2.4 Respiratory System

Respiratory System
Trading with the Environment

Every organism must exchange materials with its
environment
And this exchange ultimately occurs at the cellular level
In unicellular organisms
These exchanges occur directly with the environment
For most of the cells making up multicellular organisms
Direct exchange with the environment is not possible
Concept : Gas exchange occurs across

specialized respiratory surfaces
Gas exchange
The movement of O2 and CO2 between the animal
and its environment.
Supplies oxygen for cellular respiration and
disposes of carbon dioxide
Respiratory
medium
(air of water)
O2
CO2
Respiratory
surface
Organismal
level
Circulatory system
Cellular level
Energy-rich
molecules
from food
Figure 42.19
Cellular respiration
ATP
Respiratory surface
Portion of the animal surface where gas
exchange with the respiratory medium occurs.
Oxygen diffuses in; carbon dioxide diffuses
out.
Gills in Aquatic Animals

Animals require large, moist respiratory
surfaces for the adequate diffusion of
respiratory gases
Between their cells and the respiratory medium,
either air or water
Gills are outfoldings of the body surface

Specialized for gas exchange
Eg: Gills in invertebrates (sea star), segmented
worm, clamp & crayfish.
The feathery gills projecting from a salmon

Are an example of a specialized exchange
system found in animals
Figure 42.1
Ventilation
Any method of increasing the flow of the
respiratory medium over the respiratory
surface; brings in a fresh supply of O2 and
removes CO2.
The effectiveness of gas exchange in

some gills, including those of fishes
Is increased by ventilation and countercurrent
flow of blood and water
Oxygen-poor
blood
Lamella
%
% 15
40
Figure 42.21
30
%
60
%
Operculum
90
%
Water
flow
10
0%
70
%
Gill
arch
Blood
vessel
Oxygen-rich
blood
5%
Gill arch
Water flow
over lamellae
showing % O2
Gill
filaments
O2
Blood flow
through capillaries
in lamellae
showing % O2
Countercurrent exchange
Tracheal Systems in Insects

The tracheal system of insects
Consists of tiny branching tubes that
penetrate the body
Air sacs
Tracheae
Spiracle
(a) The respiratory system of an insect consists of branched internal

tubes that deliver air directly to body cells. Rings of chitin reinforce
the largest tubes, called tracheae, keeping them from collapsing.
Enlarged portions of tracheae form air sacs near organs that require
a large supply of oxygen. Air enters the tracheae through openings
called spiracles on the insects body surface and passes into smaller
tubes called tracheoles. The tracheoles are closed and contain fluid
(blue-gray). When the animal is active and is using more O2, most of
the fluid is withdrawn into the body. This increases the surface area
of air in contact with cells.
Figure 42.22a
The tracheal tubes

Supply O2 directly to body cells
Body
cell
Air
sac
Tracheole
Trachea
Air
Tracheoles
Mitochondria
Body wall
Myofibrils
(b) This micrograph shows cross

sections of tracheoles in a tiny
piece of insect flight muscle (TEM).
Each of the numerous mitochondria
in the muscle cells lies within about
5 m of a tracheole.
Figure 42.22b
2.5 m
Lungs
Spiders, land snails, and most terrestrial vertebrates
Have internal lungs
In mammals, air inhaled through the nostrils

Passes through the pharynx into the trachea, bronchi, bronchioles, and
dead-end alveoli, where gas exchange occurs
The process that ventilates the lungs is breathing

The alternate inhalation and exhalation of air
Mammalian Respiratory Systems: A

Closer Look
A system of branching ducts
Conveys air to the lungs
Nasal
cavity
Pharynx
Branch
from the
pulmonary
artery
(oxygen-poor
blood)
Branch
from the
pulmonary
vein
(oxygen-rich
blood)
Terminal
bronchiole
Left
lung
Alveoli
50 m
Larynx
Esophagus
Trachea
Right lung
50 m
Bronchus
Bronchiole
Diaphragm
Heart
SEM
Figure 42.23
Colorized SEM
How an Amphibian Breathes

An amphibian such as a frog
Ventilates its lungs by positive pressure
breathing, which forces air down the trachea
Air is pulled into the mouth by lowering the
floor region; this enlarges the oral cavity.
The nostrils and mouth is raised, forcing air
down the trachea.
Air is exhaled by elastic recoil of lungs and by
compression of the lungs by the muscular
body wall.
How a Mammal Breathes

Mammals ventilate their lungs
By negative pressure breathing, which pulls air into the lungs
Lung volume increases

As the rib muscles and diaphragm contract
Rib cage
expands as
rib muscles
contract
Air inhaled
Rib cage gets

smaller as
rib muscles
relax
Air exhaled
Lung
Diaphragm
INHALATION
Diaphragm contracts
(moves down)
Figure 42.24
EXHALATION
Diaphragm relaxes
(moves up)
How a Bird Breathes

Besides lungs, bird have eight or nine air sacs
That function as bellows that keep air flowing through
the lungs
Air
Air
Anterior
air sacs
Trachea
Posterior
air sacs
Lungs
Lungs
Air tubes
(parabronchi)
in lung
INHALATION
Air sacs fill
EXHALATION
Air sacs empty; lungs fill
Figure 42.25
1 mm
Control of Breathing in Humans

The main breathing control centers
Are located in two regions of the brain, the
medulla oblongata and the pons
Cerebrospinal
fluid
1 The control center in the

medulla sets the basic
rhythm, and a control center
in the pons moderates it,
smoothing out the
transitions between
inhalations and exhalations.
Pons
2 Nerve impulses trigger
muscle contraction. Nerves
from a breathing control center
in the medulla oblongata of the
brain send impulses to the
diaphragm and rib muscles,
stimulating them to contract
and causing inhalation.
Breathing
control
centers
Medulla
oblongata
4 The medullas control center

also helps regulate blood CO2 level.
Sensors in the medulla detect
changes
in the pH (reflecting CO2
concentration)
of the blood and cerebrospinal fluid
changes in
5 Nerve
bathing
theimpulses
surface ofrelay
the brain.
CO2 and O2 concentrations. Other
sensors in the walls of the aorta
and carotid arteries in the neck
detect changes in blood pH and
send nerve impulses to the medulla.
In response, the medullas breathing
control center alters the rate and
depth of breathing, increasing both
to dispose of excess CO2 or decreasing
both if CO2 levels are depressed.
Carotid
arteries
Figure 42.26
3 In a person at rest, these

nerve impulses result in
about 10 to 14 inhalations
per minute. Between
inhalations, the muscles
relax and the person exhales.
Aorta
Diaphragm
Rib muscles
6 The sensors in the aorta and

carotid arteries also detect changes
in O2 levels in the blood and signal
the medulla to increase the breathing
rate when levels become very low.
The centers in the medulla

Regulate the rate and depth of breathing in
response to pH changes in the cerebrospinal fluid
The medulla adjusts breathing rate and depth

To match metabolic demands
Sensors in the aorta and carotid arteries

Monitor O2 and CO2 concentrations in the blood
Exert secondary control over breathing
The amount of air inhaled and exhaled

depends upon size, activity level and state
of health.
Tidal volume is the volume of air an animal
inhales and exhales with each breath during
normal quiet breathing. Averages about 500
ml in humans.
Vital volume is the maximum air volume that
can be inhaled and exhaled during forced
breathing. Averages 3400-4800 ml in collegeage females and males, respectively.
Residual volume is the amount of air that
remain in the lungs even after forced
exhalation.
Concept: Respiratory pigments bind and

transport gases
The metabolic demands of many
organisms
Require that the blood transport large
quantities of O2 and CO2
The Role of Partial Pressure

Gradients
Gases diffuse down pressure gradients
In the lungs and other organs
Diffusion of a gas
Depends on differences in a quantity called partial
pressure
A gas always diffuses from a region of higher

partial pressure
To a region of lower partial pressure
In the lungs and in the tissues

O2 and CO2 diffuse from where their partial pressures
are higher to where they are lower
Inhaled air
160
O2
Exhaled air
0.2
CO2
O2
104
Alveolar
epithelial
cells
O2
40
O2
CO2
O2
Alveolar
capillaries
of lung
45
O2
CO2
Blood
leaving
alveolar
capillaries
104
40
O2
CO2
Pulmonary
veins
Pulmonary
arteries
Systemic
arteries
Systemic
veins
Heart
Tissue
capillaries
CO2
Blood
leaving
tissue
capillaries
40
O2
Figure 42.27
CO2
40
CO2
CO
Blood
entering
alveolar
capillaries
120 27
Alveolar spaces
45
O2
Blood
entering
tissue
capillaries
O2
CO2
100
O2
CO2
Tissue
cells
<40 >45
O2
CO2
40
CO2
Animals
Respiratory
Medium
Respiratory
Surface.
Amoeba
Water
Plasma
membrane.
Grasshopper
Air
Trachea
Fish
Water
Gills
Frog
Air
Lungs and Skin
Human
Air
Lungs
Air has several advantages over water as

a respiratory medium
A higher oxygen concentration
Oxygen and carbon dioxide diffuse faster
through air than water
Respiratory surfaces do not have to be
ventilate as thoroughly
Disadvantage
The respiratory surfaces are continually
desiccated.

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Respiratory System

Trading with the Environment

Concept : Gas exchange occurs across

Gills in Aquatic Animals

Gills are outfoldings of the body surface

The feathery gills projecting from a salmon

The effectiveness of gas exchange in

Tracheal Systems in Insects

(a) The respiratory system of an insect consists of branched internal

The tracheal tubes

(b) This micrograph shows cross

In mammals, air inhaled through the nostrils

The process that ventilates the lungs is breathing

Mammalian Respiratory Systems: A

Conveys air to the lungs

How an Amphibian Breathes

How a Mammal Breathes

Lung volume increases

Rib cage gets

How a Bird Breathes

Control of Breathing in Humans

1 The control center in the

4 The medullas control center

3 In a person at rest, these

6 The sensors in the aorta and

The centers in the medulla

The medulla adjusts breathing rate and depth

Sensors in the aorta and carotid arteries

The amount of air inhaled and exhaled

Concept: Respiratory pigments bind and

The Role of Partial Pressure

A gas always diffuses from a region of higher

In the lungs and in the tissues

Lungs and Skin

Air has several advantages over water as

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