FISHES of The WORLD

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FISHES

FISH
†Superclass ANASPIDOMORPHI
classification †Class ANASPIDA
†Order ANASPIDIFORMES

†Superclass THELODONTOMORPHI
PHYLUM CHORDATA †Class THELODONTI
†Order ARCHIPELEPIDIFORMES
†Order FURCACAUDIFORMES FORK-TAILED THELODONTS
SUBPHYLUM UROCHORDATA TUNICATES †Order THELODONTIFORMES
Class ASCIDIACEA ASCIDIANS
Class THALIACEA SALPS †Superclass OSTEOSTRACOMORPHI
Order PYROSOMIDA †Class CEPHALASPIDOMORPHI
Order DOLIOLIDA †Order CEPHALASPIDIFORMES
Order SALPIDA †Order GALEASPIDIFORMES
Class APPENDICULARIA †Order PITURIASPIDIFORMES

Superclass GNATHOSTOMATA JAWED VERTEBRATES


SUBPHYLUM CEPHALOCHORDATA †GRADE PLACODERMIOMORPHI
Order AMPHIOXIFORMES LANCELETS †Class PLACODERMI
Family BRANCHIOSTOMATIDAE †Order PSEUDOPETALICHTHYIFORMES
Family EPIGONICHTHYIDAE †Order ACANTHOTHORACIFORMES
†Order RHENANIFORMES
†Order ANTIARCHIFORMES ANTIARCHS
†SUBPHYLUM CONODONTOPHORIDA CONODONTS †Suborder Yunnanolepidoidei
†Class CONODONTA †Family CHUCHINOLEPIDIDAE
†Family YUNNANOLEPIDIDAE
†Suborder Bothriolepidoidei
SUBPHYLUM CRANIATA †Infraorder Sinolepidida
†Family SINOLEPIDAE
INFRAPHYLUM MYXINOMORPHI †Infraorder Euantiarcha
Class MYXINI †Family MICROBRACHIIDAE
Order MYXINIFORMES (1) HAGFISHES †Family BOTHRIOLEPIDIDAE
Family MYXINIDAE (1) HAGFISHES †Family ASTEROLEPIDIDAE
Subfamily Rubicundinae †Family GERDALEPIDIDAE
Subfamily Eptatretinae †Order PETALICHTHYIFORMES
Subfamily Myxininae
†Order PTYCTODONTIFORMES
†Order ARTHRODIRIFORMES ARTHRODIRES
†Suborder Actinolepidoidei
INFRAPHYLUM VERTEBRATA VERTEBRATES †Family ACTINOLEPIDIDAE etc.
†Anatolepis †Suborder Phyllolepidoidei
†Family PHYLLOLEPIDIDAE
Superclass PETROMYZONTOMORPHI
Class PETROMYZONTIDA †Suborder Phlyctaenioidei
†Family PHLYCTAENIIDAE
Order PETROMYZONTIFORMES (2) LAMPREYS
†Family MAYOMYZONTIDAE †Suborder Brachythoracoidei
Family PETROMYZONTIDAE (2) NORTHERN LAMPREYS
†Eubrachythoraci
†Pachyosteomorpha
Subfamily Petromyzontinae
†Coccosteomorpha
Subfamily Lampetrinae
†Family BUCHANOSTEIDAE
Family GEOTRIIDAE (3) SOUTHERN LAMPREYS
†Family CAMUROPISCIDAE
Family MORDACIIDAE (4) SOUTHERN TOPEYED LAMPREYS
†Family COCCOSTEIDAE
†Superclass PTERASPIDOMORPHI †Family DINICHTHYIDAE
†Class PTERASPIDOMORPHA †Family DUNKLEOSTEIDAE
Subclass ASTRASPIDA †Family MYLOSTOMIDAE
†Order ASTRASPIDIFORMES †Family PANXIOSTEIDAE
Subclass ARANDASPIDA †Family SELENOSTEIDAE
†Order ARANDASPIDIFORMES †Family TITANICHTHYIDAE
Subclass HETEROSTRACI †Family WUTTAGOONASPIDIDAE
†Order CARDIPELTIDA
†Order CORVASPIDIDA EUGNATHOSTOMATA
†Order LEPIDASPIDIDA GRADE CHONDRICHTHYOMORPHI
†Order TESSERASPIDIDA Class CHONDRICHTHYES CARTILAGINOUS FISHES
†Order TRAQUAIRASPIDIFORMES Incertae sedis basal chondrichthyan taxa:
†Order TOLYPELEPIDIDA †Obtusacanthus
†Order CYATHASPIDIFORMES †Family PROTODONTIDAE
†Family AMPHIASPIDIDAE †Family KATHEMACANTHIDAE
†Family CYATHASPIDIDAE †Order POLYMEROLEPIDIFORMES
†Order PTERASPIDIFORMES †Order OMALODONTIFORMES
†Family ANCHIPTERASPIDIDAE †Family AZTECODONTIDAE
†Family PROTASPIDIDAE †Family OMALODONTIDAE
†Family PROTOPTERASPIDIDAE †Order ANTARCTILAMNIFORMES
†Family PSAMMOSTEIDAE Incertae sedis stem chondrichthyan taxa:
†Family PTERASPIDIDAE
†Mongolepidida
†Family KANNATHALEPIDIDAE
1
FISHES
†Family WELLINGTONELLIDAE Family HETERODONTIDAE (8) BULLHEAD SHARKS
†Pupacampella Order ORECTOLOBIFORMES (5) CARPET SHARKS
†Plesioselachus Suborder Parascyllioidei
†Gogoselachus Family PARASCYLLIIDAE (9) COLLARED CARPET SHARKS
†Family BANDRINGIDAE Suborder Orectoloboidei
†Order PHOEBODONTIFORMES Superfamily Orectoloboidea
†Family PHOEBODONTIDAE Family BRACHAELURIDAE (10) BLIND SHARKS
†Family JALODONTIDAE Family ORECTOLOBIDAE (11) WOBBEGONGS
†Superorder CLADOSELACHIMORPHA Superfamily Hemiscyllioidea
†Order CLADOSELACHIFORMES Family HEMISCYLLIIDAE (12) BAMBOO SHARKS
†Family CLADOSELACHIDAE Family GINGLYMOSTOMATIDAE (13) NURSE SHARKS
†Order SYMMORIIFORMES Family STEGOSTOMATIDAE (14) ZEBRA SHARKS
†Family SYMMORIIDAE Family RHINCODONTIDAE (15) WHALE SHARKS
†Family FALCATIDAE Order LAMNIFORMES (6) MACKEREL SHARKS
†Superorder CTENACANTHIMORPHA †Family OTODONTIDAE
†Order CTENACANTHIFORMES †Family XIPHODOLAMIIDAE
†Family CTENACANTHIDAE †Family CARDABIODONTIDAE
†Order SQUATINACTIFORMES †Family CRETOXYRHINIDAE
†Family SQUATINACTIDAE
†Family ARCHAEOLAMNIDAE
†Superorder XENACANTHIMORPHA †Family PSEUDOSCAPANORHYNCHIDAE
†Order BRANSONELLIFORMES
†Family ANACORACIDAE
†Order XENACANTHIFORMES
†Family PSEUDOCORACIDAE
†Family DIPLODOSELACHIDAE
Family MITSUKURINIDAE (16) GOBLIN SHARKS
†Family XENACANTHIDAE
Family ODONTASPIDIDAE (17) SAND TIGER SHARKS
Crown-group chondrichthyans in two subclasses:
Family PSEUDOCARCHARIIDAE (18) CROCODILE SHARKS
Subclass HOLOCEPHALI
Family ALOPIIDAE (19) THRESHER SHARKS
†Order INIOPTERYGIFORMES
†Order ORODONTIFORMES Family MEGACHASMIDAE (20) MEGAMOUTH SHARKS
†Order EUGENEODONTIFORMES Family CETORHINIDAE (21) BASKING SHARKS
†Order PETALODONTIFORMES Family LAMNIDAE (22) MACKEREL SHARKS
†Order DEBEERIIFORMES Order CARCHARHINIFORMES (7) GROUND SHARKS
†Order HELODONTIFORMES Family SCYLIORHINIDAE (23) CAT SHARKS
Superorder HOLOCEPHALIMORPHA Family PROSCYLLIIDAE (24) FINBACK CAT SHARKS
†Order PSAMMODONTIFORMES Family PSEUDOTRIAKIDAE (25) FALSE CAT SHARKS
†Order COPODONTIFORMES Family LEPTOCHARIIDAE (26) BARBELED HOUND SHARKS
†Order SQUALORAJIFORMES Family TRIAKIDAE (27) HOUND SHARKS
†Order CHONDRENCHELYIFORMES Family HEMIGALEIDAE (28) WEASEL SHARKS
†Order MENASPIFORMES Family CARCHARHINIDAE (29) REQUIEM SHARKS
†Order COCHLIODONTIFORMES Family SPHYRNIDAE (30) HAMMERHEAD SHARKS
Order CHIMAERIFORMES (3) CHIMAERAS Superorder SQUALOMORPHI
†Suborder Echinochimaeroidei Series Hexanchida
†Family ECHINOCHIMAERIDAE Order HEXANCHIFORMES (Notidanoidei) (8) SIX-GILL SHARKS
†Suborder Myriacanthoidei Family CHLAMYDOSELACHIDAE (31) FRILLED SHARKS
†Family CHIMAEROPSIDAE Family HEXANCHIDAE (32) COW SHARKS
†Family MYRIACANTHIDAE Series Squalida
Suborder Chimaeroidei Order SQUALIFORMES (9) DOGFISH SHARKS
Superfamily Callorhinchoidea Family CENTROPHORIDAE (33) GULPER SHARKS
Family CALLORHINCHIDAE (5) PLOWNOSE CHIMAERAS Family ETMOPTERIDAE (34) LANTERN SHARKS
Superfamily Chimaeroidea Family SOMNIOSIDAE (35) SLEEPER SHARKS
Family RHINOCHIMAERIDAE (6) LONGNOSE CHIMAERAS Family OXYNOTIDAE (36) ROUGH SHARKS
Family CHIMAERIDAE (7) SHORTNOSE CHIMAERAS / RATFISHES Family DALATIIDAE (37) KITEFIN SHARKS
Subclass EUSELACHII SHARKS, RAYS, RELATED FOSSILS Family SQUALIDAE (38) DOGFISH SHARKS
†Acronemus Series Squatinida
†Order PROTACRODONTIFORMES †Order PROTOSPINACIFORMES
†Infraclass HYBODONTA Order ECHINORHINIFORMES (10) BRAMBLE SHARKS
†Order HYBODONTIFORMES Family ECHINORHINIDAE (39) BRAMBLE SHARKS
†Family HYBODONTIDAE Order SQUATINIFORMES (11) ANGEL SHARKS
†Family TRISTYCHIIDAE Family SQUATINIDAE (40) ANGEL SHARKS
†Family DISTOBATIDAE Order PRISTIOPHORIFORMES (12) SAW SHARKS
†Family ACRODONTIDAE Family PRISTIOPHORIDAE (41) SAW SHARKS
†Family POLYACRODONTIDAE Division BATOMORPHI RAYS
†Family LONCHIDIIDAE Order TORPEDINIFORMES (13) ELECTRIC RAYS
†Family STEINBACHODONTIDAE †Family ARCHAEOBATIDAE
†Family PSEUDODALATIIDAE Family TORPEDINIDAE (42) TORPEDO ELECTRIC RAYS
†Family PTYCHODONTIDAE Family NARCINIDAE (43) NUMBFISHES
†Family HOMALODONTIDAE Subfamily Narcininae
Infraclass ELASMOBRANCHII Subfamily Narkinae
Division SELACHII SHARKS Order RAJIFORMES (14) SKATES
Superorder GALEOMORPHI †Family CYCLOBATIDAE
†Order SYNECHODONTIFORMES
Family RAJIDAE (44) SKATES
†Family PALAEOSPINACIDAE
Subfamily Rajinae
Order HETERODONTIFORMES (4) BULLHEAD SHARKS
2
FISHES
Arhynchobatinae
Subfamily †Family HOLOPTYCHIIDAE
Order PRISTIFORMES (15) GUITARFISHES, SAWFISHES Superorder DIPNOI
†Family SCLERORHYNCHIDAE Early, stem-group dipnoans include:
†Family PTYCHOTRYGONIDAE †Youngolepis, †Powichthys, †Glyptolepis
Family “RHINOBATIDAE” (45) GUITARFISHES †Order DIABOLEPIDIFORMES
Family RHINIDAE (46) BOWMOUTH GUITARFISHES †Family DIABOLEPIDIDAE
Family RHYNCHOBATIDAE (47) WEDGEFISHES †Order DIPNORHYNCHIFORMES
Family PRISTIDAE (48) SAWFISHES †Family URANOLOPHIDAE
Order MYLIOBATIFORMES (16) STINGRAYS †Family DIPNORHYNCHIDAE
Suborder Platyrhinoidei †Family CHIRODIPTERIDAE
Family PLATYRHINIDAE (49) THORNBACKS †Order DIPTERIFORMES
Suborder Myliobatoidei †Family STOMIAHYKIDAE
Family ZANOBATIDAE (50) PANRAYS †Family DIPTERIDAE
Family PLESIOBATIDAE (51) DEEPWATER STINGRAYS †Family CHIRODIPTERIDAE
Family UROLOPHIDAE (52) ROUND STINGRAYS †Family RHYNCHODIPTERIDAE
Family HEXATRYGONIDAE (53) SIXGILL STINGRAYS †Family PHANEROPLEURIDAE
Family DASYATIDAE (Trygonidae) (54) WHIPTAIL STINGRAYS †Family FLEURANTIIDAE
Family POTAMOTRYGONIDAE (55) RIVER STINGRAYS †Order CTENODONTIFORMES
Family GYMNURIDAE (56) BUTTERFLY RAYS †Family URONEMIDAE
Family UROTRYGONIDAE (57) AMERICAN ROUND STINGRAYS †Family CTENODONTIDAE
Family MYLIOBATIDAE (58) EAGLE RAYS Order CERATODONTIFORMES (18) LIVING LUNGFISHES & FOSSILS
Subfamily Myliobatinae †Family ARGANODONTIDAE
Subfamily Rhinopterinae †Family CERATODONTIDAE
Subfamily Mobulinae †Family ASIATOCERATODONTIDAE
Family LEPIDOSIRENIDAE (61) SOUTH AMERICAN LUNGFISHES
GRADE TELEOSTOMI Family PROTOPTERIDAE (62) AFRICAN LUNGFISHES
†Class ACANTHODII ACANTHODIANS †Infraclass RHIZODONTIDA
Incertae sedis early-branching acanthodians include: †Order RHIZODONTIFORMES
†Lupopsyroides, †Lupopsyrus †Family RHIZODONTIDAE
†Order CLIMATIIFORMES †Infraclass OSTEOLEPIDIDA
†Family BROCHOADMONIDAE †Order OSTEOLEPIDIFORMES
†Family CLIMATIIDAE †Family CANOWINDRIDAE
†Family GYRACANTHIDAE †Family MEGALICHTHYIDAE
†Family EUTHACANTHIDAE †Family OSTEOLEPIDIDAE
†Order DIPLACANTHIFORMES †Family TRISTICHOPTERIDAE
†Family CULMACANTHIDAE †Family RHIZODOPSIDAE
†Family DIPLACANTHIDAE †Infraclass ELPISTOSTEGALIA
†Family GLADIOBRANCHIDAE †Order ELPISTOSTEGALIFORMES
†Order ISCHNACANTHIFORMES †Family ELPISTOSTEGALIDAE
†Family ISCHNACANTHIDAE Infraclass TETRAPODA tetrapods
†Family PORACANTHODIDAE Subclass ACTINOPTERYGII RAY-FINNED FISHES
†Order ACANTHODIFORMES Early, stem-group actinopterygians include:
†Family MESACANTHIDAE †Family HAPLOLEPIDAE
†Family CHEIRACANTHIDAE †Coccolepis
†Family ACANTHODIDAE †Order CHEIROLEPIDIFORMES
Class OSTEICHTHYES BONY FISHES, TETRAPODS †Order PALAEONISCIFORMES
Stem-group osteichthyans include: †Family AEDUELLIDAE
†Lophosteus, †Andreolepis †Family ACROLEPIDAE
†Order DIALIPINIFORMES †Family AMBLYPTERIDAE
Subclass SARCOPTERYGII LOBE-FINNED FISHES, TETRAPODS †Family BIRGERIIDAE
Stem-group sarcopterygians include: †Family COMMENTRYIDAE
†Psarolepis, †Achoania, †Guiyu †Family ELONICHTHYIDAE
Infraclass ACTINISTIA COELACANTHS (Coelacanthida) †Family PALAEONISCIDAE
Order COELACANTHIFORMES (17) COELACANTHS †Family PYGOPTERIDAE
†Family MIGUASHAIIDAE †Family RHABDOLEPIDAE
†Family DIPLOCERCIDAE †Family RHADINICHTHYIDAE
†Family HADRONECTORIDAE †Family AESOPICHTHYIDAE
†Family RHABDODERMATIDAE †Family STEGOTRACHELIDAE
†Family LAUGIIDAE †Order DORYPTERIFORMES
†Family WHITEIIDAE †Family DORYPTERIDAE
†Family REBELLATRICIDAE †Order PLATYSOMIFORMES
†Family COELACANTHIDAE †Family PLATYSOMIDAE
†Family MAWSONIIDAE †Family BOBASATRANIIDAE
Family LATIMERIIDAE (59) GOMBESSAS / COELACANTHS †Order TARRASIIFORMES
Infraclass ONYCHODONTIDA †Family TARRASIIDAE
†Order ONYCHODONTIFORMES †Order GUILDAYICHTHYIFORMES
†Family ONYCHODONTIDAE †Family GUILDAYICHTHYIDAE
Infraclass DIPNOMORPHA †Order PHANERORHYNCHIFORMES
†Superorder POROLEPIMORPHA †Family PHANERORHYNCHIDAE
†Order POROLEPIFORMES †Order SAURICHTHYIFORMES
†Family POROLEPIDAE †Family SAURICHTHYIDAE
3
FISHES
†Order REDFIELDIIFORMES †Family PACHYCORMIDAE
†Family REDFIELDIIDAE Division TELEOSTEI
†Order PTYCHOLEPIDIFORMES †Order PHOLIDOPHORIFORMES
†Family PTYCHOLEPIDIDAE †Family PHOLIDOPHORIDAE
†Order PHOLIDOPLEURIFORMES †Order DORSETICHTHYIFORMES
†Family PHOLIDOPLEURIDAE †Family DORSETICHTHYIDAE
†Order PERLEIDIFORMES †Order LEPTOLEPIDIFORMES
†Family CEPHALOXENIDAE †Family LEPTOLEPIDIDAE
†Family CLEITHROLEPIDIDAE †Order CROSSOGNATHIFORMES
†Family COLOBODONTIDAE †Family VARASICHTHYIDAE
†Family PERLEIDIDAE †Family CROSSOGNATHIDAE
†Family PLATYSIAGIDAE †Family NOTELOPIDAE
†Order LUGANOIFORMES †Family PACHYRHIZODONTIDAE
†Family LUGANOIDAE †Order ICHTHYODECTIFORMES
†Order PERLEIDIFORMES †Family ALLOTHRISSOPIDAE
†Family CEPHALOXENIDAE †Family CLADOCYCLIDAE
†Family COLOBODONTIDAE †Family SAURODONTIDAE
†Family PLATYSIAGIDAE †Family ICHTHYODECTIDAE
†Family PELTOPLEURIDAE †Order TSELFATIIFORMES
†Family CLEITHROLEPIDAE †Family PLETHODIDAE
†Family PERLEIDIDAE †Family PROTOBRAMIDAE
†Family THORACOPTERIDAE †Family TSELFATIIDAE
Infraclass CLADISTIA †Order ARARIPICHTHYIFORMES
Order POLYPTERIFORMES (19) BICHIRS †Family ARARIPICHTHYIDAE
Family POLYPTERIDAE (63) BICHIRS Supercohort TELEOCEPHALA CROWN-GROUP TELEOSTEI
Infraclass CHONDROSTEI Cohort ELOPOMORPHA
†Order CHONDROSTEIFORMES Order ELOPIFORMES (23) TENPOUNDERS
†Family CHONDROSTEIDAE Family ELOPIDAE (68) TENPOUNDERS (LADYFISHES)
Order ACIPENSERIFORMES (20) PADDLEFISHES, STURGEONS, Family MEGALOPIDAE (69) TARPONS
†Suborder Peipiaosteoidei Order ALBULIFORMES (24) BONEFISHES
†Family PEIPIAOSTEIDAE Family ALBULIDAE (70)
Suborder Acipenseroidei Order NOTACANTHIFORMES (25) HALOSAURS, DEEP-SEA SPINY EELS
Family POLYODONTIDAE (64) PADDLEFISHES Family HALOSAURIDAE (71) HALOSAURS
†Subfamily Protosephurinae Family NOTACANTHIDAE (72) DEEP-SEA SPINY EELS
Subfamily Polyodontinae Order ANGUILLIFORMES (26) EELS
Family ACIPENSERIDAE (65) STURGEONS Suborder Protanguilloidei
Subfamily Acipenserinae Family PROTANGUILLIDAE (74) PRIMITIVE CAVE EELS
Subfamily Scaphirhynchinae Suborder Synaphobranchoidei
Subfamily Pseudoscaphirhynchinae Family SYNAPHOBRANCHIDAE (74) CUTTHROAT EELS
Subfamily Husinae Subfamily Simenchelyinae
NEOPTERYGII Subfamily Ilyophinae
†Order PYCNODONTIFORMES Subfamily Synaphobranchinae
Infraclass HOLOSTEI GARS, BOWFINS, RELATIVES Suborder Muraenoidei
Division GINGLYMODI Family HETERENCHELYIDAE (75) MUD EELS
†Order DAPEDIIFORMES Family MYROCONGRIDAE (76) MYROCONGER EELS
†Family DAPEDIIDAE Family MURAENIDAE (77) MORAY EELS
Order LEPISOSTEIFORMES (21) GARS Subfamily Uropteryginae
†Family OBAICHTHYIDAE Subfamily Muraeninae
Family LEPISOSTEIDAE (66) GARS Suborder Chlopsoidei
†Order SEMIONOTIFORMES Family CHLOPSIDAE (78) FALSE MORAYS
Incertae sedis in Semionotiformes: †Sangiorgioichthys Suborder Congroidei
†Family SEMIONOTIDAE Family DERICHTHYIDAE (79) LONGNECK / NARROWNECK, SHORTTAIL EELS
†Family CALLIPURBECKIDAE Family OPHICHTHIDAE (80) SNAKE EELS, WORM EELS
†Order MACROSEMIIFORMES Subfamily Myrophinae
†Family MACROSEMIIDAE Subfamily Ophichthinae
Division HALECOMORPHI Family MURAENESOCIDAE (81) PIKE CONGERS
†Order PARASEMIONOTIFORMES Family NETTASTOMATIDAE (82) DUCKBILL EELS
†Order IONOSCOPIFORMES Family CONGRIDAE (83) CONGER EELS
Order AMIIFORMES (22) BOWFINS Subfamily Congrinae
†Suborder Caturoidei Subfamily Heterocongrinae
†Family CATURIDAE Subfamily Bathymyrinae
†Suborder Amioidei Suborder Moringuoidei
†Family LIODESMIDAE Family MORINGUIDAE (84) SPAGHETTI EELS
†Family SINAMIIDAE Suborder Saccopharyngoidei
Family AMIIDAE (67) BOWFINS Family CYEMATIDAE (85) BOBTAIL SNIPE EELS
Infraclass TELEOSTEOMORPHA Family MONOGNATHIDAE (86)
Oldest, most primitive teleosteomorph: †Prohalecites Family SACCOPHARYNGIDAE (87) SWALLOWERS
†Division ASPIDORHYNCHEI Family EURYPHARYNGIDAE (88) GULPERS / PELICAN EELS
†Order ASPIDORHYNCHIFORMES
Suborder Anguilloidei
†Family ASPIDORHYNCHIDAE
Family NEMICHTHYIDAE (89) SNIPE EELS
†Order PACHYCORMIFORMES
Family SERRIVOMERIDAE (90) SAWTOOTH EELS
4
FISHES
Family ANGUILLIDAE (91) FRESHWATER EELS Subfamily Cyprininae
OSTEOGLOSSOCEPHALA Subfamily Leuciscinae
Cohort OSTEOGLOSSOMORPHA Subfamily Acheilognathinae
†Order LYCOPTERIFORMES Subfamily Gobioninae
†Family LYCOPTERIDAE Subfamily Tincinae
Order HIODONTIFORMES (27) MOONEYES Subfamily Opsariichthyinae
Family HIODONTIDAE (92) MOONEYES Subfamily Cultrinae
Order OSTEOGLOSSIFORMES (28) BONYTONGUES Subfamily Squaliobarbinae
Family PANTODONTIDAE (93) BUTTERFLYFISHEs Subfamily Xenocyprinae
Family OSTEOGLOSSIDAE (94) OSTEOGLOSSIDS / BONYTONGUES Family PSILORHYNCHIDAE (110) MOUNTAIN CARPS
Subfamily Heterotidinae Superfamily Cobitoidea
Subfamily Osteoglossinae †Family JIANGHANICHTHYIDAE
Family NOTOPTERIDAE (95) Family GYRINOCHEILIDAE (111) ALGAE EATERS
Subfamily Notopterinae Family CATOSTOMIDAE (112) SUCKERS
Subfamily Xenomystinae Subfamily Myxocyprininae
Family MORMYRIDAE (96) ELEPHANTFISHES Subfamily Cycleptinae
Family GYMNARCHIDAE (97) ABAS Subfamily Ictiobinae
Cohort CLUPEOCEPHALA Subfamily Catostominae
OTOCEPHALA Tribe Erimyzontini
Superorder CLUPEOMORPHA Tribe Catostomini
†Order ELLIMMICHTHYIFORMES Tribe Thoburniini
†Suborder Armigatoidei Tribe Moxostomatini
†Family ARMIGATIDAE Family BOTIIDAE (113) BOTIID LOACHES
†Suborder Ellimichthyoidei Family VAILLANTELLIDAE (114) LONG-FIN LOACHES
†Family SORBINICHTHYIDAE Family COBITIDAE (115) LOACHES
†Family PARACLUPEIDAE Family BALITORIDAE (116) HILLSTREAM / RIVER LOACHES
†Subfamily Scutatospinosinae Family GASTROMYZONTIDAE (117) SUCKER LOACHES
†Subfamily Thorectichthyinae Family NEMACHEILIDAE (118) STONE LOACHES
†Subfamily Ellimminae Family BARBUCCIDAE (120) FIRE-EYED LOACHES
†Subfamily Paraclupeinae Family ELLOPOSTOMATIDAE (119) STURGEON-MOUTHED LOACHES
Order CLUPEIFORMES (29) HERRINGS Family SERPENTICOBITIDAE (121) SERPENT LOACHES
Suborder Denticipitoidei Subseries Characiphysi
Family DENTICIPITIDAE (98) DENTICLE HERRINGS, Order CHARACIFORMES (33) CHARACINS
Suborder Clupeoidei Suborder Citharinoidei
Family PRISTIGASTERIDAE (99) LONGFIN HERRINGS Family DISTICHODONTIDAE (122) DISTICHODONTIDS
Subfamily Pelloninae Family CITHARINIDAE (123) CITHARINIDS
Subfamily Pristigasterinae Suborder Characoidei
Family ENGRAULIDAE (100) ANCHOVIES Superfamily Crenuchoidea
Subfamily Coiliinae Family CRENUCHIDAE (124) SOUTH AMERICAN DARTERS
Subfamily Engraulinae Subfamily Crenuchinae
Family CHIROCENTRIDAE (101) WOLF HERRINGS Subfamily Characidiinae
Family CLUPEIDAE (102) HERRINGS Superfamily Alestoidea
(SHADS, SPRATS, SARDINES, PILCHARDS, MENHADENS)
Family ALESTIDAE (125) AFRICAN TETRAS
Subfamily Dussumieriinae
Family HEPSETIDAE (126) AFRICAN PIKES
Subfamily Ehiravinae
Superfamily Erythrinoidea
Subfamily Pellonulinae
Family ERYTHRINIDAE (127) TRAHIRAS
Subfamily Clupeinae
Family PARODONTIDAE (128) PARODONTIDS
Subfamily Alosinae
Family CYNODONTIDAE (129) CYNODONTIDS
Subfamily Dorosomatinae
Family SERRASALMIDAE (130) PACUS, SILVER DOLLARS, PIRANHAS
Superorder ALEPOCEPHALI
Family HEMIODONTIDAE (131) HEMIODONTIDS
Order ALEPOCEPHALIFORMES (30) – SLICKHEADS, TUBESHOULDERS
Subfamily Anodontinae
Family PLATYTROCTIDAE (103) TUBESHOULDERS
Subfamily Hemiodontinae
Family BATHYLACONIDAE (104) BATHYLACONIDS
Tribe Hemiodontini
Family ALEPOCEPHALIDAE (105) SLICKHEADS
Tribe Bibibranchiini
Superorder OSTARIOPHYSI
Family ANOSTOMIDAE (132) TOOTHED HEADSTANDERS
Series Anotophysi
Family CHILODONTIDAE (133) HEADSTANDERS
Order GONORYNCHIFORMES (31) MILKFISHES
Family CURIMATIDAE (134) TOOTHLESS CHARACIFORMS
Family CHANIDAE (106) MILKFISHES
Family PROCHILODONTIDAE (135) FLANNEL-MOUTH CHARACIFORMS
†Subfamily Rubiesichthyinae
Superfamily Characoidea
Subfamily Chaninae
Family LEBIASINIDAE (136) PENCIL FISHES
Family GONORYNCHIDAE (107) BEAKED SANDFISHES
Subfamily Lebiasininae
Family KNERIIDAE (108) KNERIAS, SNAKE MUDHEADS
Subfamily Pyrrhulininae
Subfamily Kneriinae
Family CTENOLUCIIDAE (137) PIKE-CHARACIDS
Subfamily Phractolaeminae
Family ACESTRORHYNCHIDAE (138) ACESTRORHYNCHIDS
Series Otophysi
Subfamily Acestrorhynchinae
Subseries Cypriniphysi
Subfamily Heterocharacinae
Order CYPRINIFORMES (32) CARPS, LOACHES, MINNOWS, RELATIVES
Subfamily Roestinae
Superfamily Cyprinoidea
Family CYPRINIDAE (109) MINNOWS, CARPS, LOACHES Family CHARACIDAE (139) CHARACINS
Subfamily Iguanodectinae
Subfamily Danioninae
Subfamily Agoniatinae
Subfamily Labeoninae
5
FISHES
Subfamily Clupeacharacinae Family MALAPTERURIDAE (163) ELECTRIC CATFISHES
Subfamily Bryconinae Family MOCHOKIDAE (164) SQUEAKERS / UPSIDE-DOWN CATFISHES
Subfamily Stethaprioninae Family SCHILBEIDAE (165) SCHILBEID / BUTTER CATFISHES
Subfamily Rhoadsiinae Family AUCHENOGLANIDIDAE (166) AUCHENOGLANIDIDS
Subfamily Characinae Family CLAROTEIDAE (167) CLAROTEIDS
Subfamily Tetragonopterinae Family LACANTUNIIDAE (168) CHIAPAS CATFISHES
Subfamily Aphyocharacinae Superfamily Clarioidea
Subfamily Aphyoditeinae Family CLARIIDAE (169) AIRBREATHING CATFISHES
Subfamily Cheirodontinae Family HETEROPNEUSTIDAE (170) AIRSAC CATFISHES
Subfamily Gymnocharacinae Superfamily Arioidea
Subfamily Stevardiinae Family ANCHARIIDAE (171) MALAGASY CATFISHES
Family GASTEROPELECIDAE (140) FRESHWATER HATCHETFISHES Family ARIIDAE (172) SEA CATFISHES
Subseries Siluriphysi Superfamily Doradoidea
Order SILURIFORMES (34) CATFISHES Family ASPREDINIDAE (173) BANJO CATFISHES
†Family ANDINICHTHYIDAE Subfamily Bunocephalinae
Suborder Diplomystoidei Subfamily Aspredininae
†Family BACHMANNIIDAE Subfamily Hoplomyzontinae
Family DIPLOMYSTIDAE (141) VELVET CATFISHES Family DORADIDAE (174) THORNY CATFISHES
Suborder Hypsidoroidei Family AUCHENIPTERIDAE (175) DRIFTWOOD CATFISHES
†Family HYPSIDORIDAE Subfamily Auchenipterinae
Family CETOPSIDAE (142) WHALE CATFISHES Subfamily Centromochlinae
Subfamily Helogeneinae Superfamily Ictaluroidea
Subfamily Cetopsinae Family CRANOGLANIDIDAE (176) ARMORHEAD CATFISHES
Suborder Loricarioidei Family ICTALURIDAE (177) NORTH AMERICAN CATFISHES
Family TRICHOMYCTERIDAE (143) – PENCIL / PARASITIC CATFISHES Superfamily Pimelodoidea
Subfamily Copionodontinae Family HEPTAPTERIDAE (178) HEPTAPTERIDS
Subfamily Trichogeninae Family PIMELODIDAE (179) LONG-WHISKERED CATFISHES
Subfamily Trichomycterinae Family PSEUDOPIMELODIDAE (180) BUMBLEBEE CATFISHES
Subfamily Vandelliinae Order GYMNOTIFORMES (35) NEOTROPICAL KNIFEFISHES
Subfamily Stegophilinae Suborder Gymnotoidei
Subfamily Tridentinae Family GYMNOTIDAE (181) NAKEDBACK KNIFEFISHES
Subfamily Glanapteryginae Suborder Sternopygoidei
Subfamily Sarcoglanidinae Superfamily Rhamphichthyoidea
Family NEMATOGENYIDAE (144) MOUNTAIN CATFISHES Family RHAMPHICHTHYIDAE (182) SAND KNIFEFISHES
Family CALLICHTHYIDAE (145) CALLICHTHYID ARMORED CATFISHES Family HYPOPOMIDAE (183) BLUNTNOSE KNIFEFISHES
Subfamily Callichthyinae Superfamily Apteronotoidea
Subfamily Corydoradinae Family STERNOPYGIDAE (184) GLASS KNIFEFISHES
Family SCOLOPLACIDAE (146) SPINY DWARF CATFISHES Family APTERONOTIDAE (185) GHOST KNIFEFISHES
Family ASTROBLEPIDAE (147) CLIMBING CATFISHES EUTELEOSTEI
Family LORICARIIDAE (148) SUCKERMOUTH ARMORED CATFISHES Order LEPIDOGALAXIIFORMES (36) SALAMANDERFISHES
Subfamily Lithogeneinae Family LEPIDOGALAXIIDAE (186) SALAMANDERFISHES
Subfamily Neoplecostominae Superorder PROTACANTHOPTERYGII
Subfamily Hypoptopomatinae Order SALMONIFORMES (37) TROUT, SALMON, WHITEFISH
Subfamily Loricariinae Family SALMONIDAE (187) TROUT, SALMON, WHITEFISH
Subfamily Ancistrinae Subfamily Coregoninae
Subfamily Hypostominae Subfamily Thymallinae
Subfamily Delturinae Subfamily Salmoninae
Suborder Siluroidei Order ESOCIFORMES (38) PIKES
Family SILURIDAE (149) SHEATFISHES †Family PALAEOESOCIDAE
Family AUSTROGLANIDIDAE (150) ROCK CATFISHES Family ESOCIDAE (188) PIKES
Family PANGASIIDAE (151) SHARK CATFISHES Family UMBRIDAE (189) MUDMINNOWS
Family CHACIDAE (152) SQUAREHEAD, ANGLER, / FROGMOUTH CATFISHES ZOROTELEOSTEI
Family PLOTOSIDAE (153) EELTAIL CATFISHES Superorder OSMEROMORPHA
Family RITIDAE (154) RITAS, NANOBAGRIDS Order ARGENTINIFORMES (39) MARINE SMELTS
Family AILIIDAE (155) ASIAN “SCHILBEIDS” Family ARGENTINIDAE (190) ARGENTINES / HERRING SMELTS
Family HORABAGRIDAE (156) IMPERIAL / SUN CATFISHES Family OPISTHOPROCTIDAE (191) BARRELEYES / SPOOKFISHES
Family BAGRIDAE (157) BAGRID CATFISHES Family MICROSTOMATIDAE (192) PENCILSMELTS
Superfamily Sisoroidea Family BATHYLAGIDAE (193) DEEPSEA SMELTS
Family AKYSIDAE (158) STREAM CATFISHES Order GALAXIIFORMES (40) GALAXIIFORMS
Subfamily Akysinae Family GALAXIIDAE (194) GALAXIIDS
Subfamily Parakysinae Subfamily Galaxiinae
Family AMBLYCIPITIDAE (159) TORRENT CATFISHES Subfamily Lovettiinae
Family SISORIDAE (160) SISORID CATFISHES Subfamily Aplochitoninae
Subfamily Sisorinae Order OSMERIFORMES (41) FRESHWATER SMELTS
Subfamily Glyptosterninae Suborder Osmeroidei
Family ERETHISTIDAE (161) ERETHISTID CATFISHES Family OSMERIDAE (195) NORTHERN HEMISPHERE SMELTS
Family AMPHILIIDAE (162) LOACH CATFISHES Family PLECOGLOSSIDAE (196) AYU / SWEETFISH
Subfamily Amphiliinae Family SALANGIDAE (197) ICEFISHES / NOODLEFISHES
Subfamily Leptoglaninae Suborder Retropinnoidei
Subfamily Doumeinae Family PROTOTROCTIDAE (198) SOUTHERN GRAYLINGS

6
FISHES
Family RETROPINNIDAE (199) SOUTHERN SMELTS Family LOPHOTIDAE (225) CRESTFISHES
Order STOMIIFORMES (42) DRAGONFISHES Family RADIICEPHALIDAE (226) TAPERTAILS
Suborder Gonostomatoidei Family TRACHIPTERIDAE (227) RIBBONFISHES
Family GONOSTOMATIDAE (200) BRISTLEMOUTHS Family REGALECIDAE (228) OARFISHES
Family STERNOPTYCHIDAE (201) MARINE HATCHETFISHES EUACANTHOMORPHA
Subfamily Maurolicinae Superorder PARACANTHOPTERYGII
Subfamily Sternoptychinae Order POLYMIXIIFORMES (47) BEARDFISHES
Family PHOSICHTHYIDAE (202) LIGHTFISHES Family POLYMIXIIDAE (229) BEARDFISHES
Family STOMIIDAE (203) BARBELED DRAGONFISHES †Order SPHENOCEPHALIFORMES
Subfamily Stomiinae †Family SPHENOCEPHALIDAE
Tribe Stomiini Order PERCOPSIFORMES (48) TROUT-PERCHES
Tribe Chauliodontini Family PERCOPSIDAE (230) TROUT-PERCHES
Subfamily Melanostomiinae †Family MCCONICHTHYIDAE
Subfamily Idiacanthinae Family APHREDODERIDAE (231) PIRATE PERCHES
Subfamily Malacosteinae Family AMBLYOPSIDAE (232) CAVEFISHES
NEOTELEOSTEI Order ZEIFORMES (49) DORIES
Superorder ATELEOPODOMORPHA †Family CRETAZEIDAE
Order ATELEOPODIFORMES (43) JELLYNOSE FISHES Suborder Cyttoidei
Family ATELEOPODIDAE (204) JELLYNOSE FISHES Family CYTTIDAE (233) LOOKDOWN DORIES
EURYPTERYGII Suborder Zeoidei
Superorder CYCLOSQUAMATA Family OREOSOMATIDAE (234) OREOS
Order AULOPIFORMES (44) LIZARDFISHES Family PARAZENIDAE (235) SMOOTH DORIES
†Suborder Ichthyotringoidei Subfamily Parazeninae
†Family ICHTHYOTRINGIDAE Subfamily Cyttopsinae
†Family DERCETIDAE Family ZENIONTIDAE (236) ARMOREYE DORIES
†Family PRIONOLEPIDIDAE Family GRAMMICOLEPIDIDAE (237) TINSELFISHES
†Suborder Halecoidei Subfamily Macrurocyttinae
†Family HALECIDAE Subfamily Grammicolepidinae
Suborder Aulopoidei Family ZEIDAE (238) DORIES
Family SYNODONTIDAE (205) LIZARDFISHES Order STYLEPHORIFORMES (50) TUBE-EYES / THREAD-TAILS
Subfamily Synodontinae Family STYLEPHORIDAE (239) TUBE-EYES / THREAD-TAILS
Subfamily Harpadontinae Order GADIFORMES (51) CODS, HAKES
Family AULOPIDAE (206) FLAGFINS Suborder Melanonoidei
Family PSEUDOTRICHONOTIDAE (207) SANDIVING LIZARDFISHES Family MELANONIDAE (240) PELAGIC CODS
Suborder Paraulopoidei Suborder Macruroidei
Family PARAULOPIDAE (208) CUCUMBER FISHES Family STEINDACHNERIIDAE (241) LUMINOUS, SOUTHERN HAKES
Suborder Alepisauroidei Family BATHYGADIDAE (242) RATTAILS
Superfamily Ipnopoidea Family MACROURIDAE (243) GRENADIERS (RATTAILS)
Family IPNOPIDAE (209) DEEPSEA TRIPOD FISHES Family TRACHYRINCIDAE (244) WHIPTAILS, TRACHYRINCINES
Subfamily Ipnopinae Subfamily Macruroidinae
Subfamily Bathysauropsinae Subfamily Trachyrincinae
Family GIGANTURIDAE (210) TELESCOPEFISHES Suborder Gadoidei
Family BATHYSAUROIDIDAE (211) LARGESCALE DEEP-SEA LIZARDFISH Family EUCLICHTHYIDAE (245) EUCLA COD
Family BATHYSAURIDAE (212) DEEPSEA LIZARDFISHES Family MORIDAE (246) DEEPSEA CODS, CODLINGS, HAKELINGS
Superfamily Chlorophthalmoidea Family MACRURONIDAE (247) SOUTHERN HAKES
Family CHLOROPHTHALMIDAE (213) GREENEYES Family MERLUCCIIDAE (248) MERLUCCIID HAKES
Superfamily Notosudoidea Superfamily Gadoidea
Family NOTOSUDIDAE (214) WARYFISHES Family RANICIPITIDAE (249) TADPOLE CODS
Superfamily Alepisauroidea Family BREGMACEROTIDAE (250) CODLETS
Family SCOPELARCHIDAE (215) PEARLEYES Family MURAENOLEPIDIDAE (251) EEL CODS, MORAY CODS
Family EVERMANNELLIDAE (216) SABERTOOTH FISHES Family GADIDAE (252) CODS
Family SUDIDAE (217) SUDID BARRACUDINAS Subfamily Phycinae
Family PARALEPIDIDAE (218) BARRACUDINAS Subfamily Gaidropsarinae
†Family ENCHODONTIDAE Subfamily Lotinae
†Subfamily Cimolichthyinae Subfamily Gadinae
†Subfamily Enchodontinae Superorder ACANTHOPTERYGII
Family ALEPISAURIDAE (219) LANCETFISHES Series Berycida
Family LESTIDIIDAE (220) NAKED BARRACUDINAS Order HOLOCENTRIFORMES (52) SQUIRRELFISHES
CTENOSQUAMATA Family HOLOCENTRIDAE (253) SQUIRRELFISHES
Superorder SCOPELOMORPHA Subfamily Holocentrinae
Order MYCTOPHIFORMES (45) LANTERNFISHES Subfamily Myripristinae
Family NEOSCOPELIDAE (221) BLACKCHINS Order TRACHICHTHYIFORMES (53) ROUGHIES
Family MYCTOPHIDAE (222) LANTERNFISHES Suborder Anoplogastroidei
Subfamily Myctophinae Family ANOPLOGASTRIDAE (254) FANGTOOTHS
Subfamily Lampanyctinae Family DIRETMIDAE (255) SPINYFINS
ACANTHOMORPHA Suborder Trachichthyoidei
†Order CTENOTHRISSIFORMES Family ANOMALOPIDAE (256) FLASHLIGHT / LANTERNEYE FISHES
Superorder LAMPRIMORPHA †Family PSEUDOMONOCENTRIDAE
Order LAMPRIFORMES (46) OPAHS Family MONOCENTRIDAE (257) PINECONE FISHES
Family VELIFERIDAE (223) VELIFERS Family TRACHICHTHYIDAE (258) ROUGHIES
Family LAMPRIDAE (224) OPAHS
7
FISHES
Order BERYCIFORMES (54) BERYCIFORMS Subfamily Anisochrominae
Suborder Stephanoberycoidei Subfamily Congrogadinae
Superfamily Stephanoberycoidea Family OPISTOGNATHIDAE (290) JAWFISHES
Family GIBBERICHTHYIDAE (259) GIBBERFISHES Order MUGILIFORMES (59) MULLETS
Family STEPHANOBERYCIDAE (260) PRICKLEFISHES Family MUGILIDAE (291) MULLETS
Family HISPIDOBERYCIDAE (261) HISPIDOBERYCIDS Order CICHLIFORMES (60)
Superfamily Cetomimoidea Family CICHLIDAE (292) CICHLIDS
Family RONDELETIIDAE (262) REDMOUTH WHALEFISHES Family PHOLIDICHTHYIDAE (293) CONVICT BLENNY
Family BARBOURISIIDAE (263) RED (REDVELVET) WHALEFISHES Order BLENNIIFORMES (61) BLENNIES
Family CETOMIMIDAE (264) FLABBY WHALEFISHES Family TRIPTERYGIIDAE (294) TRIPLEFIN BLENNIES
Suborder Berycoidei Family DACTYLOSCOPIDAE (295) SAND STARGAZERS
Family MELAMPHAIDAE (265) BIGSCALE FISHES Family BLENNIIDAE (296) COMBTOOTH BLENNIES
Family BERYCIDAE (266) ALFONSINOS Subfamily Salarinae
Series Percomorpha Subfamily Ablenniinae
Subseries Ophidiida Family CLINIDAE (297) KELP BLENNIES
Order OPHIDIIFORMES (55) CUSK-EELS Subfamily Ophiclininae
Suborder Ophidioidei Subfamily Clininae
Family CARAPIDAE (267) PEARLFISHES Subfamily Myxodinae
Subfamily Pyramodontinae Family LABRISOMIDAE (298) LABRISOMID BLENNIES
Subfamily Carapinae Family CHAENOPSIDAE (299) TUBE BLENNIES
Subfamily Tetragondacninae Order GOBIESOCIFORMES (62) CLINGFISHES
Family OPHIDIIDAE (268) CUSK-EELs Family GOBIESOCIDAE (300) CLINGFISHES
Subfamily Brotulinae Subfamily Gobiesocinae
Subfamily Brotulotaeniinae Subfamily Cheilobranchinae
Subfamily Ophidiinae Infraseries Atherinomorpha
Subfamily Neobythitinae Order ATHERINIFORMES (63) SILVERSIDES
Suborder Bythitoidei Suborder Atherinopsoidei
Family BYTHITIDAE (269) VIVIPAROUS BROTULAS Family ATHERINOPSIDAE (301) NEW WORLD SILVERSIDES
Subfamily Bythitinae Subfamily Atherinopsinae
Subfamily Brosmophycinae Tribe Atherinopsini
Family APHYONIDAE (270) APHYONIDS, BLIND CUSK-EEL Tribe Sorgentinini
Family PARABROTULIDAE (271) FALSE BROTULAS Subfamily Menidiinae
Subseries Batrachoidida Tribe Menidiini
Order BATRACHOIDIFORMES (Haplodoci) (56) TOADFISHES Tribe Membradini
Family BATRACHOIDIDAE (272) TOADFISHES Family NOTOCHEIRIDAE (302) SURF SILVERSIDES
Subfamily Porichthyinae Suborder Atherinoidei
Subfamily Thalassophryninae Family ISONIDAE (303) SURF SARDINES
Subfamily Batrachoidinae Family MELANOTAENIIDAE (304) RAINBOWFISHES, BLUE EYES
Subfamily Halophryninae Subfamily Bedotiinae
Subseries Gobiida Subfamily Melanotaeniinae
Order KURTIFORMES (57) NURSERYFISHES, CARDINALFISHES Subfamily Pseudomugilinae
Family KURTIDAE (273) NURSERYFISHES Subfamily Telmatherininae
Family APOGONIDAE (274) CARDINALFISHES Family ATHERIONIDAE (305) PRICKLENOSE SILVERSIDES
Subfamily Apogoninae Family DENTATHERINIDAE (306) MERCER’S TUSKED SILVERSIDE
Subfamily Pseudamiinae Family PHALLOSTETHIDAE (307) PRIAPIUMFISHES
Order GOBIIFORMES (58) GOBIES Subfamily Phallostethinae
Family RHYACICHTHYIDAE (275) LOACH GOBIES Subfamily Gulaphallinae
Family ODONTOBUTIDAE (276) FRESHWATER SLEEPERS Family ATHERINIDAE (308) OLD WORLD SILVERSIDES
Family MILYERINGIDAE (277) BLIND CAVE GOBIES Subfamily Atherinomorinae
Family ELEOTRIDAE (278) SPINYCHEEK SLEEPERS Subfamily Craterocephalinae
Family BUTIDAE (279) BUTID SLEEPERS Subfamily Bleheratherininae
Family THALASSELEOTRIDIDAE (280) OCEAN SLEEPERS Subfamily Atherininae
Family OXUDERCIDAE (281) GOBIONELLUS-LIKE, MUDSKIPPER GOBIES Order BELONIFORMES (64) NEEDLEFISHES
Family GOBIIDAE (282) GOBIES Suborder Adrianichthyoidei
Subseries Ovalentaria Family ADRIANICHTHYIDAE (309) ADRIANICHTHYIDS
Family AMBASSIDAE (Chandidae) (283) ASIATIC GLASSFISHES Subfamily Oryziinae
Family EMBIOTOCIDAE (284) SURFPERCHES Subfamily Adrianichthyinae
Family GRAMMATIDAE (285) BASSLETS Suborder Exocoetoidei
Family PLESIOPIDAE (286) ROUNDHEADS Superfamily Exocoetoidea
Subfamily Plesiopinae Family EXOCOETIDAE (310) FLYINGFISHES
Subfamily Acanthoclininae Subfamily Fodiatorinae
Family POLYCENTRIDAE (287) SOUTH AMERICAN LEAFFISHES Subfamily Parexocoetinae
Family POMACENTRIDAE (288) DAMSELFISHES Subfamily Exocoetinae
Subfamily Amphiprioninae Subfamily Cypselurinae
Subfamily Chrominae Family HEMIRAMPHIDAE (311) HALFBEAKS
Subfamily Lepidozyginae Family ZENARCHOPTERIDAE (312) VIVIPAROUS HALFBEAKS
Subfamily Pomacentrinae Superfamily Scomberesocoidea
Family PSEUDOCHROMIDAE (289) DOTTYBACKS Family BELONIDAE (313) NEEDLEFISHES
Subfamily Pseudochrominae Family SCOMBERESOCIDAE (314) SAURIES
Subfamily Pseudoplesiopinae Order CYPRINODONTIFORMES (65) KILLIFISHES
8
FISHES
Suborder Aplocheiloidei Family CHANNIDAE (340) SNAKEHEADS
Family APLOCHEILIDAE (315) ASIAN RIVULINES Family NANDIDAE (341) ASIAN LEAFFISHES
Family NOTHOBRANCHIIDAE (316) AFRICAN RIVULINES Family BADIDAE (342) CHAMELEONFISHES
Family RIVULIDAE (317) NEW WORLD RIVULINES Family PRISTOLEPIDIDAE (343) (MALAYAN) LEAFFISHEs
Suborder Cyprinodontoidei Order PLEURONECTIFORMES (70) FLATFISHES
Superfamily Funduloidea Suborder Psettodoidei
Family PROFUNDULIDAE (318) MIDDLE AMERICAN KILLIFISHES Family PSETTODIDAE (344) SPINY TURBOTS
Family GOODEIDAE (319) GOODEIDS Suborder Pleuronectoidei
Subfamily Empetrichthyinae Superfamily Citharoidea
Subfamily Goodeinae Family CITHARIDAE (345) LARGESCALE FLOUNDERS
Family FUNDULIDAE (320) TOPMINNOWS Superfamily Pleuronectoidea
Superfamily Valencioidea Family SCOPHTHALMIDAE (346) TURBOTS
Family VALENCIIDAE (321) VALENCIA TOOTHCARPS Family PARALICHTHYIDAE (347) SAND FLOUNDERS
Superfamily Cyprinodontoidea Family PLEURONECTIDAE (348) RIGHTEYE FLOUNDERS
Family CYPRINODONTIDAE (322) PUPFISHES Subfamily Hippoglossinae
Subfamily Cubanichthyinae Subfamily Eopsettinae
Subfamily Cyprinodontinae Subfamily Lyopsettinae
Tribe Orestiini Subfamily Hippoglossoidinae
Tribe Cyprinodontini Subfamily Pleuronectinae
Superfamily Poecilioidea Tribe Psettichthyini
Family ANABLEPIDAE (323) FOUR-EYED FISHES Tribe Isopsettini
Subfamily Anablepinae Tribe Microstomini
Subfamily Oxyzygonectinae Tribe Pleuronectini
Family POECILIIDAE (324) LIVEBEARERS Family BOTHIDAE (349) LEFTEYE FLOUNDERS
Subfamily Aplocheilichthyinae Superfamily Soleoidea
Subfamily Procatopodinae Family PARALICHTHODIDAE (350) MEASLES / PEPPERED FLOUNDERS
Tribe Fluviphylacini Family POECILOPSETTIDAE (351) BIGEYE FLOUNDERS
Tribe Procatopodini Family RHOMBOSOLEIDAE (352) RHOMBOSOLEIDS
Subfamily Poeciliinae Family ACHIROPSETTIDAE (353) SOUTHERN FLOUNDERS
Tribe Alfarini Family SAMARIDAE (354) CRESTED FLOUNDERS
Tribe Priapellini Family ACHIRIDAE (355) AMERICAN SOLES
Tribe Gambusiini Family SOLEIDAE (356) SOLES
Tribe Heterandrini Family CYNOGLOSSIDAE (357) TONGUEFISHES
Tribe Girardini Subfamily Symphurinae
Tribe Poeciliini Subfamily Cynoglossinae
Tribe Cnesterodontini Proposed clade sister to all percomorphs except
Tribe Scolichthyini Ophidiida, Batrachoidida, Gobiida
Tribe Xenodexini Order SYNGNATHIFORMES (71) PIPEFISHES, SEAHORSES
Proposed sister clade to subseries Ovalentaria Suborder Syngnathoidei
Order SYNBRANCHIFORMES (66) SWAMP EELS Superfamily Pegasoidea
Suborder Synbranchoidei Family PEGASIDAE (358) SEAMOTHS
Family SYNBRANCHIDAE (325) SWAMP EELS
Superfamily Syngnathoidea
Suborder Mastacembeloidei
Family SOLENOSTOMIDAE (359) GHOST PIPEFISHES
Family CHAUDHURIIDAE (326) EARTHWORM EELS
Family SYNGNATHIDAE (360) PIPEFISHES, SEAHORSES
Family MASTACEMBELIDAE (327) FRESHWATER SPINY EELS
Subfamily Syngnathinae
Order CARANGIFORMES (67) JACKS
Subfamily Hippocampinae
Family NEMATISTIIDAE (328) ROOSTERFISHES
Suborder Aulostomoidei
Family CORYPHAENIDAE (329) DOLPHINFISHES
Superfamily Aulostomoidea
Family RACHYCENTRIDAE (330) cobias
Family AULOSTOMIDAE (361) TRUMPETFISHES
Family ECHENEIDAE (331) REMORAS (SHARKSUCKERS)
Family FISTULARIIDAE (362) CORNETFISHES
Family CARANGIDAE (332) jACKS, POMPANOS
Superfamily Centriscoidea
Subfamily Trachinotinae
Family MACRORAMPHOSIDAE (363) SNIPEFISHES
Subfamily Scomberoidinae
Family CENTRISCIDAE (364) SHRIMPFISHES
Subfamily Naucratinae
Family DACTYLOPTERIDAE (365) FLYING GURNARDS
Subfamily Caranginae
Order ICOSTEIFORMES (72) RAGFISHES
Family MENIDAE (333) MOONFISHES
Family ICOSTEIDAE (366) RAGFISHES
Order ISTIOPHORIFORMES (68) BARRACUDAS, BILLFISHES Order CALLIONYMIFORMES (73)
Family SPHYRAENIDAE (334) BARRACUDAS Family CALLIONYMIDAE (367) DRAGONETS
Family XIPHIIDAE (335) SWORDFISHES Family DRACONETTIDAE (368) SLOPE DRAGONETS
Family ISTIOPHORIDAE (336) BILLFISHES Order SCOMBROLABRACIFORMES (74) LONGFIN ESCOLARS
Order ANABANTIFORMES (69) LABYRINTH FISHES Family SCOMBROLABRACIDAE (369) LONGFIN ESCOLARS
Suborder Anabantoidei Order SCOMBRIFORMES (75) MACKERELS
Family ANABANTIDAE (337) CLIMBING GOURAMIES
Suborder Scombroidei
Family HELOSTOMATIDAE (338) KISSING GOURAMIES Family GEMPYLIDAE (370) SNAKE MACKERELS
Family OSPHRONEMIDAE (339) GOURAMIES, FIGHTING FISHES Family TRICHIURIDAE (371) CUTLASSFISHES
Subfamily Osphroneminae
Subfamily Aphanopodinae
Subfamily Belontiinae
Subfamily Lepidopodinae
Subfamily Macropodusinae
Subfamily Trichiurinae
Subfamily Luciocephalinae
Family SCOMBRIDAE (372) MACKERELS, TUNAS
Subfamily Trichogastrinae
Subfamily Gasterochismatinae
Suborder Channoidei
9
FISHES
Scombrinae
Subfamily Family DINOPERCIDAE (418) CAVEBASSES
Tribe Scombrini Family BANJOSIDAE (419) BANJOFISHES
Tribe Scomberomorini Family CENTRARCHIDAE (420) SUNFISHES
Tribe Sardini Subfamily Elassomatinae
Tribe Thunnini Subfamily Centrarchinae
Suborder Stromateoidei Subfamily Lepominae
Family AMARSIPIDAE (373) AMARSIPAS Family SERRANIDAE (421) SEA BASSES
Family CENTROLOPHIDAE (374) MEDUSAFISHES Subfamily Serraninae
Family NOMEIDAE (375) DRIFTFISHES Subfamily Anthiinae
Family ARIOMMATIDAE (376) ARIOMMATIDS Subfamily Epinephelinae
Family TETRAGONURIDAE (377) SQUARETAILS Tribe Niphonini
Family STROMATEIDAE (378) BUTTERFISHES Tribe Epinephelini
Proposed clade of all remaining percomorph orders: Tribe Diploprionini
Order TRACHINIFORMES (76) Tribe Liopropomini
Family CHIASMODONTIDAE (379) SWALLOWERS Tribe Grammistini
Family CHAMPSODONTIDAE (380) GAPERS Family PERCIDAE (422) PERCHES
Family PINGUIPEDIDAE (381) SANDPERCHES Subfamily Percinae
Family CHEIMARRICHTHYIDAE (382) NEW ZEALAND TORRENTFISHES Subfamily Acerinae
Family TRICHONOTIDAE (383) SANDDIVERS Subfamily Percarininae
Family CREEDIIDAE (384) SANDBURROWERS Subfamily Luciopercinae
Family PERCOPHIDAE (385) DUCKBILLS Tribe Luciopercini
Subfamily Percophinae Tribe Romanichthyini
Subfamily Bembropinae Subfamily Etheostomatinae
Family LEPTOSCOPIDAE (386) SOUTHERN SANDFISHES Family LACTARIIDAE (423) FALSE TREVALLIES
Family AMMODYTIDAE (387) SAND LANCES Family DINOLESTIDAE (424) LONG-FINNED PIKES
Family TRACHINIDAE (388) WEEVERFISHES Family SCOMBROPIDAE (425) GNOMEFISHES
Family URANOSCOPIDAE (389) STARGAZERS Family POMATOMIDAE (426) BLUEFISHES
Proposed clade of all remaining percomorph orders: Family BRAMIDAE (427) POMFRETS
Order LABRIFORMES (77) WRASSES, RELATIVES Subfamily Braminae
Family LABRIDAE (390) WRASSES Subfamily Pteraclinae
Family ODACIDAE (391) CALES Family CARISTIIDAE (428) MANEFISHES
Family SCARIDAE (392) PARROTFISHES Possible relatives of Acanthuriformes:
Suggested clade of perciform relatives: Family MONODACTYLIDAE (429) MOONFISHES (FINGERFISHES)
Order PERCIFORMES (78) PERCHES Family PRIACANTHIDAE (430) BIGEYES (CATALUFAS)
Suborder Percoidei Seven families proposed relationship to
Superfamily Percoidea Acanthuroidei, Monodactylidae, Priacanthidae
Family CENTROPOMIDAE (393) SNOOKS Family LEIOGNATHIDAE (431) PONYFISHES, SLIMYS / SLIPMOUTHS
Family LATIDAE (394) LATES Family CHAETODONTIDAE (432) BUTTERFLYFISHES
Family GERREIDAE (395) MOJARRAS Family POMACANTHIDAE (433) ANGELFISHES
Family CENTROGENYIDAE (396) FALSE SCORPIONFISHES Family MALACANTHIDAE (434) TILEFISHES
Family PERCILIIDAE (397) SOUTHERN BASSES Subfamily Malacanthinae
Family HOWELLIDAE (398) OCEANIC BASSLETS Subfamily Latilinae
Family ACROPOMATIDAE (399) LANTERNBELLIES Family HAEMULIDAE (435) GRUNTS
Family EPIGONIDAE (400) DEEPWATER CARDINALFISHES Family HAPALOGENYIDAE (436) BARBELED GRUNTERS
Family POLYPRIONIDAE (401) WRECKFISHES Family LUTJANIDAE (437) SNAPPERS
Family LATEOLABRACIDAE (402) ASIAN SEAPERCHES Subfamily Etelinae
Family MULLIDAE (403) GOATFISHES Subfamily Apsilinae
Family GLAUCOSOMATIDAE (404) PEARL PERCHES Subfamily Paradicichthyinae
Family PEMPHERIDAE (405) SWEEPERS Subfamily Lutjaninae
Family SYMPHYSANODONTIDAE (405a) SLOPEFISHES Family CAESIONIDAE (438) FUSILIERS
Family OSTRACOBERYCIDAE (405b) OSTRACOBERYCIDS Superfamily Cirrhitoidea
Family OPLEGNATHIDAE (406) KNIFEJAWS Family CIRRHITIDAE (439) HAWKFISHES
Family KUHLIIDAE (407) FLAGTAILS Family CHIRONEMIDAE (440) KELPFISHES
Family LEPTOBRAMIDAE (407a) BEACHSALMON Family APLODACTYLIDAE (441) MARBLEFISHES
Family BATHYCLUPEIDAE (408) BATHYCLUPEIDS Family CHEILODACTYLIDAE (442) MORWONGS
Family POLYNEMIDAE (408a) THREADFINS Family LATRIDAE (443) TRUMPETERS
Family TOXOTIDAE (409) ARCHERFISHES Superfamily Cepoloidea
Family ARRIPIDAE (410) AUSTRALASIAN SALMON (KAHAWAI) Family CEPOLIDAE (444) BANDFISHES
Family DICHISTIIDAE (411) GALJOEN FISHES Subfamily Cepolinae
Family KYPHOSIDAE (412) SEA CHUBS Subfamily Owstoniinae
Subfamily Girellinae Superfamily Siganoidea
Subfamily Kyphosinae Family SCATOPHAGIDAE (445) SCATS
Subfamily Scorpidinae Family SIGANIDAE (446) RABBITFISHES
Subfamily Microcanthinae Suborder Notothenioidei
Subfamily Parascorpidinae Family BOVICHTIDAE (447) TEMPERATE ICEFISHES
Family TERAPONTIDAE (413) GRUNTERS / TIGERPERCHES Family PSEUDAPHRITIDAE (448) CATADROMOUS ICEFISHES
Family PERCICHTHYIDAE (414) TEMPERATE PERCHES Family ELEGINOPSIDAE (449) PATAGONIAN BLENNIES
Family SINIPERCIDAE (415) CHINESE PERCHES Family NOTOTHENIIDAE (450) COD ICEFISHES
Family ENOPLOSIDAE (416) OLDWIVES Family HARPAGIFERIDAE (451) SPINY PLUNDERFISHES
Family PENTACEROTIDAE (417) ARMORHEADS Family ARTEDIDRACONIDAE (452) BARBELED PLUNDERFISHES

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FISHES
Family BATHYDRACONIDAE (453) ANTARCTIC DRAGONFISHES Family SCORPAENICHTHYIDAE (487) CABEZON
Family CHANNICHTHYIDAE (454) CROCODILE ICEFISHES Family AGONIDAE (488) POACHERS, SEARAVENS
Order SCORPAENIFORMES (79) MAIL-CHEEKED FISHES Subfamily Hemilepidotinae
Suborder Scorpaenoidei Subfamily Hemitripterinae
Family SCORPAENIDAE (455) SCORPIONFISHES (ROCKFISHES) Subfamily Bothragoninae
Subfamily Sebastinae Subfamily Hypsagoninae
Tribe Sebastini Subfamily Anoplagoninae
Tribe Sebastolobini Subfamily Brachyopsinae
Subfamily Setarchinae Subfamily Agoninae
SubfamilyNeosebastinae Subfamily Bathyagoninae
SubfamilyScorpaeninae Family COTTIDAE (489) SCULPINS
Tribe Scorpaenini Subfamily Cottinae
Tribe Pteroini Subfamily Comephorinae
Subfamily Caracanthinae Subfamily Abyssocottinae
Subfamily Apistinae Family PSYCHROLUTIDAE (490) FATHEAD SCULPINS
Subfamily Tetraroginae Subfamily Cottunculinae
Subfamily Synanceiinae Subfamily Psychrolutinae
Tribe Minoini Family BATHYLUTICHTHYIDAE (491) ANTARCTIC SCULPINS
Tribe Choridodactylini Superfamily Cyclopteroidea
Tribe Synanceiini
Family CYCLOPTERIDAE (492) LUMPFISHES (LUMPSUCKERS)
Subfamily Plectrogeniinae
Family LIPARIDAE (493) SNAILFISHES
Family APLOACTINIDAE (456) VELVETFISHES
Proposed clade of the remaining orders of Percomorpha:
Family ESCHMEYERIDAE (457) COFISHES Order MORONIFORMES (80) TEMPERATE BASSES
Family PATAECIDAE (458) AUSTRALIAN PROWFISHES Family MORONIDAE (494) TEMPERATE BASSES
Family GNATHANACANTHIDAE (459) RED VELVETFISHES Family DREPANEIDAE (495) SICKLEFISHES
Family CONGIOPODIDAE (460) RACEHORSES (PIGFISHES / HORSEFISHES) Family EPHIPPIDAE (496) SPADEFISHES
Suborder Platycephaloidei Order ACANTHURIFORMES (81) SURGEONFISHES, RELATIVES
Family TRIGLIDAE (461) SEAROBINS (GURNARDS)
Suborder Sciaenoidei
Subfamily Prionotinae
Family EMMELICHTHYIDAE (497) ROVERS
Subfamily Pterygotriglinae
Family SCIAENIDAE (498) DRUMS (CROAKERS)
Subfamily Triglinae
Suborder Acanthuroidei
Family PERISTEDIIDAE (462) ARMORED SEAROBINS Family LUVARIDAE (499) LOUVAR
Family BEMBRIDAE (463) DEEPWATER FLATHEADS Family ZANCLIDAE (500) MOORISH IDOLS
Family PLATYCEPHALIDAE (464) FLATHEADS †Family MASSALONGIIDAE
Family HOPLICHTHYIDAE (465) GHOST FLATHEADS Family ACANTHURIDAE (501) SURGEONFISHES
Suborder Normanichthyoidei Subfamily Nasinae
Family NORMANICHTHYIDAE (466) BAREHEAD SCORPIONFISHES / Subfamily Acanthurinae
MOTE SCULPINS
Suborder Zoarcoidei Tribe Prionurini
Family BATHYMASTERIDAE (467) RONQUILS Tribe Zebrasomini
Family EULOPHIIDAE (468) Tribe Acanthurini

Family ZOARCIDAE (469) EELPOUTS Order SPARIFORMES (82) BREAMS, PORGIES


Family STICHAEIDAE (470) PRICKLEBACKS Family CALLANTHIIDAE (502) SPLENDID PERCHES / GROPPOS
Family CRYPTACANTHODIDAE (471) WRYMOUTHS Family SILLAGINIDAE (503) SILLAGOS (WHITINGS, SMELT-WHITINGS)
Family PHOLIDAE (472) GUNNELS Family LOBOTIDAE (504) TRIPLETAILS
Family ANARHICHADIDAE (473) WOLFFISHES Proposed clade of three families formerly called the “sparoid lineage”:
Family NEMIPTERIDAE (505) THREADFIN BREAMS
Family PTILICHTHYIDAE (474) QUILLFISHES
Family LETHRINIDAE (506) EMPERORS / EMPEROR BREAMS
Family ZAPRORIDAE (475) PROWFISHES
Family SPARIDAE (507) PORGIES
Family SCYTALINIDAE (476) GRAVELDIVERS
Proposed clade of the last three orders:
Suborder Gasterosteoidei
Family HYPOPTYCHIDAE (477) SAND EEL
Order CAPROIFORMES (83) BOARFISHES
Family CAPROIDAE (508) BOARFISHES
Family AULORHYNCHIDAE (478) TUBESNOUTS
Family GASTEROSTEIDAE (479) STICKLEBACKS
Antigoniinae
Subfamily

Family INDOSTOMIDAE (480) ARMORED STICKLEBACKS


Caproinae
Subfamily

Suborder Cottoidei
Order LOPHIIFORMES (84) ANGLERFISHES
Superfamily Anoplopomatoidea Suborder Lophioidei
Family LOPHIIDAE (509) GOOSEFISHES
Family ANOPLOPOMATIDAE (481) SABLEFISHES
Suborder Antennarioidei
Superfamily Zaniolepidoidea
Family ANTENNARIIDAE (510) FROGFISHES
Family ZANIOLEPIDIDAE (482) COMBFISHES
Family TETRABRACHIIDAE (511) TETRABRACHIID FROGFISHES
Subfamily Zaniolepidinae
Family LOPHICHTHYIDAE (512) LOPHICHTHYID FROGFISHES
Subfamily Oxylebiinae
Family BRACHIONICHTHYIDAE (513) HANDFISHES (WARTY ANGLERS)
Superfamily Hexagrammoidea
Suborder Chaunacoidei
Family HEXAGRAMMIDAE (483) GREENLINGS
Family CHAUNACIDAE (514) COFFINFISHES / SEA TOADS
Subfamily Ophiodontinae
Suborder Ogcocephaloidei
Subfamily Hexagramminae
Family OGCOCEPHALIDAE (515) BATFISHES
Subfamily Pleurogramminae
Suborder Ceratioidei
Superfamily Trichodontoidea Family CAULOPHRYNIDAE (516) FANFINS
Family TRICHODONTIDAE (484) SANDFISHES
Family NEOCERATIIDAE (517) SPINY SEADEVILS
Superfamily Cottoidea Family MELANOCETIDAE (518) BLACK SEADEVILS
Family JORDANIIDAE (485) LONGFIN SCULPIN
Family HIMANTOLOPHIDAE (519) FOOTBALLFISHES
Family RHAMPHOCOTTIDAE (486) GRUNT SCULPINS
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FISHES
FamilyDICERATIIDAE (520) DOUBLE ANGLERS
FamilyONEIRODIDAE (521) DREAMERS
FamilyTHAUMATICHTHYIDAE (522) WOLFTRAP ANGLERS
FamilyCENTROPHRYNIDAE (523) PRICKLY SEADEVILS
FamilyCERATIIDAE (524) WARTY SEADEVILS
FamilyGIGANTACTINIDAE (525) WHIPNOSE ANGLERS
FamilyLINOPHRYNIDAE (526) LEFTVENTS
Order TETRAODONTIFORMES (85) PLECTOGNATHS
†Suborder Plectocretacicoidei
†Family Cretatriacanthidae
†Family Plectocretacicidae
†Family Protriacanthidae
Suborder Triodontoidei
Family TRIODONTIDAE (527) THREETOOTH PUFFERS
Suborder Triacanthoidei
Family TRIACANTHODIDAE (528) SPIKEFISHES
Subfamily Hollardinae
Subfamily Triacanthodinae
Family TRIACANTHIDAE (529) TRIPLESPINES
Suborder Ostracioidea
Family ARACANIDAE (530) DEEPWATER BOXFISHES
Family OSTRACIIDAE (531) BOXFISHES (COWFISHES, TRUNKFISHES)
Suborder Balistoidei
Family BALISTIDAE (532) TRIGGERFISHES
Family MONACANTHIDAE (533) FILEFISHES
Suborder Tetraodontoidei
Family MOLIDAE (534) MOLAS
Family TETRAODONTIDAE (535) PUFFERS
Subfamily Tetraodontinae
Subfamily Canthigastrinae
Family DIODONTIDAE (536) PORCUPINEFISHES (BURRFISHES)

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