15305443
Z31060 M1a1a 18107412 15995532
Z31062 M1a1a 18208202 16096322
Z31079 M1a1a 19339054 17227174
Z31151 M1a1a 22953120 20791234
Z31158 M1a1a 23150645 20988759
P94 M1a1a1~ 14869681 12757751
Z30928 M1a1a2 8022892 8154851
Z31007 M1a1a2 15576350 13464470
Z31146 M1a1a2 22808950 20647064
P87 M1a1a2a rs868292303 15019719 12907808
M104_1^^ M1a1a2a1 P22_1^^ 8619592 8751551
P22_1^^ M1a1a2a1 M104_1^^ 8619592 8751551
M104_2^^ M1a1a2a1 P22_2^^ 8620324 8752283
P22_2^^ M1a1a2a1 M104_2^^ 8620324 8752283
F2561 M1a2a2a1~ Page67 rs35852845 17678803 15566923
Page67 M1a2a2a1~ F2561 rs35852845 17678803 15566923
FGC24709 M1a2a2a2~ 8212218 8344177
FGC36703 M1a2a2a2~ 9840359 10002750
SK1877 M1a2a2a2~ rs907039672 8545089 8677048
SK1879 M1a2a2a2~ 17844157 15732277
Z42236 M1a2a2a2~ 7519846 7651805
Z42237 M1a2a2a2~ 7528865 7660824
Z42238 M1a2a2a2~ 7583062 7715021
Z42239 M1a2a2a2~ 7680906 7812865
Z42240 M1a2a2a2~ 7707247 7839206
Z42241 M1a2a2a2~ 7803606 7935565
Z42242 M1a2a2a2~ 7870593 8002552
Z42243 M1a2a2a2~ 8243993 8375952
Z42244 M1a2a2a2~ 8372862 8504821
Z42245 M1a2a2a2~ 8733591 8865550
Z42246 M1a2a2a2~ 9891791 10054182
Z42247 M1a2a2a2~ 14073836 11953130
Z42248 M1a2a2a2~ 14097247 11976541
Z42249 M1a2a2a2~ 14116928 11996222
Z42250 M1a2a2a2~ 14238977 12118271
Z42251 M1a2a2a2~ 14265828 12145122
Z42252 M1a2a2a2~ 14265948 12145242
Z42253 M1a2a2a2~ 14292115 12171409
Z42254 M1a2a2a2~ 14772806 12660875
Z42255 M1a2a2a2~ 15361173 13249293
Z42256 M1a2a2a2~ 16355236 14243356
Z42257 M1a2a2a2~ rs1013506222 16541971 14430091
Z42258 M1a2a2a2~ 16625031 14513151
Z42259 M1a2a2a2~ 16663490 14551610
Z42260 M1a2a2a2~ 16744143 14632263
Z42261 M1a2a2a2~ 16966249 14854369
Z42262 M1a2a2a2~ rs1252039405 17167617 15055737
Z42263 M1a2a2a2~ 17852575 15740695
Z42264 M1a2a2a2~ 18186626 16074746
Z42265 M1a2a2a2~ 18725433 16613553
Z42266 M1a2a2a2~ 19068368 16956488
Z42267 M1a2a2a2~ 19315641 17203761
Z42268 M1a2a2a2~ 19341447 17229567
Z42269 M1a2a2a2~ 19444636 17332756
Z42270 M1a2a2a2~ 21145277 18983391
Z42271 M1a2a2a2~ 17513642 15401762
Z42272 M1a2a2a2~ 21221362 19059476
Z42273 M1a2a2a2~ 21231346 19069460
Z42274 M1a2a2a2~ 21357947 19196061
Z42275 M1a2a2a2~ 21448467 19286581
Z42276 M1a2a2a2~ 21514652 19352766
Z42277 M1a2a2a2~ rs1041652979 21557719 19395833
Z42278 M1a2a2a2~ 22784216 20622330
Z42279 M1a2a2a2~ 22791140 20629254
Z42280 M1a2a2a2~ 22802316 20640430
M16 M1a1a2a2a rs3907 21610301 19448415
M83 M1a1a2a2b rs2032641 21873287 19711401
SK1881 M1a1a1a1b~ 7999341 8131300
Z42285 M1a1a1a1b~ 6807417 6939376
Z42286 M1a1a1a1b~ 6838413 6970372
Z42287 M1a1a1a1b~ 7676637 7808596
SK1880 M1a1a1a1c~ 17568668 15456788
Z42281 M1a1a1a1c~ 8145744 8277703
Z42282 M1a1a1a1c~ 8202129 8334088
Z42283 M1a1a1a1c~ 15597434 13485554
Z42284 M1a1a1a1c~ 17567694 15455814
Z33330 M1a1a3~ rs896050429 16960342 14848462
Z33345 M1a1a3~ 21284118 19122232
Z42288 M1a1a3~ 23795714 21633828
P51 M1a1a3a~ rs376386703 14492042 12380239
SK1872 M1a1a3a~ 2720899 2852858
Z42289 M1a1a3a~ 2672414 2804373
Z42290 M1a1a3a~ 6880665 7012624
Z42291 M1a1a3a~ 6994851 7126810
Z42292 M1a1a3a~ 7574376 7706335
B256 M1a1a3b 2723942 2855901
SK1882 M1a1a3b 18110438 15998558
Z31135 M1a1a3b 22580315 20418429
Z33309 M1a1a3b 2851943 2983902
Z33313 M1a1a3b 8369136 8501095
Z33318 M1a1a3b 14050849 11930143
Z33319 M1a1a3b 14139507 12018801
Z33323 M1a1a3b 15059490 12947582
Z33325 M1a1a3b 15733463 13621583
Z33331 M1a1a3b 16970846 14858966
Z33336 M1a1a3b 18050131 15938251
Z33343 M1a1a3b 19478645 17366765
Z33347 M1a1a3b 22806304 20644418
Z33348 M1a1a3b 22994606 20832720
Z33350 M1a1a3b 23281355 21119469
Z33310 M1a1a3b~ 6984103 7116062
Z33311 M1a1a3b~ 7990628 8122587
Z33317 M1a1a3b~ 13996958 11876252
Z33320 M1a1a3b~ 14632848 12520917
Z33321 M1a1a3b~ 14743029 12631097
Z33324 M1a1a3b~ 15108821 12996908
Z33326 M1a1a3b~ 15910717 13798837
Z33327 M1a1a3b~ 16277784 14165904
Z33332 M1a1a3b~ 17014713 14902833
Z33337 M1a1a3b~ 18185841 16073961
Z33341 M1a1a3b~ 18630858 16518978
Z33351 M1a1a3b~ 28563495 26417348
Z33322 M1a1a3b1 14751334 12639403
Z33328 M1a1a3b1 16334244 14222364
Z33114 M1a1a3b1~ 8456245 8588204
Z33315 M1a1a3b1~ 8491698 8623657
Z33316 M1a1a3b1~ 9843427 10005818
Z33317 M1a1a3b1~ 13996958 11876252
Z33329 M1a1a3b1~ 16481545 14369665
Z33333 M1a1a3b1~ 17246382 15134502
Z33339 M1a1a3b1~ rs953852329 18561672 16449792
Z33340 M1a1a3b1~ 18612566 16500686
Z33342 M1a1a3b1~ 19401320 17289440
SK1889 M1a1a3b1a~ 6779737 6911696
Z42293 M1a1a3b1a~ 2683463 2815422
Z42294 M1a1a3b1a~ 7585279 7717238
Z42295 M1a1a3b1a~ 7698724 7830683
Z42296 M1a1a3b1a~ 8012216 8144175
Z42297 M1a1a3b1b 6876680 7008639
Z42298 M1a1a3b1b 7106708 7238667
Z42299 M1a1a3b1b 7107917 7239876
Z42300 M1a1a3b1b 7750092 7882051
Z42301 M1a1a3b1b 8121320 8253279
Z42302 M1a1a3b1b 8179883 8311842
Z42303 M1a1a3b1b 8770794 8902753
Z42304 M1a1a3b1b 14117547 11996841
Z42305 M1a1a3b1b 14126987 12006281
Z42306 M1a1a3b1b 14383921 12263217
Z42307 M1a1a3b1b 16210714 14098834
Z42308 M1a1a3b1b 16214328 14102448
Z42309 M1a1a3b1b 16267038 14155158
Z42310 M1a1a3b1b 16747765 14635885
Z42311 M1a1a3b1b 16889638 14777758
Z42312 M1a1a3b1b 16936767 14824887
Z42313 M1a1a3b1b 17048618 14936738
Z42314 M1a1a3b1b 17723819 15611939
Z42315 M1a1a3b1b 17875975 15764095
Z42316 M1a1a3b1b 21275295 19113409
Z42317 M1a1a3b1b 21327140 19165254
Z42318 M1a1a3b1b 21547520 19385634
Z42319 M1a1a3b1b 22076024 19914138
Z42320 M1a1a3b1b 22871185 20709299
Z42321 M1a1a3b1b 23304124 21142238
Z42322 M1a1a3b1b 28734954 26588807
Z42326 M1a1a3b1b1~ Y26302 7579846 7651805
Y26302 M1a1a3b1b1~ Z42326 7579846 7651805
Z42323 M1a1a3b1b1~ 6828273 6960232
Z42324 M1a1a3b1b1~ 8257926 8389885
Z42325 M1a1a3b1b1~ 8644753 8776712
Z42327 M1a1a3b1b1~ 15017486 12905575
BY3630.2 M1a1a3b1b2~ rs1016979501 7694785 7826744
F1295.2 M1a1a3b1b2~ rs1016979501 8545494 8677453
Z42328 M1a1a3b1b2~ 7156933 7288892
Z42329 M1a1a3b1b2~ 7243541 7375500
Z42330 M1a1a3b1b2~ 8484349 8616308
B269 M1a1a3b1c 2885637 3017596
Z31204.2 M1a1a3b1c 7325371 7457330
Z33269 M1a1a3b1c 7327175 7459134
Z33270 M1a1a3b1c 7651974 7783933
Z33271 M1a1a3b1c 8058221 8190180
Z33272 M1a1a3b1c 8493986 8625945
Z33273 M1a1a3b1c rs918535566 8591531 8723490
Z33274 M1a1a3b1c 8847987 8979946
Z33276 M1a1a3b1c 9125962 9288353
Z33277 M1a1a3b1c 14056352 11935646
Z33278 M1a1a3b1c 14129203 12008497
Z33279 M1a1a3b1c 14175024 12054318
Z33280 M1a1a3b1c 14183482 12062776
Z33281 M1a1a3b1c 14469116 12348385
Z33282 M1a1a3b1c 14925718 12813783
Z33283 M1a1a3b1c 14975906 12863981
Z33284 M1a1a3b1c 15010476 12898561
Z33285 M1a1a3b1c 15125312 13013399
Z33286 M1a1a3b1c 15240791 13128877
Z33288 M1a1a3b1c 15311169 13199274
Z33289 M1a1a3b1c 16523817 14411937
Z33290 M1a1a3b1c 16569772 14457892
Z33291 M1a1a3b1c 16913805 14801925
Z33292 M1a1a3b1c 17241587 15129707
Z33293 M1a1a3b1c 18676823 16564943
Z33295 M1a1a3b1c 19356211 17244331
Z33296 M1a1a3b1c 21262082 19100196
Z33297 M1a1a3b1c 21574060 19412174
Z33298 M1a1a3b1c 22791814 20629928
Z33299 M1a1a3b1c 23147416 20985530
Z33300 M1a1a3b1c 23185558 21023672
B271 M1a1a3b1c1~ 7269443 7401402
Z33302 M1a1a3b1c1~ 8156524 8288483
Z33305 M1a1a3b1c1~ 7341503 7473462
Z33307 M1a1a3b1c1~ 8537044 8669003
Z33308 M1a1a3b1c1~ 9880005 10042396
B270 M1a1a3b1c2~ 14038578 11917872
Z38339 M1a1a3b1c2~ 14296545 12175839
Z38340 M1a1a3b1c2~ rs777121309 14328165 12207459
Z38341 M1a1a3b1c2~ 15042639 12930729
B272 M1a1a3b1d~ 6903117 7035076
Z33740 M1a1a3b1d~ 2889563 3021522
Z33741 M1a1a3b1d~ 7159520 7291479
Z33742 M1a1a3b1d~ 7638937 7770896
Z33743 M1a1a3b1d~ 8608287 8740246
Z43837 M1a1a3b2 Y26293 18177988 16066108
Y26293 M1a1a3b2 Z43837 18177988 16066108
Z43838 M1a1a3b2 10007133 10169524
M10781 M1a1a3b2~ S20812; ZS4770 rs200810549 18209106 16097226
S20812 M1a1a3b2~ M10781; ZS4770 rs200810549 18209106 16097226
ZS4770 M1a1a3b2~ M10781; S20812 rs200810549 18209106 16097226
Z42538 M1a1a3b2a~ 2722393 2854352
Z42539 M1a1a3b2a~ 6702580 6834539
Z42540 M1a1a3b2a~ 7077655 7209614
Z42541 M1a1a3b2a~ 7307366 7439325
Z42542 M1a1a3b2a~ 7637584 7769543
Z43832 M1a1a3b2b~ 6735210 6867169
Z43833 M1a1a3b2b~ 7284544 7416503
Z43834 M1a1a3b2b~ 7643536 7775495
Z43835 M1a1a3b2b~ 8098451 8230410
Z43836 M1a1a3b2b~ 8400418 8532377
Z43411 M1a2~ 2751052 2883011
Z43412 M1a2~ 2807488 2939447
Z43413 M1a2~ 2913130 3045089
Z43414 M1a2~ 6699692 6831651
Z43415 M1a2~ 6891091 7023050
FT254675 M1b~ 14344207 12223502
M353 M2 rs796470087 17309267 15197387
M387 M2 rs868120210 2740274 2872233
PF4350.1 M2~ 7746566 7878525
M177^^ M2a SRY9138^^ 2658869 2790828
SRY9138^^ M2a M177^^ 2658869 2790828
P117 M3 14819693 12707760
 Subclades - 2019-2020

han copying. Links may not copy to new sheet.

Main Page.

ision date for this specific page: 9 April 2020

nges amd will be updated when possible

Email Ray Banks

Papers/Presentations Cited Listing Criteria

nfirmed within subclade Investigational items

meet quality guidelines for minimum number of reads.

y be helpful.

usage, testing done in multiple labs, and/or being

eneration sequencing reports consistently showing as
up, but some confirmatory evidence is not yet

182, Z31183, Z31184, Z31185, Z31187, Z31188, Z31189, Z31190. Z31191, Z31192. Z31193,
45, Z30949, Z30955, Z30966, Z30978, Z31010, Z31026, Z31057, Z31070, Z31071, Z31093,

8.1, SK1869, Z30835, Z30907, Z30917, Z30920, Z30928, Z30937, Z31000, Z31005, Z31007, Z31046, Z31060, Z31062,
Z31158

0928, Z31007, Z31146

P87
M1a1a2a1 M104_1^^/P22_1^^, M104_2^^/P22_2^^
M1a2a2a1~ F2561/Page67
Z42248, Z42249, Z42250, Z42251, Z42252, Z42253, Z42254, Z42255, Z42256, Z42257, Z42258, Z42259, Z42
M1a2a2a2~ Z42265, Z42266, Z42267, Z42268, Z42269, Z42270, Z42271, Z42272, Z42273, Z42274, Z42275, Z42276, Z42
M1a1a2a2a M16
M1a1a2a2b M83
M1a1a1a1b~ SK1881, Z42285, Z42286, Z42287
M1a1a1a1c~ SK1880, Z42281, Z42282, Z42283, Z42284
3330, Z33345, Z42288
P51, SK1872, Z42289, Z42290, Z42291, Z42292
B256, SK1882, Z31135, Z33309, Z33313, Z33318, Z33319, Z33323, Z33325, Z33331, Z33336, Z33343, Z33347, Z33348
Z33310, Z33311, Z33317, Z33320, Z33321, Z33324, Z33326, Z33327, Z33332, Z33337, Z33341, Z33351
M1a1a3b1 Z33322, Z33328
M1a1a3b1~ Z33114. Z33315, Z33316, Z33317, Z33329, Z33333, Z33339, Z33340, Z33342
M1a1a3b1a~ SK1889, Z42293, Z42294,
Z42312, Z42313, Z42314,Z42295,
Z42315,Z42296
Z42307, Z42308, Z42309, Z42310, Z42311, Z42312, Z42
M1a1a3b1b Z42318, Z42319, Z42320, Z42321, Z42322
M1a1a3b1b1~ Y26302/Z42326, Z42323, Z42324, Z42325, Z42327
M1a1a3b1b2~ BY3630.2,
Z33283, Z33284, Z33277, F1295.2, Z42328, Z42329,
Z33278, Z33279, Z33280, Z42330
Z33281, Z33282, Z33283, Z33284, Z33
M1a1a3b1c Z33291, Z33292, Z33293, Z33295, Z33296, Z33297, Z33298, Z33299, Z33300
M1a1a3b1c1~ B271, Z33302, Z33305, Z33307, Z33308
M1a1a3b1c2~ B270, Z38339, Z38340, Z38341
M1a1a3b1d~ B272, Z33740. Z33741, Z33742, Z33743
M1a1a3b2 Y26293/Z43837, Z43838
M1a1a3b2~ M10781/S20812/ZS4770
M1a1a3b2a~ Z42538, Z42539, Z42540, Z42541, Z42542
M1a1a3b2b~ Z43832. Z43833, Z43834, Z43835, Z43836

he order of discovery,

cent chromosome position 8685231.

two-thirds of population.
cent arrivals among Aborigines (Bergstrom).
 ogy, 26:809-813, (2016).
istory of New Guinea Science, 357:1160-1163 (2017).
Affiliation in Paired Austronesian- and Papuan-Speaking Communities from Solomon Islands. (pdf) American Journal of Human
ion Structure Revealed by mtDNA. American Journal of Human Genetics, 63:1793-1806, 1998.
tics, 160:289-303, January 2002.
ange in Culture. Genome Research, doi: 10.1101/gr.186684.114, published online March 13, 2015.
Y-Chromosomal Haplogroup Tree. Abstract. Genome Research, published online April 2, 2008. Supplementary Material.
omosome Diversity in the Admiralty Islands of Melanesia. Molecular Biology Evolution, 25(7):1362-1374, 2008.
lia. American Journal of Human Genetics, 68:173-190, 2001.
radients across the Pacific. MBE Advance Access published August 21, 2006.
No. 20, September 2000.
opulations from West New Guinea. American Journal of Human Genetics, 72:281-302, 2003.
othesis. Mol Biol Evol. 2007 Sep 10; [Epub ahead of print]
le-Copy Genes, Implying Effective Purifying Selection. American Journal of Human Genetics. 2009 December 11; 85(6): 923-9
Society for Molecular Biology, 2006.

10 April 2020.
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1046, Z31060, Z31062,

Z42256, Z42257, Z42258, Z42259, Z42260, Z42261, Z42262, Z42263, Z42264,

Z42273, Z42274, Z42275, Z42276, Z42277, Z42278, Z42279, Z42280
1, Z33336, Z33343, Z33347, Z33348, Z33350
7, Z33341, Z33351

2309, Z42310, Z42311, Z42312, Z42313, Z42314, Z42315, Z42316, Z42317.

3281, Z33282, Z33283, Z33284, Z33285, Z33286, Z33288, Z33289, Z33290,

3298, Z33299, Z33300
. (pdf) American Journal of Human Biology, 18:35-50, 2006.

08. Supplementary Material.

7):1362-1374, 2008.

cs. 2009 December 11; 85(6): 923-928.